Senckenberg – Leibniz Institution for Biodiversity and Earth System Research

Board of Directors

The Board of Directors manage and administrate the Senckenberg – Leibniz Institution for Biodiversity and Earth System Research, as well as lead staff meetings. The board consists of the Director General who serves as chairman and up to five other members. The additional members of the Board of Directors include the Deputy Director General and the Administrative Director, who are appointed by the Board of Directors for a period of up to five years. The final two members represent the Senckenberg Society for Nature Research. 

Members of the Board of Directors

Tockner Jun 2021
Prof. Dr. Klement Tockner
Director General of Senckenberg – Leibniz Institution for Biodiversity and Earth System Research

Klement Tockner Publications


Summary Information (Status: March 2021)

Total: ~ 250, hereof 180 ISI-publications (1997-2020: in average >10 publications per year; >7 ISI publications per year)

h-Index (Google Scholar): 81; Total # of citations: 27,687 (maximum 1,869 citations per publication); highly cited papers with more than 200 citations each: 36


Drenckhahn, D., Arneth, A., Filser, J., Haberl, H., Hansjürgens, B., Herrmann, B., … Tockner, K. (2020). Globale Biodiversität in der Krise: Was können Deutschland und die EU dagegen tun? Deutsche Akademie der Naturforscher Leopoldina eVNationale Akademie der Wissenschaften.

Mehner, T., Likens, G. E., & Tockner, K. (Eds.). (2009). Encyclopedia of Inland Waters. Academic Press.

Tockner, K., Uehlinger, U., Robsinson, C. T. (Eds.). (2009). Rivers of Europe. Academic Press. (2nd Edition: 2021).

Woolsey, S., Weber, C., Gonser, T., Hoehn, E., Hostmann, M., Junker, B., Roulier, C., Schweizer, S., Tiegs, S., Tockner, K., Peter, A. (2005). Handbook for evaluating rehabilitation projects in rivers and streams: A publication by the Rhone-Thur project. Eawag, WSL, LCH-EPFL, VAW-ETHZ.

Special issues

Lewandowski, J., Nützmann, G., & Tockner, K. (Eds.). (2014). Frontiers in real-time ecohydrology. Fundamental and Applied Limnology, 184, 3.

Milner, A. M., & Tockner, K. (Eds.). (2010). Freshwater Benthic Science – what has it contributed to ecological theory? River Research and Applications, 26.

Gurnell, A., Tubino, M., & Tockner, K. (Eds.). (2009). Linkages and feedbacks in highly dynamic alpine fluvial systems. Aquatic Sciences, 71.

Tockner, K., Kollmann, J., Edwards, P. E., & Ward, J. V. (Eds.). (2002). Riverine Landscapes. Freshwater Biology, 47.


Peer-reviewed journal articles (ISI listed)



Cantonati, M., Fensham, R. J., Stevens, L. E., Gerecke, R., Glazier, D. S., Goldscheider, N., … Tockner, K. (2021). Urgent plea for global protection of springs. Conservation Biology, 35(1), 378-382.

Jähnig, S. C., Baranov, V., Altermatt, F., Cranston, P., Friedrichs‐Manthey, M., … Tockner, K., … Domisch, S. (2021). Revisiting global trends in freshwater insect biodiversity. WIREs Water, 8(2), e1506.

Peters, R., Berlekamp, J., Lucía, A., Stefani, V., Tockner, K., & Zarfl, C. (2021). Integrated Impact Assessment for Sustainable Hydropower Planning in the Vjosa Catchment (Greece, Albania). Sustainability, 13(3), 1514.


He, F., Langhans, S. D., Zarfl, C., Wanke, R., Tockner, K., & Jähnig, S. C. (2020). Combined effects of life‐history traits and human impact on extinction risk of freshwater megafauna. Conservation Biology, 35(2), 643-653.

Heger, T., Bernard-Verdier, M., Gessler, A., Greenwood, A. D., Grossart, H.-P., … Tockner, K., … Jeschke, J. M. (2020). Clear Language for Ecosystem Management in the Anthropocene: A Reply to Bridgewater and Hemming. BioScience, 70(5), 374-376.

Moleón, M., Sánchez-Zapata, J. A., Donázar, J. A., Revilla, E., Martín-López, B., Gutiérrez-Cánovas, C., … Tockner, K. (2020). Rethinking megafauna. Proceedings of the Royal Society B, 287(1922), 20192643.

Sánchez-Montoya, M. M., Tockner, K., von Schiller, D., Miñano, J., Catarineu, C., Lencina, J. L., … Ruhi, A. (2020). Dynamics of ground-dwelling arthropod metacommunities in intermittent streams: The key role of dry riverbeds. Biological Conservation, 241, 108328.

Serlet, A. J., López Moreira M, G. A., Zolezzi, G., Wharton, G., Hölker, F., … Tockner, K., … Zarfl, C. (2020). SMART Research: Toward Interdisciplinary River Science in Europe. Frontiers in Environmental Science, 8, 63.  

Thieme, M. L., Khrystenko, D., Qin, S., Golden Kroner, R. E., Lehner, B., … Tockner, K., … Mascia, M. B. (2020). Dams and protected areas: Quantifying the spatial and temporal extent of global dam construction within protected areas. Conservation Letters, 13(4), e12719.

Tickner, D., Opperman, J. J., Abell, R., Acreman, M., Arthington, A. H., … Tockner, K., … Young, L. (2020). Bending the Curve of Global Freshwater Biodiversity Loss: An Emergency Recovery Plan. BioScience, 70(4), 330-342.  


Grill, G., Lehner, B., Thieme, M., Geenen, B., Tickner, D., … Tockner, K., … Zarfl, C. (2019). Mapping the world’s free flowing rivers. Nature, 569(7755), 215-221.

He, F., Zarfl, C., Bremerich, V., David, J. N. W., Hogan, Z., … Tockner, K., Jähnig, S. C. (2019). The global decline of freshwater megafauna. Global Change Biology, 25(11), 3883-3892.

Heger, T., Bernard-Verdier, M., Gessler, A., Greenwood, A. D., Grossart, H.-P., … Tockner, K., … Jeschke, J. M. (2019). Towards an Integrative, Eco-Evolutionary Understanding of Ecological Novelty: Studying and Communicating Interlinked Effects of Global Change. BioScience, 69(11), 888-899.

Hickisch, R., Hodgetts, T., Johnson, P. J., Sillero‐Zubiri, C., Tockner, K., & Macdonald, D. (2019). Effects of publication bias on conservation planning. Conservation Biology, 33(5), 1151-1163.

Jeschke, J. M., Börner, K., Stodden, V., & Tockner, K. (2019). Open Access journals need to become first choice, in invasion ecology and beyond. NeoBiota, 52, 1-8.

Jeschke, J. M., Lokatis, S., Bartram, I. & Tockner, K. (2019). Knowledge in the dark: scientific challenges and ways forward. FACETS, 4(1), 423-441.

