Senckenberg am Meer Wilhelmshaven

Marine Botany

JoP      Cover      Journal of Eukaryotic Microbiology          Cover

Peer-reviewed publications  


WILKE, T., ZINSSMEISTER, C. & HOPPENRATH, M. (2018): Morphological variability within the marine dinoflagellate Ornithocercus quadratus (Dinophysales, Dinophyceae). Phycologia 57(?): in press.

YAMAGUCHI, A., WAKEMAN, K.C., HOPPENRATH, M., HORIGUCHI, T. & KAWAI H. (2018): Molecular phylogeny of the benthic dinoflagellate Cabra matta (Dinophyceae) from Okinawa, Japan. Phycologia 57(?): in press.

ELBRÄCHTER, M., HOPPENRATH, M., JAHN, R. & KUSBER, W.-H. (2018): Stability of the generic names Alexandrium Halim and Gessnerium Halim at risk because of Peridinium splendor-maris Ehrenberg, the first documented bloom of Alexandrium (Dinophyceae). Notulae algarum No. 60

GOTTSCHLING, M., TILLMANN, U., KUSBER, W.-H., HOPPENRATH, M. & ELBRÄCHTER, M. (2018): (2607) Proposal to conserve the name Heterocapsa (Dinophyceae) with a conserved type. Taxon 67(3): .

GOTTSCHLING, M., TILLMANN, U., KUSBER, W.-H., HOPPENRATH, M. & ELBRÄCHTER, M. (2018): A Gordian knot: Nomenclature and taxonomy of Heterocapsa triquetra (Ehrenb.) Stein (Heterocapsaceae, Peridiniales). Taxon 67(1): 179-185.

WAKEMAN, K., HOPPENRATH, M., YAMAGUCHI, A., GAVELIS, G.S., LEANDER, B.S. & NOZAKI, H. (2018): Ultrastructure of the marine benthic dinoflagellate Plagiodinium belizeanum (Dinophyceae) from the southeast Pacific island of Okinawa, Japan. Phycologia 57(2): 209-222.

BRANDT, A., SCHOLZ, J., ALLSPACH, A., BRENKE, N., BRIX, S., GEORGE, K.H., HÖRNSCHEMEYER, T., HOLST, S., HOPPENRATH, M., IWAN, F., JANSSEN, A., JANSSEN, R., JANUSSEN, D., JESKULKE, K., FIEGE, D., KAISER, S., KIENEKE, A., KIHARA, T.C., KRÖNCKE, I., KRUPP, F., MARTHA, S.O., MARTÍNEZ ARBIZU, P.M., MEIßNER, K., MILJUTINA, M., MILJUTIN, D., RENZ, J., RIEHL, T., SAEEDI, H., SIEGLER, V., SONNEWALD, M., STUCKAS, H. & VEIT-KÖHLER, G. (2018): 200 years of marine research at Senckenberg: selected highlights. Marine Biodiversity: DOI 10.1007/s12526-017-0839-4

MERTENS, K.N., CARBONELL-MOORE, C., POSPELOVA, V., HEAD, M.J., HIGHFIELD, A., SCHROEDER, D., GU, H., ANDREE, K.B., YAMAGUCHI, A., TAKANO, Y., NÉZAN, E., BILIEN, G., OKOLODKOV, Y., KOIKE, K., HOPPENRATH, M., PFAFF, M., PITCHER G., AL-MUFTAH, A., ROCHON, A., LIM, P.T., LEAW, C.P., LIM, Z.F. & ELLEGAARD, M. (2018): The potentially toxic Pentaplacodinium saltonense gen. et sp. nov. (Dinophyceae), and its relationship to the cyst-defined Operculodinium psilatum and the yessotoxin-producing Protoceratium reticulatum. Harmful Algae 71: 57-77.


VERMA, A., KOHLI, G.S., HOPPENRATH, M., HARWOOD, T., KUZHIUMPARAMBIL, U., RALPH, P.J. & MURRAY, S.A. (2017): Systematics and diversity of genus Ostreopsis in the East Australian Current region. In: Proenca, L.A.O. & Hallegraeff, G.M. (eds) Marine and fresh-water harmful algae. Proceedings of the 17th International Conference on Harmful Algae. International Society for the Study of Harmful Algae 2017: 84-87.