Kyba, C. M. C., Giuliani, G., Franziskakis, F., Tockner, K., & Lacroix, P. (2019). Artisanal and Small-Scale Mining Sites in the Democratic Republic of the Congo Are Not Associated with Nighttime Light Emissions. J Multidisciplinary Science Journal, 2(2), 152-161.

Larsen, S., Karaus, U., Claret, C., Sporka, F., Hamerlík, L., & Tockner, K. (2019). Flooding and hydrologic connectivity modulate community assembly in a dynamic riverfloodplain ecosystem. PLoS ONE, 14(4), e0213227.

Pinter, N., Brasington, J., Gurnell, A., Kondolf, G. M., Tockner, K., Wharton, G., & Yarnell, S. M. (2019). River research and applications across borders. River  Research and Applications, 35(6), 768-775.

Reid, A. J., Carlson, A. K., Creed, I. F., Eliason, E. J., Gell, P. A., … Tockner, K., … Cooke, S. J. (2019). Emerging threats and persistent conservation challenges for freshwater biodiversity. Biological Reviews, 94(3), 849-873.

Schmidt-Kloiber, A., Bremerich, V., De Wever, A., Jähnig, S. C., Martens, K., … Tockner, K., Hering, D. (2019). The Freshwater Information Platform: a global online network providing data, tools and resources for science and policy support. Hydrobiologia, 838(1), 1-11.

Schmidt-Kloiber, A., De Wever, A., Bremerich, V., Strackbein, J., Hering, D., Jähnig, S., … Tockner, K. (2019). The Freshwater Information Platform: An online network supporting freshwater biodiversity research and data publishing. Biodiversity Information Science and Standards, 3, e37378.

Shumilova, O., Tockner, K., Gurnell, A. M., Langhans, S. D., Righetti, M., Lucía, A., & Zarfl, C. (2019). Floating matter: a neglected component of the ecological integrity of rivers. Aquatic Sciences, 81(2), 25.

Shumilova, O., Zak, D., Datry, T., von Schiller, D., Corti, R., … Tockner, K., … Zarfl, C. (2019). Simulating rewetting events in intermittent rivers and ephemeral streams: A global analysis of leached nutrients and organic matter. Global  Change Biology, 25(5), 1591-1611.

Tydecks, L., Ibelings, B. W., & Tockner, K. (2019). A global survey of freshwater biological field stations. River Research and Applications, 35(8), 1314-1324.

von Schiller, D., Datry, T., Corti, R., Foulquier, A., Tockner, K., Marcé, R., … Zoppini, A. (2019). Sediment Respiration Pulses in Intermittent Rivers and Ephemeral Streams. Global Biogeochemical Cycles, 33(10), 1251-1263.

Zarfl, C., Berlekamp, J., He, F., Jähnig, S. C., Darwall, W., & Tockner, K. (2019). Future large hydropower dams impact global freshwater megafauna. Scientific Reports, 9(1), 18531.


Darwall, W., Bremerich, V., De Wever, A., Dell, A. I., Freyhof, J., … Tockner, K., … Weyl, O. (2018). The Alliance for Freshwater Life: A global call to unite efforts for freshwater biodiversity science and conservation. Aquatic Conservation, 28(4), 1015-1022.

Datry, T., Boulton, A. J., Bonada, N., Fritz, K., Leigh, C., … Tockner, K., … Dahm, C. N. (2018). Flow intermittence and ecosystem services in rivers of the Anthropocene. Journal of Applied Ecology, 55(1), 353-364.

Datry, T., Foulquier, A., Corti, R., von Schiller, D., Tockner, K., Mendoza-Lera, C., … Zoppini, A. (2018). A global analysis of terrestrial plant litter dynamics in non-perennial waterways. Nature Geoscience, 11(7), 497-503.

de la Fuente, M., Bonada, N., Bêche, L., Dahm, C. N., Mendez, P. K., Tockner, K., … Acuña, V. (2018). Evolutionary responses of aquatic macroinvertebrates to two contrasting flow regimes. Hydrobiologia, 808(1), 353-370.

He, F., Bremerich, V., Zarfl, C., Geldmann, J., Langhans, S. D., … Tockner, K., Jähnig, S. C. (2018). Freshwater megafauna diversity: Patterns, status and threats. Diversity and Distributions, 24(10), 1395-1404.

Lumsdon, A. E., Artamonov, I., Bruno, M. C., Righetti, M., Tockner, K., Tonolla, D., & Zarfl, C. (2018). Soundpeaking – Hydropeaking induced changes in river soundscapes. River Research and Applications, 34(1), 3-12.

Marshall, J. C., Acuña, V., Allen, D. C., Bonada, N., Boulton, A. J., … Tockner, K., Vander Vorste, R. (2018). Protecting U.S. temporary waterways. Science, 361(6405), 856-857.

Sánchez-Montoya, M. M., von Schiller, D., Barberá, G. G., Díaz, A. M., Arce, M. I., del Campo, R., & Tockner, K. (2018). Understanding the effects of predictability, duration, and spatial pattern of drying on benthic invertebrate assemblages in two contrasting intermittent streams. PLoS ONE, 13(3), e0193933.

Shumilova, O., Tockner, K., Thieme, M., Koska, A., & Zarfl, C. (2018). Global Water Transfer Megaprojects: A Potential Solution for the Water-Food-Energy Nexus? Frontiers in Environmental Science, 6, 150.

Turnbull, L., Hütt M.-C., Ioannides, A. A., Kininmonth, S., Poeppl, R., Tockner, K., … Parsons, A. J. (2018). Connectivity and complex systems: learning from a multidisciplinary perspective. Applied Network Science, 3(1), 11.

Tydecks, L., Jeschke, J. M., Wolf, M., Singer, G., & Tockner, K. (2018). Spatial and topical imbalances in biodiversity research. PLoS ONE, 13(7), e0199327.


Carrizo, S. F., Jähnig, S. C., Bremerich, V., Freyhof, J., Harrison, I., … Tockner, K., … Darwall, W. (2017). Freshwater Megafauna: Flagships for Freshwater Biodiversity under Threat. BioScience, 67(10), 919-927.

He, F., Zarfl, C., Bremerich, V., Henshaw, A., Darwall, W., Tockner, K., & Jähnig, S.C. (2017). Disappearing giants: a review of threats to freshwater megafauna. WIREs Water, 4(3), e1208.

Leigh, C., Laporte, B., Bonada, N., Fritz, K., Pella, H., … Tockner, K., Datry, T. (2017). IRBAS: An online database to collate, analyze, and synthesize data on the biodiversity and ecology of intermittent rivers worldwide. Ecology and Evolution, 7(3), 815-823.

Skoulikidis, N. T., Sabater, S., Datry, T., Morais, M. M., Buffagni, A., Dörflinger, G., … Tockner, K. (2017). Non-perennial Mediterranean rivers in Europe: Status, pressures, and challenges for research and management. Science of the Total  Environment, 577, 1-18.