TILLMANN, U., HOPPENRATH, M., GOTTSCHLING, M., KUSBER, W.-H. & ELBRÄCHTER, M. (2017): Plate pattern clarification of the marine dinophyte Heterocapsa triquetra sensu Stein (Dinophyceae) collected at the Kiel Fjord (Germany). Journal of Phycology 53: 1305-1324.

FRICKE, A., KIHARA, T.C., KOPPRIO, G.A. & HOPPENRATH, M. (2017): Anthropogenically driven habitat formation by a tube dwelling diatom on the Northern Patagonian Atlantic coast. Ecological Indicators 77: 8-13.

HOPPENRATH, M., YUBUKI, N., STERN, R. & LEANDER, B.S. (2017): Ultrastructure and molecular phylogenetic position of a new marine sand-dwelling dinoflagellate from British Columbia, Canada: Pseudadenoides polypyrenoides sp. nov. (Dinophyceae). European Journal of Phycology 52(2): 208-224.

FRICKE, A., KIHARA, T.C. & HOPPENRATH, M. (2017): Studying mesoalgal structures – a non-destructive approach based on confocal laser scanning microscopy (CLSM). Botanica Marina 60(2): 181-195.

ZINSSMEISTER, C., WILKE, T. & HOPPENRATH, M. (2017): Species diversity of dinophysoid dinoflagellates of the Clarion Clipperton Fracture Zone. Marine Biodiversity 47(2): 271-287.

KRETZSCHMAR, A.L., VERMA, A., HARWOOD, T., HOPPENRATH, M. & MURRAY, S.A. (2017): Characterisation of Gambierdiscus lapillus sp. nov. (Gonyaulacales, Dinophyceae): a new toxic dinoflagellate from the Great Barrier Reef (Australia). Journal of Phycology 53: 283-297.

HOPPENRATH, M. (2017): Dinoflagellate taxonomy - a review and proposal of a revised classification. Marine Biodiversity 47(2): 381-403.


VERMA, A., HOPPENRATH, M., DORANTES-ARANDA, J.-J., HARWOOD, T. & MURRAY S.A. (2016) Molecular phylogenetic characterization of Ostreopsis (Dinophyceae) and the description of a new species, Ostreopsis rhodesae sp. nov., from a subtropical Australian lagoon. Harmful Algae 60: 116-130.

YAMAGUCHI, A., YOSHIMATSU, S., HOPPENRATH, M., WAKEMAN K.C. & KAWAI, H. (2016): Molecular phylogeny of the benthic dinoflagellate genus Amphidiniopsis and its relationship with the family Protoperidiniaceae. Protist 167(6): 568-583.

VERMA, A., HOPPENRATH, M., HARWOOD, T., BRETT, S., RHODES, L. & MURRAY S.A. (2016) Molecular phylogeny, morphology and toxigenicity of Ostreopsis cf. siamensis (Dinophyceae) from temperate south-east Australia. Phycological Research 64: 146-159.


SELINA, M., CHOMÉRAT, N. & HOPPENRATH, M. (2015): Morphology and spatial distribution of Cabra species (Dinophyceae, Peridiniales) from Peter the Great Bay (northwestern Sea of Japan), including the description of C. levis sp. nov. European Journal of Phycology 50: 80-91.


WISSHAK, M., ALEXANDRAKIS, E. & HOPPENRATH, M. (2014): The diatom attachment scar Ophthalmichnus lyolithon igen. et isp. n. Ichnos 21: 111-118.

KOHLI, G.S., NEILAN, B.A., BROWN, M.V., HOPPENRATH, M.  & MURRAY S.A. (2014): Cob gene pyrosequencing enables characterisation of benthic dinoflagellate diversity and biogeography. Environmental Microbiology 16: 467-485.

MURRAY S.A., HOPPENRATH, M., ORR, R.J., BOLCH, C., JOHN, U., DIWAN, R., YAUWENAS, R., HARWOOD, T., DE SALAS, M., NEILAN, B.A. & HALLEGRAEFF, G. (2014): Alexandrium diversaporum nov. sp., a new non-saxitoxin producing species: phylogeny, morphology and sxtA genes. Harmful Algae 31: 54-65.