Sommerwerk, N., Wolter, C., Freyhof, J., & Tockner, K. (2017). Components and drivers of change in European freshwater fish faunas. Journal of Biogeography, 44(8), 1781-1790.

Tockner, K., & Jeschke, J. M. (2017). Wissenschaft in Umbruchzeiten. Laborjournal, 78, 16-17. 

Tockner, K., & Jeschke, J. M. (2017). Freie und unabhängige Wissenschaft: Fundament aufgeklärter Gesellschaften. In H. Hösele & L. Wieser (Eds.), Europa. USA. 3.0: Werte, Interessen, Perspektiven (pp. 199-213).Wieser.


Arora, R., Tockner, K., & Venohr, M. (2016). Changing river temperatures in northern Germany: trends and drivers of change. Hydrological Processes, 30(17), 3084-3096

Bismuth, C., Hansjürgens, B., Moss, T., Hoechstetter, S., Tockner, K., Yorke, V., … Yaramanci, U. (2016). Lessons Learnt, Open Research Questions and Recommendations. In R. F. Hüttl, O. Bens, C. Bismuth, & S. Hoechstetter (Eds.), Society – Water – Technology: Water Resources Development and Management (pp. 279-292). Springer.

Gurnell, A. M., Bertoldi, W., Tockner, K., Wharton, G., & Zolezzi, G. (2016). How large is a river? Conceptualizing river landscape signatures and envelopes in four dimensions. WIREs Water, 3(3), 313-325.

Hansjürgens, B., Droste, N., & Tockner, K. (2016). Neglected Values of Major Water Engineering Projects: Ecosystem Services, Social Impacts, and Economic Valuation. In R. F. Hüttl, O. Bens, C. Bismuth, & S. Hoechstetter (Eds.), Society – Water – Technology: Water Resources Development and Management (pp. 65-78). Springer.

Jackson, M. C., Weyl, O. L. F., Altermatt, F., Durance, I., Friberg, N, … Tockner, K., … Woodward, G. (2016). Chapter Twelve – Recommendations for the Next Generation of Global Freshwater Biological Monitoring Tools. Advances in  Ecological Research, 55, 615-636.

Jeschke, J. M., Kaushal, S. S., & Tockner, K. (2016, April 8). Diversifying Skills and Promoting Teamwork in Science. Eos.

Le, Q. N., Fujii, M., Yoshimura, C., & Tockner, K. (2016). Dissolved nitrogen release from coarse and amphipod-produced fine particulate organic matter in freshwater column. Limnology, 17(1), 33-46.

Sánchez‐Montoya, M. M., Moleón, M., Sánchez‐Zapata, J. A., & Tockner, K. (2016). Dry riverbeds: corridors for terrestrial vertebrates. Ecosphere, 7(10), e01508.

Sánchez‐Montoya, M. M., von Schiller, D., Ruhí, A., Pechar, G. S., Proia, L., Miñano, J., … Tockner, K. (2016). Responses of ground-dwelling arthropods to surface flow drying in channels and adjacent habitats along Mediterranean streams. Ecohydrology, 9(7), 1376-1387.

Tockner, K., Bernhardt, E. S., Koska, A., & Zarfl, C. (2016). A Global View on Future Major Water Engineering Projects. In R. F. Hüttl, O. Bens, C. Bismuth, & S. Hoechstetter (Eds.), Society – Water – Technology: Water Resources Development and Management (pp. 47-64). Springer.

Tydecks, L., Bremerich, V., Jentschke, I., Likens, G. E., & Tockner, K. (2016). Biological Field Stations: A Global Infrastructure for Research, Education, and Public Engagement. BioScience, 66(2), 164-171.

Tydecks, L., Bremerich, V., Jentschke, I., Likens, G.E., & Tockner, K. (2016). Global Database on Biological Field Stations a pivotal infrastructure for environmental research, education and public information. Limnology and Oceanography Bulletin, 25(3), 88-88.



Diener, S., Zurbrügg, C., & Tockner, K. (2015). Bioaccumulation of heavy metals in the black soldier fly, Hermetia illucens and effects on its life cycle. Journal of Insects as Food and Feed, 1(4), 261-270.

Krause, S., Lewandowski, J., Dahm, C. N., & Tockner, K. (2015). Frontiers in real-time ecohydrology – a paradigm shift in understanding complex environmental systems. Ecohydrology, 8(4), 529-537.

Rillig, M. C., Kiessling, W., Borsch, T., Gessler, A., Greenwood, A. D., … Tockner, K., … Jeltsch, F. (2015). Biodiversity research: data without theory – theory without data. Frontiers in Ecology and Evolution, 3, 20.

Rosado, J., Morais, M., & Tockner, K. (2015). Mass Dispersal of Terrestrial Organisms During First Flush Events in a Temporary Stream. River Research and  Applications, 31(7), 912-917.

von Schiller, D., Graeber, D., Ribot, M., Timoner, X., Acuña, V., Martí, E., … Tockner, K. (2015). Hydrological transitions drive dissolved organic matter quantity and composition in a temporary Mediterranean stream. Biogeochemistry, 123(3), 429-446.

Zarfl, C., Lumsdon, A. E., Berlekamp, J., Tydecks, L., & Tockner, K. (2015). A global boom in hydropower dam construction. Aquatic Sciences, 77(1), 161-170.


Acuña, V., Datry, T., Marshall, J., Barceló, D., Dahm, C. N., … Tockner, K., Palmer, M. A. (2014). Why Should We Care About Temporary Waterways? Science, 343(6175), 1080-1081.  

Datry, T., Larned, S. T., & Tockner, K. (2014). Intermittent Rivers: A Challenge for Freshwater Ecology. BioScience, 64(3), 229-235.

Langhans, S. D., & Tockner, K. (2014). Is the unsaturated sediment a neglected habitat for riparian arthropods? Evidence from a large gravel-bed river. Global Ecology and Conservation, 2, 129-137.

Langhans, S. D., & Tockner, K. (2014). Edge Effects Are Important in Supporting Beetle Biodiversity in a Gravel-Bed River Floodplain. PLOS ONE, 9(12), e114415.

Muehlbauer, J. D., Collins, S. F., Doyle, M. W., & Tockner, K. (2014). How wide is a stream? Spatial extent of the potential “stream signature” in terrestrial food webs using meta‐analysis. Ecology, 95(1), 44-55.

Perkin, E. K., Hölker, F., & Tockner, K. (2014). The effects of artificial lighting on adult aquatic and terrestrial insects. Freshwater Biology, 59(2), 368-377.

Perkin, E. K., Hölker, F., Tockner, K., & Richardson, J. S. (2014). Artificial light as a disturbance to light-naïve streams. Freshwater Biology, 59(11), 2235-2244.

Prat, N., Gallart, F., Von Schiller, D., Polesello, S., García-Roger, E. M., … Tockner, K., Froebrich, J. (2014). The Mirage Toolbox: An integrated assessment tool for temporary streams. River Research and Applications, 30(10), 1318-1334.

Settele, J., Scholes, R., Betts, R. A., Bunn, S., Leadley, P., … Tockner, K., … Winter, M. (2014). Terrestrial and Inland Water Systems. In C. B. Field, V. R. Barros, D. J. Dokken, K. J. Mach, M. D. Mastrandrea, T. E. Bilir, … L. L. White (Eds.), Climate  Change 2014: Impacts, Adaptation, and Vulnerability. Part A: Global and  Sectoral Aspects. Contribution of Working Group II to the Fifth Assessment  Report of the Intergovernmental Panel on Climate Change (pp. 271-359). Cambridge University Press.

Shrestha, J., Niklaus, P. A., Pasquale, N., Huber, B., Barnard, R. L., …Tockner, K., Luster, J. (2014). Flood pulses control soil nitrogen cycling in a dynamic river floodplain. Geoderma, 228-229, 14-24.

Tockner, K. (2014). Novel freshwater ecosystems in a changing climate: a challenge for research and conservation. In H. Korn, J. Stadler, A. Bonn, K. Bockmühl, & N. Macgregor (Eds.), Proceedings of the European Conference “Climate Change and Nature Conservation in Europe – an ecological, policy and economic perspective” (BfN-Skripten 367, pp. 46-47). Bundesamt für Naturschutz. Tockner, K. (2014). Der Wert der Vielfalt. Leibniz-Journal, 3, 14-15.


Doering, M., Uehlinger, U., & Tockner, K. (2013). Vertical hydrological exchange, and ecosystem properties and processes at two spatial scales along a floodplain river (Tagliamento, Italy). Freshwater Science, 32(1), 12-25.

Hanafiah, M. M., Leuven, R. S. E. W., Sommerwerk, N., Tockner, K., & Huijbregts, M. A. J. (2013). Including the Introduction of Exotic Species in Life Cycle Impact Assessment: The Case of Inland Shipping. Environmental Science & Technology, 47(24), 13934-13940.

Karaus, U., Larsen, S., Guillong, H., & Tockner, K. (2013). The contribution of lateral aquatic habitats to insect diversity along river corridors in the Alps. Landscape Ecology, 28(9), 1755-1767.

Langhans, S. D., Richard, U., Rueegg, J., Uehlinger, U., Edwards, P., Doering, M., & Tockner, K. (2013). Environmental heterogeneity affects input, storage, and transformation of coarse particulate organic matter in a floodplain mosaic. Aquatic Sciences, 75(3), 335-348.

Ostojić, A., Rosado, J., Miliša, M., Morais, M., & Tockner, K. (2013). Release of nutrients and organic matter from river floodplain habitats: simulating seasonal inundation dynamics. Wetlands, 33(5), 847-859.

Pahl-Wostl, C., Arthington, A., Bogardi, J., Bunn, S. E., Hoff, H., … Tockner, K., Tsegai, D. (2013). Environmental flows and water governance: managing sustainable water uses. Current Opinion in Environmental Sustainability, 5(3-4), 341-351.

Tockner, K. (2013). Managing river flood plains as ecosystems of global strategic importance. In European Environment Agency (Ed.), Late lessons from early warnings: science, precaution, innovation (EEA Report No. 1, pp. 349-350). EEA.

Tockner, K., & Grossart, H.-P. (2013). Biodiversität der Binnengewässer. In E. Beck (Ed.), Die Vielfalt des Lebens: Wie hoch, wie komplex, warum? (pp. 121-128). Wiley-VCH.


Beniston, M., Stoffel, M., Harding, R., Kernan, M., Ludwig, R., Moors, E., … Tockner, K. (2012). Obstacles to data access for research related to climate and water: Implications for science and EU policy-making. Environmental Science & Policy, 17, 41-48.

De Wever, A., Schmidt-Kloiber, A., Gessner, M. O., & Tockner, K. (2012). Freshwater Journals Unite to Boost Primary Biodiversity Data Publication. BioScience, 62(6), 529-530.

Handcock, R. N., Torgersen, C. E., Cherkauer, K. A., Gillespie, A. R., Tockner, K., Faux, R. N., & Tan, J. (2012). Thermal Infrared Remote Sensing of Water Temperature in Riverine Landscapes. In P. E. Carbonneau & H. Piégay (Eds.), Fluvial Remote Sensing for Science and Management (pp. 85-113). Wiley-Blackwell. 

Peter, S., Rechsteiner, R., Lehmann, M. F., Brankatschk, R., Vogt, T., … Tockner, K., Durisch-Kaiser, E. (2012). Nitrate removal in a restored riparian groundwater system: functioning and importance of individual riparian zones. Biogeosciences, 9(11), 4295-4307.

Ricaurte, L. F., Boesch, S., Jokela, J., & Tockner, K. (2012). The distribution and environmental state of vegetated islands within human-impacted European rivers. Freshwater Biology, 57(12), 2539-2549.

Shrestha, J., Niklaus, P. A., Frossard, E., Samaritani, E., Huber, B., … Tockner, K., Luster, J. (2012). Soil Nitrogen Dynamics in a River Floodplain Mosaic. Journal of Environmental Quality, 41(6), 2033-2045.

Steward, A.L., von Schiller, D., Tockner, K., Marshall, J.C., & Bunn, S.E. (2012). When the river runs dry: human and ecological values of dry riverbeds. Frontiers in  Ecology and the Environment, 10(4), 202-209.

Tockner, K. (2012). Vom struktur- zum funktionsbasierten Gewässermanagement. Natur und Mensch, 54(2), 18-21.

Tockner, K., Gessner, J., Pusch, M., & Wolter, C. (2012). Domestizierte Ökosysteme und neuartige Lebensgemeinschaften: Herausforderungen für das Gewässermanagement. In U. Grünewald, O. Bens, H. Fischer, R.F. Hüttl, K. Kaiser, & A. Knierim (Eds.), Wasserbezogene Anpassungsmaßnahmen an den Landschafts- und Klimawandel (pp. 33-42). Schweizerbart.

Tonolla, D., Wolter, C., Ruhtz, T., & Tockner, K. (2012). Linking fish assemblages and spatiotemporal thermal heterogeneity in a river-floodplain landscape using high-resolution airborne thermal infrared remote sensing and in-situ measurements. Remote Sensing of Environment, 125, 134-146.


Diener, S., Studt Solano, N. M., Roa Gutiérrez, F., Zurbrügg, C., & Tockner, K. (2011). Biological Treatment of Municipal Organic Waste using Black Soldier Fly Larvae. Waste and Biomass Valorization, 2(4), 357-363.

Diener, S., Zurbrügg, C., Gutiérrez, F. R., Nguyen, D. H., Morel, A., Koottatep, T., & Tockner, K. (2011). Black soldier fly larvae for organic waste treatment – prospects and constraints. In M. Alamgir, Q. H. Bari, I. M. Rafizul, S. M. T. Islam,

  1. Sarkar, & M. K. Howlader (Eds.), Proceedings of the WasteSafe 2011 – 2nd International Conference on Solid Waste Management in the Developing Countries, 13-15 February 2011, Khulna, Bangladesh (pp. 52 (1-8)).