HOPPENRATH, M., CHOMÉRAT, N., HORIGUCHI, T., SCHWEIKERT, M., NAGAHAMA, Y. & MURRAY S. (2013): Taxonomy and phylogeny of the benthic Prorocentrum species (Dinophyceae) - a proposal and review. Harmful Algae 27: 1-28.

TILLMANN, U. & HOPPENRATH, M. (2013): The life-cycle of the heterotrophic dinoflagellate Polykrikos kofoidii. J. Phycol. 49: 298-317.

SELINA, M. & HOPPENRATH, M. (2013): Morphology and taxonomy of seven marine sand-dwelling Amphidiniopsis species (Peridiniales, Dinophyceae), including two new species A. konovalovae sp. nov. and A. striata sp. nov from the Sea of Japan, Russia. Marine Biodiversity 43: 87-104.

HOPPENRATH, M., CHOMÉRAT, N. & LEANDER, B.S. (2013): Molecular phylogeny of Sinophysis: Evaluating the possible early evolutionary history of dinophysoid dinoflagellates. In: Lewis, J.M., Marret, F. & Bradley, L. (eds.) Biological and Geological Perspectives of Dinoflagellates. The Micropalaeontological Society, Special publications. Geological Society, London: 207-214.


ADL, S.M, SIMPSON, A.G.B., LANE, C.E., LUKES, J., BASS, D., BOWSER, S.S., BROWN, M.W., BURKI, F., DUNTHORN, M., HAMPL, V., HEISS, A., HOPPENRATH, M., LARA, E., GALL, L.L., LYNN, D.H., MCMANUS, H.,MITCHELL, E.A.D., MOZLEY-STANRIDGE, S.E., PARFREY, L.W., PAWLOWSKI, J., RUECKERT, S., SHADWICK, L., SCHOCH, C.L., SMIRNOV, A. & SPIEGEL, F.W. (2012): The revised classification of eukaryotes. The Journal of Eukaryotic Microbiology 59: 429-493.

MURRAY, S.A., GARBY, T., HOPPENRATH, M. & NEILAN, B.A. (2012): Genetic diversity, morphological uniformity and polyketide production in dinoflagellates (Amphidinium, Dinoflagellata). PLoS ONE 7(6): e38253.

RÖDER, K., HANTZSCHE, F.M., GEBÜHR, C., MIENE, C., HELBIG, T., KROCK, B., HOPPENRATH, M., LUCKAS, B. & GERDTS, G. (2012): Effects of salinity, temperature and nutrients on growth, cellular characteristics and yessetoxin production of Protoceratium reticulatum. Harmful Algae 15: 59-70.

HOPPENRATH, M., MURRAY, S., SPARMANN, S. & LEANDER, B.S. (2012): Morphology and molecular phylogeny of Ankistrodinium gen. nov. (Dinophyceae), a new genus of marine sand-dwelling dinoflagellates formerly classified within Amphidinium. J. Phycol. 48: 1143-1152. 

SABUROVA, M., CHOMÉRAT, N. & HOPPENRATH, M. (2012): Morphology and SSU rDNA phylogeny of Durinskia agilis (Kofoid & Swezy) comb. nov. (Peridiniales, Dinophyceae), a thecate, marine, sand-dwelling dinoflagellate formerly classified within Gymnodinium. Phycologia 51: 287-302.

HOPPENRATH, M., SELINA, M., YAMAGUCHI, A. & LEANDER, B.S. (2012): Morphology and molecular phylogeny of Amphidiniopsis rotundata sp. nov. (Peridiniales, Dinophyceae), a new benthic marine dinoflagellate. Phycologia 51: 157-167.


YAMAGUCHI, A., HOPPENRATH, M., POSPELOVA, V., HORIGUCHI, T. & LEANDER, B.S. (2011): Molecular phylogeny of the marine sand-dwelling dinoflagellate Herdmania litoralis and an emended description of the closely related planktonic genus Archaeperidinium Jörgensen. Europ. J. Phycol. 46: 98-112. 


HOPPENRATH, M. & LEANDER, B.S. (2010): Dinoflagellates phylogeny as inferred from heat shock protein 90 and ribosomal gene sequences. PLoS ONE 5, e13220 (12 pages).

STERN, R., HORAK, A., ANDREW, R., COFFROTH, M.-A., ANDERSEN, R., KÜPPER, F., JAMESON, I., HOPPENRATH, M., VÉRON, B., KASAI, F., BRAND, J., JAMES, E., KEELING, P.J. (2010): Environmental barcoding reveals massive dinoflagellate diversity in marine environments. PLoS ONE 5, e13991 (14 pages).