Dieter, D., von Schiller, D., García-Roger, E. M., Sánchez-Montoya, M. M., Gómez, R., Mora-Gómez, J., … Tockner, K. (2011). Preconditioning effects of intermittent stream flow on leaf litter decomposition. Aquatic Sciences, 73(4), 599-609.

Doering, M., Uehlinger, U., Ackermann, T., Woodtli, M., & Tockner, K. (2011). Spatiotemporal heterogeneity of soil and sediment respiration in a riverfloodplain mosaic (Tagliamento, NE Italy). Freshwater Biology, 56(7), 1297-1311.

Harner, M. J., Opitz, N., Geluso, K., Tockner, K., & Rillig, M. C. (2011). Arbuscular mycorrhizal fungi on developing islands within a dynamic river floodplain: an investigation across successional gradients and soil depth. Aquatic Sciences, 73(1), 35-42.

Pahl-Wostl, C., Nilsson, C., Gupta, J., & Tockner, K. (2011). Societal Learning Needed to Face the Water Challenge. AMBIO, 40(5), 549-553.

Perkin, E. K., Hölker, F., Richardson, J. S., Sadler, J. P., Wolter, C., & Tockner, K. (2011). The influence of artificial light on stream and riparian ecosystems: questions, challenges, and perspectives. Ecosphere, 2(11), 122.

Peter, S., Rechsteiner, R., Lehmann, M. F., Tockner, K., Vogt, T., Wehrli, B., & DurischKaiser, E. (2011). Denitrification hot spot and hot moments in a restored riparian system. In M. Schirmer, E. Hoehn, & T. Vogt (Eds.), GQ10: groundwater quality management in a rapidly changing world (pp. 433-436). International Association of Hydrological Sciences. 

Samaritani, E., Shrestha, J., Fournier, B., Frossard, E., Gillet, F., … Tockner, K., … Luster, J. (2011). Heterogeneity of soil carbon pools and fluxes in a channelized and a restored floodplain section (Thur River, Switzerland). Hydrology and Earth System Sciences, 15(6), 1757-1769.

Steward, A. L., Marshall, J. C., Sheldon, F., Harch, B., Choy, S., Bunn, S. E., & Tockner, K. (2011). Terrestrial invertebrates of dry river beds are not simply subsets of riparian assemblages. Aquatic Sciences, 73(4), 551-566.

Tockner, K. (2011). Obituary: Internat. Rev. Hydrobiol. 5/2011. International Review of Hydrobiology, 96(5), 427-434.

Tonolla, D., Lorang, M. S., Heutschi, K., Gotschalk, C. C., & Tockner, K. (2011). Characterization of spatial heterogeneity in underwater soundscapes at the river segment scale. Limnology and Oceanography, 56(6), 2319-2333.

von Schiller, D., Acuña, V., Graeber, D., Martí, E., Ribot, M., Sabater, S., … Tockner, K. (2011). Contraction, fragmentation and expansion dynamics determine nutrient availability in a Mediterranean forest stream. Aquatic Sciences, 73(4), 485-497.


Acuña, V., & Tockner, K. (2010). The effects of alterations in temperature and flow regime on organic carbon dynamics in Mediterranean river networks. Global Change Biology, 16(9), 2638-2650.

Blettler, M., Sukhodolov, A., & Tockner, K. (2010). Hydraulic conditions over bed forms control the benthic fauna distribution in a lowland river (Spree River, Germany). In A. Dittrich, K. Koll, J. Aberle, & P. Geisenhainer (Eds.), Riverflow 2010 (Vol. 2, pp. 1463-1467). Bundesanstalt für Wasserbau.

Hölker, F., Moss, T., Griefahn, B., Kloas, W., Voigt, C. C., Henckel, D., … Tockner, K. (2010). The Dark Side of Light: A Transdisciplinary Research Agenda for Light Pollution. Ecology and Society, 15(4), 13.

Hölker, F., Perkin, E. K., Voigt, C. C., Monaghan, M. T., Wolter, C., & Tockner, K. (2010). Artificial light and the environment. In F. Hölker & K. Tockner (Eds.), Loss of the Night: Transdisciplinary Research on Light Pollution (pp. 14-15). Mediabogen. 

Hölker, F., & Tockner, K. (2010). Loss of the Night. In F. Hölker & K. Tockner (Eds.), Loss of the Night: Transdisciplinary Research on Light Pollution (pp. 4-5). Mediabogen.

Hölker, F., Wolter, C., Perkin, E. K., & Tockner, K. (2010). Light pollution as a biodiversity threat. Trends in Ecology & Evolution, 25(12), 681-682.

Indermaur, L., Schaub, M., Jokela, J., Tockner, K., & Schmidt, B. R. (2010). Differential response to abiotic conditions and predation risk rather than competition avoidance determine breeding site selection by anurans. Ecography, 33(5), 887-

Indermaur, L., Schmidt, B. R., Tockner, K., & Schaub, M. (2010). Spatial variation in abiotic and biotic factors in a floodplain determine anuran body size and growth rate at metamorphosis. Oecologia, 163(3), 637-649.

Larned, S. T., Datry, T., Arscott, D. B., & Tockner, K. (2010). Emerging concepts in temporary-river ecology. Freshwater Biology, 55(4), 717-738.

Milner, A. M., & Tockner, K. (2010). River science – What has it contributed to general ecological theory? River Research and Applications, 26(1), 1-4.

Sabater, S., & Tockner, K. (2009). Effects of Hydrologic Alterations on the Ecological Quality of River Ecosystems. In S. Sabater & D. Barceló (Eds.), Water Scarcity in the Mediterranean: Perspectives Under Global Change (pp. 15-39). Springer

Sommerwerk, N., Bloesch, J., Paunović, M., Baumgartner, C., Venohr, M., SchneiderJacoby, M., … Tockner, K. (2010). Managing the world’s most international river: the Danube River Basin. Marine and Freshwater Research, 61(7), 736-748.

Thieme, M. L., Turak, E., McIntyre, P., Darwall, W., Tockner, K., Cordeiro, J.,& Butchart, S. H. M. (2010). Freshwater Ecosystems Under Threat: The Ultimate Hotspot. In A. Mittermeier, T. A. Farrell, I. J. Harrison, A. J. Upgren, & T. M. Brooks (Eds.),Fresh Water: The Essence of Life (pp. 118-151). Conservation International 

Tockner, K., Lorang, M. S., & Stanford, J. A. (2010). River flood plains are model ecosystems to test general hydrogeomorphic and ecological concepts. River Research and Applications, 26(1), 76-86.

Tockner, K., Pusch, M. T., Borchardt, D., & Lorang, M. S. (2010). Multiple stressors in coupled river-floodplain ecosystems. Freshwater Biology, 55(s1), 135-151.