GARCÈS, E. & HOPPENRATH, M. (2010): Ultrastructure of the intracellular parasite Parvilucifera sinerae (Alveolata, Myzozoa) infecting the marine toxic planktonic dinoflagellate Alexandrium minutum (Dinophyceae). Harmful Algae 10: 64-70.

CHANTANGSI, C., HOPPENRATH, M. & LEANDER, B.S. (2010): Evolutionary relationships among marine cercozoans as inferred from combined SSU and LSU rDNA sequences and polyubiquitin insertions. Mol. Phylo. Evol. 57, 518-527.

BREGLIA, S.A., YUBUKI, N., HOPPENRATH, M. & LEANDER, B.S. (2010): Ultrastructure and molecular phylogenetic position of a novel euglenozoan with extrusive episymbiotic bacteria: Bihospites bacati n. gen. et sp. (Symbiontida). BMC Microbiology 10: 145 (21 pages).

HOPPENRATH, M., YUBUKI, N., BACHVAROFF, T.R., & LEANDER, B.S. (2010): Re-classification of Pheoploykrikos hartmannii as Polykrikos (Dinophyceae) based partly on the ultrastructure of complex extrusomes. Europ. J. Protistol. 46: 29-37.


2009): Molecular phylogeny of ocelloid-bearing dinoflagellates (Warnowiaceae) as inferred from SSU and LSU rDNA sequences. BMC Evolutionary Biology 9: 116 (15pages).

HOPPENRATH, M. & LEANDER, B.S. (2009): Molecular phylogeny of Parvilucifera prorocentri (Alveolata, Myzozoa): insights into Perkinsid character evolution. J. Eukaryot. Microbiol. 56: 251-256.

HOPPENRATH, M., KOEMAN R.P.T. & LEANDER, B.S. (2009). Morphology and taxonomy of a new marine sand-dwelling Amphidiniopsis species (Dinophyceae, Peridiniales), A. aculeata sp. nov., from Cap Feret, France. Mar. Biodiv. 39: 1-7.


(2008): Comparative morphology and molecular phylogeny of taxa of the new marine benthic dinoflagellate genus Apicoporus, classified formerly within Amphidinium sensu lato. Protist 159: 383-399.

YAMAGUCHI, H., HOPPENRATH, M., TAKISHITA, K. & HORIGUCHI, T. (2008): Haramonas pauciplastida sp. nov. (Raphidophyceae, Heterokontophyta) and phylogenetic analyse of Haramonas species using small subunit ribosomal RNA gene sequences. Phycol. Res. 56: 127-138.

HOPPENRATH, M. & LEANDER, B.S. (2008):  Morphology and molecular phylogeny of a new marine sand-dwelling Prorocentrum species, P. tsawwassenense sp. nov. (Dinophyceae, Prorocentrales) from British Columbia, Canada. J. Phycol. 44: 451-466.

SELINA, M. & HOPPENRATH, M.   (2008): An emended description of Amphidiniopsis arenaria Hoppenrath 2000, based on material from the Sea of Japan. Europ. J. Protistol. 44: 71-79.

LEANDER, B.S.  & HOPPENRATH, M.   (2008): Ultrastructure of a novel tube-forming, intracellular parasite of dinoflagellates: Parvilucifera prorocentri sp. nov. (Alveolata, Myzozoa). Europ. J. Protistol. 44: 55-70.

TAYLOR, F.J.R., HOPPENRATH, M. & SALDARRIAGA, J.F. (2008): Dinoflagellate diversity and distribution. Biodiv. Conserv. 17: 407-418.


OKOLODKOV, Y.B., CAMPOS-BAUTISTA, G., Gárate-Lizárraga, I., González- González, J.A.G., Hoppenrath, M. & Arenas, V. (2007): Seasonal changes of benthic and epiphytic dinoflagellates in the Veracruz reef zone, Gulf of Mexico. Aquat. Microb. Ecol. 47: 223-237.   

HOPPENRATH, M.., Beszteri, B., Drebes, G., Halliger, H., Janisch, S., van Beusekom, J.E.E. & Wiltshire, K.H. (2007): Thalassiosira species (Bacillariophyceae, Thalassiosirales) in the waters around the islands of Helgoland (German Bight) and Sylt (Northfrisian Wadden Sea) – a first approach to assessing diversity. Europ. J. Phycol. 42: 271-288.

HOPPENRATH, M., M. Elbrächter, Halliger, H., Koeman, R.P.T., Krakhmalnyy, A., Surek, B., Erler, K. & Lucas, B. (2007): First rec ords of the benthic, bloom-forming, non-toxic dinoflagellate Thecadinium yashimaense (Dinophyceae) in Europe – with special emphasis on the invasion in the North Sea. Helgol. Mar. Res. 61: 157-165.

HOPPENRATH, M., Horiguchi, T., Miyoshi, Y., Selina, M., Taylor, F.J.R. & Leander, B.S. (2007): Taxonomy, phylogeny, biogeography, ecology of Sabulodinium undulatum, including an amended description of the species. Phycol. Res. 55: 159-175.

HOPPENRATH, M. & Leander, B.S. (2007): Morphology and phylogeny of the pseudocolonial dinoflagellates Polykrikos lebourae and Polykrikos herdmanae n. sp. Protist 158: 209-227.

HOPPENRATH, M. & Leander, B.S. (2007): Character evolution in polykrikoid dinoflagellates. J. Phycol. 43: 366-377.


HOPPENRATH, M. & Leander, B.S. (2006): Dinoflagellate, euglenoid or cercomonad? The ultrastructure and molecular phylogenetic position of Protaspis grandis n. sp.  J. Eukaryot. Microbiol. 53: 327-242.

HOPPENRATH, M. & Leander, B.S. (2006): Ebriid phylogeny and the expansion of the 'Cercozoa'. Protist 157: 279-290.

Murray, S., HOPPENRATH, M., Preisfeld, A., Larsen, J., Yoshimatsu, S., Toriumi, S. & Patterson, D.J. 
(2006): Phylogenetics of Rhinodinium broomeense gen. et sp. nov., a peridinoid, sand-dwelling dinoflagellate (Dinophyceae). J. Phycol. 42: 934-942.

HOPPENRATH, M. & Selina, M. (2006): Pseudothecadinium campbellii gen. nov. et sp. nov. (Dinophyceae), a phototrophic, thecate, marine planktonic species found in the Sea of Okhotsk, Russia. Phycologia 45: 260-269.

Murray, s., HOPPENRATH, M., Larsen, J. & Patterson, D.J. (2006): Bysmatrum teres sp. nov., a new sand-dwelling dinoflagellate from north-western Australia. Phycologia 45: 161-167.

Houpt, P. & Hoppenrath, M.
(2006): First record of the marine, benthic dinoflagellate Spiniferodinium galeiforme (Dinophyceae) from a temperate region. Phycologia 45: 10-12.  


HOPPENRATH, M., Bolch, C.J.S., Yoshimatsu, S., Saldarriaga, J.F., Schweikert, M., Campbell, C.N., Toriumi, S., Dodge, J.D., Elbrächter, M. & Taylor, F.J.R. (2005): Nomenclatural note on a Thecadinium species (Dinophyceae, Gonyaulacales), which was described as new independently three times within two months. J. Phycol. 41: 1284-1286.


Selina, M. & HOPPENRATH, M. (2004):  Morphology of Sinophysis minimum sp. nov., and three Sinophysis species (Dinophyceae, Dinophysiales) from the Sea of Japan. Phycol. Res. 52: 149-159.

HOPPENRATH, M. (2004): A revised check-list of planktonic diatoms and dinoflagellates from Helgoland (North Sea, German Bight). Helgol. Mar. Res. 58:243-251.   

HOPPENRATH, M., Saldarriaga, J.F., Schweikert, M., Elbrächter, M. & Taylor, F.J.R. (2004): Description of Thecadinium mucosum sp. nov. (Dinophyceae), a new sand-dwelling, marine dinoflagellate, and an emended description of Thecadinium inclinatum Balech. J. Phycol. 40:946-961.


HOPPENRATH, M., Schweikert, M. & Elbrächter, M. 
(2003): Morphological reinvestigation and characterisation of the marine, sand-dwelling dinoflagellate Adenoides eludens (Dinophyceae). Europ. J. Phycol. 38: 385-394.