Tonolla, D., Acuña, V., Lorang, M. S., Heutschi, K., & Tockner, K. (2010). A field-based investigation to examine underwater soundscapes of five common river habitats. Hydrological Processes, 24(22), 3146-3156.

Tonolla, D., Acuña, V., Uehlinger, U., Frank, T., & Tockner, K. (2010). Thermal Heterogeneity in River Floodplains. Ecosystems, 13(5), 727-740.

Yoshimura, C., Fujii, M., Omura, T., & Tockner, K. (2010). Instream release of dissolved organic matter from coarse and fine particulate organic matter of different origins. Biogeochemistry, 100(1), 151-165.


Acuña, V., & Tockner, K. (2009) .Surface-subsurface water exchange rates along alluvial river reaches control the thermal patterns in an Alpine river network. Freshwater Biology, 54(2), 306-320.

Bertoldi, W., Gurnell, A., Surian, N., Tockner, K., Zanoni, L., Ziliani, L., & Zolezzi, G. (2009). Understanding reference processes: linkages between river flows, sediment dynamics and vegetated landforms along the Tagliamento River, Italy. River Research and Applications, 25(5), 501-516.

Diener, S., Roa Gutiérrez, F., Zurbrügg, C., & Tockner, K. (2009). Are larvae of the black soldier fly – Hermetia illucens – a financianlly viable option for organic waste management in Costa Rica? In R. Cossu, L. F. Diaz, & R. Stegmann (Eds.), Proceedings Sardinia 2009, Twelfth International Waste Management and Landfill Symposium. CISA publisher.

Diener, S., Zurbrügg, C., & Tockner, K. (2009). Conversion of organic material by black soldier fly larvae: establishing optimal feeding rates. Waste Management & Research, 27(6), 603-610.

Gurnell, A., Tubino, M., & Tockner, K. (2009). Linkages and feedbacks in highly dynamic alpine fluvial systems. Aquatic Sciences, 71(3), 251-252.

Hölker, F., Voigt, C., Wolter, C., Perkin, E., & Tockner, K. (2009). Der Verlust der Nacht ist auch ein ökologisches Problem. Leibniz Gemeinschaft Zwischenruf, 2, 18-21.

Hölker, F., Wolter, C., Perkin, E., & Tockner, K. (2009). Der Verlust der Nacht. IGBJahresforschungsbericht 2008, 12-13.

Indermaur, L., Gehring, M., Wehrle, W., Tockner, K., & Naef-Daenzer, B. (2009).

Behavior‐Based Scale Definitions for Determining Individual Space Use: Requirements of Two Amphibians. The American Naturalist, 173(1), 60-71.

Indermaur, L., Winzeler, T., Schmidt, B. R., Tockner, K., & Schaub, M. (2009). Differential resource selection within shared habitat types across spatial scales in sympatric toads. Ecology, 90(12), 3430-3444.

Monaghan, M. T., & Tockner, K. (2009). BeGenDiv: Gemeinsam die Artenvielfalt entschlüsseln. IGB-Jahresforschungsbericht 2008, 8-9.

Sommerwerk, N., Baumgartner, C., Bloesch, J., Hein, T., Ostojić, A., Paunović M, … Tockner, K. (2009). The Danube River Basin. Rivers of Europe. In K. Tockner, U. Uehlinger, & C. T. Robinson (Eds.), Rivers of Europe (pp. 59-112). Academic Press. 

Tockner, K., Robinson, C. T., Uehlinger, U. (Eds.). (2009). Rivers of Europe (1st ed.). Academic Press.

Tockner, K., Tonolla, D., Junker, B., & Heutschi, K. (2009). Der Klang der Flüsse. Nationalpark, 143, 20-21.

Tockner, K., Uehlinger, U., & Robinson, C. T. (2009). Preface and Acknowledgements. In K. Tockner, U. Uehlinger, & C. T. Robinson (Eds.), Rivers of Europe (1st ed., pp. xv-xv). Academic Press.

Tockner, K., Uehlinger, U., Robinson, C. T., Siber, R., Tonolla, D., & Peter, F. D. (2009).

European Rivers. In G. E. Likens (Ed.). Encyclopedia of Inland Waters (Vol. 3, pp. 366-377). Academic Press.

Tockner, K., Uehlinger, U., Robinson, C. T., Tonolla, D., Siber, R., & Peter, F. D. (2009). Introduction to European Rivers. In K. Tockner, U. Uehlinger, & C. T. Robinson (Eds.), Rivers of Europe (1st ed., pp. 1-21). Academic Press.

Tockner, K., Wüest, A., & Findlay, S. (2009). Aquatic Sciences celebrates its 20th anniversary. Aquatic Sciences, 71(1), 1-2.

Tonolla, D., Lorang, M. S., Heutschi, K., & Tockner, K. (2009). A flume experiment to examine underwater sound generation by flowing water. Aquatic Sciences, 71(4), 449-462.

Yarushina, M. I., Eremkina, T. V., & Tockner, K. (2009). Ural River Basin. In K. Tockner, U. Uehlinger, & C. T. Robinson (Eds.), Rivers of Europe (1st ed., pp. 673-684). Academic Press.


Acuña, V., Wolf, A., Uehlinger, U., & Tockner, K. (2008). Temperature dependence of stream benthic respiration in an Alpine river network under global warming. Freshwater Biology, 53(10), 2076-2088.

Arrigoni, A., Findlay, S., Fischer, D., & Tockner, K. (2008). Predicting carbon and nutrient transformations in tidal freshwater wetlands of the Hudson River. Ecosystems, 11(5), 790-802.

Döring, M., & Tockner, K. (2008). Morphology and dynamics of riparian zones. In D. Arizpe, A. Mendes, & J. E. Rabaça (Eds.), Sustainable Riparian Zones: A Management Guide (pp. 24-29). Generalitat Valenciana.

Indermaur, L., Schmidt, B. R., & Tockner, K. (2008). Effect of transmitter mass and tracking duration on body mass change of two anuran species. AmphibiaReptilia, 29(2), 263-269.

Langhans, S. D., Tiegs, S. D., Gessner, M. O., & Tockner, K. (2008). Leaf-decomposition heterogeneity across a riverine floodplain mosaic. Aquatic Sciences, 70(3), 337-346.

Paetzold, A., Yoshimura, C., & Tockner, K. (2008). Riparian arthropod responses to flow regulation and river channelization. Journal of Applied Ecology, 45(3), 894-903.

Tockner, K. (2008). Wenn die Nacht zum Tage wird: Lichtverschmutzung – ein stark unterschätztes Problem für Mensch und Natur. Grünblick, 77, 4-5.

Tockner, K., Bunn, S. E., Gordon, C., Naiman, R. J., Quinn, G. P., & Stanford, J. A. (2008). Flood plains: critically threatened ecosystems. In N. V. C. Polunin (Ed.), Aquatic Ecosystems: Trends and Global Prospects (pp. 45-61). Cambridge University Press.

Wantzen, K. M., Yule, C. M., Tockner, K., & Junk, W. J. (2008). Riparian Wetlands of Tropical Streams. In D. Dudgeon (Ed.), Tropical Stream Ecology (pp. 199-217). Academic Press.

Yoshimura, C., Gessner, M. O., Tockner, K., & Furumai, H. (2008). Chemical properties, microbial respiration, and decomposition of coarse and fine particulate organic matter. Journal of the North American Benthological Society, 27(3), 664-673.


Doering, M., Uehlinger, U., Rotach, A., Schlaepfer, D. R., & Tockner, K. (2007). Ecosystem expansion and contraction dynamics along a large Alpine alluvial corridor (Tagliamento River, Northeast Italy). Earth Surface Processes and Landforms, 32(11), 1693-1704.

Paetzold, A., Sabo, J. L., Sadler, J. P., Findlay, S. E., & Tockner, K. (2007). AquaticTerrestrial Subsidies along River Corridors. In P. J. Wood, D. M. Hannah, & J. P.

Sadler (Eds.), Hydroecology and Ecohydrology: Past, Present and Future (pp. 57-73). John Wiley & Sons.

Reichert, P., Borsuk, M., Hostmann, M., Schweizer, S., Spörri, C., Tockner, K., & Truffer, B. (2007). Concepts of decision support for river rehabilitation. Environmental Modelling & Software, 22(2), 188-201.

Tiegs, S. D., Langhans, S. D., Tockner, K., & Gessner, M. O. (2007). Cotton strips as a leaf surrogate to measure decomposition in river floodplain habitats. Journal of the North American Benthological Society, 26(1), 70-77.[70:CSAALS]2.0.CO;2

Tockner, K. (2007). River restoration: linking science with application. Ecology and Civil Engineering, 10(1), 15-25.

Woolsey, S., Capelli, F., Gonser, T., Hoehn, E., Hostmann, M., … Tockner, K., … Peter, (2007). A strategy to assess river restoration success. Freshwater Biology, 52(4), 752-769.


Coops, H., Tockner, K., Amoros, C., Hein, T., & Quinn, G. (2006). Restoring Lateral Connections Between Rivers and Floodplains: Lessons from Rehabilitation Projects. In J. T. A. Verhoeven, B. Beltman, R. Bobbink, & D. F. Whigham (Eds.), Wetlands and Natural Resource Management (pp. 15-32). Springer.

Langhans, S. D., Tiegs, S. D., Uehlinger, U., & Tockner, K. (2006). Environmental heterogeneity controls organic-matter dynamics in river-floodplain ecosystems. Polish Journal of Ecology, 54(4), 675-680.

Langhans, S. D., & Tockner, K. (2006). The role of timing, duration, and frequency of inundation in controlling leaf litter decomposition in a river-floodplain ecosystem (Tagliamento, northeastern Italy). Oecologia, 147(3), 501-509.

Malard, F., Uehlinger, U., Zah, R. and Tockner, K. (2006). Flood-pulse and riverscape dynamics in a braided glacial river. Ecology, 87(3), 704-716.  

Nakamura, K., Amano, K., & Tockner, K. (2006). River restoration: European perspectives and lessons for Japan. Ecology and Civil Engineering, 8(2), 201-214.

Nakamura, K., Tockner, K., & Amano, K. (2006). River and Wetland Restoration: Lessons from Japan. Bioscience, 56(5), 419-429;2

Paetzold, A., Bernet, J. F., & Tockner, K. (2006). Consumer-specific responses to riverine subsidy pulses in a riparian arthropod assemblage. Freshwater Biology, 51(6), 1103-1115.

Tockner, K., Klaus, I., Baumgartner, C., & Ward, J. V. (2006). Amphibian diversity and nestedness in a dynamic floodplain river (Tagliamento, NE-Italy). Hydrobiologia, 565(1), 121-133.

Tockner, K., Paetzold, A., Karaus, U., Claret, C., & Zettel, J. (2006). Ecology of Braided Rivers. In G. H. Sambrook Smith, J. L. Best, C. S. Bristow, & G. E. Petts (Eds.), Braided Rivers: Process, Deposits, Ecology and Management (pp. 339-359). Blackwell.

Trottmann, N., Langhans, S. D., & Tockner, K. (2006). Schwemmgut als Ausbreitungsmedium. Das Innenleben eines unterschätzten Naturstoffs. Wasser, Energie, Luft, 98(2), 153-159.

Yoshimura, C., Tockner, K., Omura, T., & Moog, O. (2006). Species diversity and functional assessment of macroinvertebrate communities in Austrian rivers. Limnology, 7(2), 63-74.


Arscott, D. B., Tockner, K., & Ward, J. V. (2005). Later organization of aquatic invertebrates along the corridor of a braided floodplain river. Journal of the North American Benthological Society, 24(4), 934-954.

Gurnell, A., Tockner, K., Edwards, P. & Petts, G. (2005). Effects of deposited wood on biocomplexity of river corridors. Frontiers in Ecology and the Environment, 3(7), 377-382.;2

Jansson, R., Backx, H., Boulton, A. J., Dixon, M., Dudgeon, D., Hughes, F. M. R., …

Tockner, K. (2005). Stating mechanisms and refining criteria for ecologically successful river restoration: a comment on Palmer et al. (2005). Journal of  Applied Ecology, 42(2), 218-222.

Karaus, U., Alder, L. & Tockner, K. (2005). “Concave islands”: Habitat heterogeneity of parafluvial ponds in a gravel-bed river. Wetlands, 25(1), 26-37.[0026:CIHHOP]2.0.CO;2

Paetzold, A., Schubert, C. J., & Tockner, K. (2005). Aquatic Terrestrial Linkages Along a Braided-River: Riparian Arthropods Feeding on Aquatic Insects. Ecosystems, 8(7), 748-759.

Paetzold, A., & Tockner, K. (2005). Effects of riparian arthropod predation on the biomass and abundance of aquatic insect emergence. Journal of the North American Benthological Society, 24(2), 395-402.

Tockner, K. (2005). Linking Pattern and Process Along River Corridors. In G. E. Petts & B. Kennedy (Eds.), Emerging Concepts for Integrating Human and Environmental Water Needs in River Basin Management (ERDC/EL TR-05-13, pp. 11-17). Environmental Laboratory.

Tockner, K., Surian, N., & Toniutti, N. (2005). Geomorphologie, Ökologie und nachaltiges Management einer Wildflusslandschaft am Beispiel des Fiume Tagliamento (Friaul, Italien) – ein Modellökosystem für den Alpenraum und ein Testfall für die EU-Wasserrahmenrichtlinie. Jahrbuch des Vereins zum Schutz der Bergwelt, 70, 3-17.

Trottmann, N., Langhans, S. D., & Tockner, K. (2005). Schwemmgut, ein wichtiger Weg der Ausbreitung. Natur und Mensch, 47(5), 8-11.