Horiguchi, T. & HOPPENRATH, M. (2003): Haramonas viridis sp. nov. (Raphidophyceae, Heterokontophyta), a new sand-dwelling raphidophyte from cold temperate waters. Phycol. Res. 51: 61-67.


(2000): Morphology and taxonomy of six marine sand-dwelling Amphidiniopsis species (Dinophyceae, Peridiniales), four of them new, from the German Bight, North Sea. Phycologia 39: 482-497.

HOPPENRATH, M. (2000): An emended description of Herdmania litoralis Dodge (Dinophyceae) including the plate formula. Nova Hedwigia 71: 481-489.

HOPPENRATH, M. (2000): Morphology and taxonomy of the marine sand-dwelling genus Thecadinium (Dinophyceae), with the description of two new species from the North German Wadden Sea. Phycologia 39: 96-108.

HOPPENRATH, M. (2000): A new marine sand-dwelling Prorocentrum species, P. clipeus sp. nov. (Dinophyceae, Prorocentrales) from Helgoland, German Bight, North Sea. Europ. J. Protistol. 36: 29-33.

HOPPENRATH, M. (2000): Morphology and taxonomy of Sinophysis (Dinophyceae, Dinophysiales) including two new marine sand-dwelling species from the North German Wadden Sea. Europ. J. Phycol. 35: 153-162.

HOPPENRATH, M. & Okolodkov, Y.B . (2000): Amphidinium glabrum sp. nov. (Dinophyceae) from the North German Waddense and European Arctic sea ice: morphology, distribution and ecology. Europ. J. Phycol. 35: 61-67.


HOPPENRATH, M. & Elbrächter, M. 
(1998): Roscoffia capitata (Dinophyceae) refound: notes on morphology and biology. Phycologia 37 (6): 450-457.

Robinson, D.G., HOPPENRATH, M., Oberbeck, K., Luykx, P. & Ratajczak, R. (1998): Localization of Pyrophosphatase and V-ATPase in Chlamydomonas reinhardtii. Bot. Acta 111: 108-122.

Luykx, P., HOPPENRATH, M. & Robinson, D.G. 
(1997): Osmoregulatory mutants that affect the function of the contractile vacuole in Chlamydomonas reinhardtii. Protoplasma 200: 99-111.

Luykx, P., HOPPENRATH, M. & Robinson, D.G. (1997): Structure and behavior of contractile vacuoles in Chlamydomonas reinhardtii. Protoplasma 198: 73-84.


not peer-reviewed publications (e.g. presentation abstracts)

(2011) The new Herbarium Senckenbergianum: old institutions under a new common roof. TAXON

HOPPENRATH, M. (2001): Biodiversity of marine sand-dwelling flagellates from the North German Wadden Sea., Phycologia 40 (4), Supplement, 114.

HOPPENRATH, M., Drebes, G., Elbrächter, M., Scharek, R., Beusekom, J. van & Wiltshire, K. (2001): The Helgoland and Sylt Marine Phytoplankton Time-Series (AWI)., Phycologia 40 (4), Supplement, 105.

(1998): Sand-dwelling dinoflagellates of the Wadden Sea (List/Sylt), with special emphasis on Roscoffia capitata., NTNU Vitensk. mus. Rapp. bot. Ser. 1998-I; 69-70.


(2011) Aufbau einer Dinoflagellaten-Sammlung - Mikrometerkleine Lebewesen kommen bei Senckenberg groß raus. SENCKENBERG nfm 141(3/4): 118-121. 

DRESSLER, S., GEBAUER, P., HOPPENRATH, M., KIENAST, F., OTTE, V. & ZIZKA, G. (2011) Das neue Herbarium Senckenbergianum - Alte Institutionen unter einem gemeinsamen neuen Dach. SENCKENBERG nfm 141(3/4): 110-113.

HOPPENRATH, M. (2004): Im Sand brodelt das Leben. Die Geißeltierchen. In: Watt’n Leben! Tiere und Pflanzen der Nordseeküste vorgestellt von der Wattenmeerstation Sylt. AWI.

HOPPENRATH, M. (2004): Ein leuchtendes Meerestier. Der Dinoflagellat Noctiluca scintillans. In: Watt’n Leben! Tiere und Pflanzen der Nordseeküste vorgestellt von der Wattenmeerstation Sylt. AWI.