Woolsey, S., Weber, C., Gonser, T., Hoehn, E., Hostmann, M., … Tockner, K., Peter, A. (2005). Handbuch für die Erfolgskontrolle bei Fliessgewässerrevitalisierungen. Eine Publikation des Rhone-Thur Projektes. Eawag, WSL, LCH-EPFL, VAW-ETHZ.

Yoshimura, C., Omura, T., Furumai, H., & Tockner, K. (2005). Present state of rivers and streams in Japan. River Research and Applications, 21(2-3), 93-112.


Kaiser, E., Arscott, D. B., Tockner, K., & Sulzberger, B. (2004). Sources and distribution of organic carbon and nitrogen in the Tagliamento River, Italy. Aquatic Sciences, 66(1), 103-116.

Reichert, P., Borsuk, M., Hostmann, M., Schweizer, S., Spörri, C., Tockner, K., & Truffer, B. (2004). Concepts of Decision Support for River Rehabilitation. In C. Pahl-Wostl, S. Schmidt, A. E. Rizzoli, & A. J. Jakeman (Eds.), Complexity and Integrated Resources Management. Transactions of the 2nd Biennial Meeting of the International Environmental Modelling and Software Society. (Vol. 2, pp. 550-555). International Environmental Modelling and Software Societey.

Robinson, C. T., Tockner, K., & Burgherr, P. (2004). Drift benthos relationships in the seasonal colonization dynamics of alpine streams. Archiv für Hydrobiologie, 160(4), 447-470.

Tockner, K., Karaus, U., Paetzold, A., & Blaser, S. (2004). Ökologischer Zustand der Rhone: Benthische Evertebraten und Uferfauna. Wasser, Energie, Luft, 96(1112), 315-317. 


Arscott, D. B., Keller, B., Tockner, K., & Ward, J. V. (2003). Habitat Structure and Trichoptera Diversity in Two Headwater Flood Plains, N.E. Italy. International Review of Hydrobiology, 88(3-4), 255-273.

Arscott, D. B., Tockner, K., & Ward, J. V. (2003). Spatio-temporal patterns of benthic invertebrates along the continuum of a braided Alpine river. Archiv für Hydrobiologie, 158(4), 431-460

Burgherr, P., Hieber, M., Klein, B., Monaghan, M. T., Robinson, C. T., & Tockner, K. (2003). Biodiversität der Wirbellosenfauna in Fliessgewässern des Val Roseg.

EAWAG news, 54d, 26-27. Burgherr, P., Hieber, M., Klein, B., Monaghan, M. T., Robinson, C. T., & Tockner, K. (2003). La biodiversité du zoobenthos des cours d’eau alpins: Le Val Roseg. EAWAG news, 54f, 26-27.

Burgherr, P., Klein, B., Robinson, C. T., & Tockner, K. (2003). Surface Zoobenthos. In J. V. Ward & U. Uehlinger (Eds.), Ecology of a Glacial Flood Plain (pp. 153-173). Kluwer Academic Publishers.

Fujii, M., Yoshimura, C., Tockner, K., & Omura, T. (2003). Dissolved Organic Matter (DOM) Formation from Particulate Organic Matter (POM) in Upstreams. Environmental Engineering Research, 40, 139-149.

Robinson, C. T., Burgherr, P., Malard, F., Tockner, K., & Uehlinger, U. (2003). Synthesis and Perspectives. In J. V. Ward & U. Uehlinger (Eds.), Ecology of a Glacial Flood Plain (pp. 259-271). Kluwer Academic Publishers.

Tockner, K. (2003). The Tagliamento River: an ecosystem of European importance. FBA News, 22, 6.

Tockner, K. (2003). Der “König der Alpenflüsse” vor seinem Ende? Natur und Mensch, 45(4), 26-27.

Tockner, K. (2003). Totholz – entsorgungspflichtig oder wertvoll? Petri-Heil, 54(3), 43.

Tockner, K., Illi, R., Malard, F., & Uehlinger, U. (2003). Nutrient Dynamics. In J. V. Ward & U. Uehlinger (Eds.), Ecology of a Glacial Flood Plain (pp. 91-107). Kluwer Academic Publishers.

Tockner, K., & Langhans, S. D. (2003). Die ökologische Bedeutung des Schwemmgutes. Wasser, Energie, Luft, 95(11-12), 353-354.

Tockner, K., & Malard F (2003). Channel Typology. In J. V. Ward & U. Uehlinger (Eds.), Ecology of a Glacial Flood Plain (pp. 57-73). Kluwer Academic Publishers. 

Tockner, K., & Peter. A. (2003) Totholz und Schwemmgut – entsorgungspflichtig oder ökologisch wertvoll? Wasser, Energie, Luft, 95(11-12), 351-374.

Tockner, K., Robinson, C. T., & Burgherr, P. (2003). Drift and colonization dynamics. In J. V. Ward & U. Uehlinger (Eds.), Ecology of a Glacial Flood Plain (pp. 243-257). Kluwer Academic Publishers. 

Tockner, K., Uehlinger, U., & Malard, F. (2003). Habitatdynamik in der Schwemmebene des Val Roseg. EAWAG news, 54d, 16-18.

Tockner, K., Uehlinger, U., & Malard, F. (2003). La dynamique des habitats de la plaine alluviale du Val Roseg. EAWAG news, 54f, 16-18.

Tockner, K., Ward, J. V., Arscott, D. B., Edwards, P. J., Kollmann, J., Gurnell, A. M., … Maiolini, B. (2003). The Tagliamento River: A model ecosystem of European importance. Aquatic Sciences, 65(3), 239-253.

Uehlinger, U., Malard, F., & Tockner, K. (2003). Val Roseg: eine glaziale Schwemmebene in den Schweizer Alpen. EAWAG news, 54d, 13-15.

Uehlinger, U., Malard, F., & Tockner, K. (2003). La Val Roseg: Une plaine alluviale glaciaire des Alpes suisses. EAWAG news, 54f, 13-15.

Uehlinger, U., Tockner, K., & Malard, F. (2003). Ökologische Zeitfenster in glazialen Fliessgewässerökosystemen. EAWAG news, 54d, 22-23.

Uehlinger, U., Tockner, K., & Malard, F. (2003). Fenêtres écologiques dans les écosystèmes fluviaux glaciaires. EAWAG news, 54f, 22-23.

van der Nat, D., Tockner, K., Edwards, P. J., & Ward, J. V. (2003). Large wood dynamics of complex Alpine river floodplains. Journal of the North American Benthological Society, 22(1), 35-50. van der Nat, D., Tockner, K., Edwards, P. J., Ward, J. V., & Gurnell, A. M. (2003).

Habitat change in braided flood plains (Tagliamento, NE-Italy). Freshwater Biology, 48(10), 1799-1812.


Arscott, D. B., Glatthaar, R., Tockner, K., & Ward, J. V. (2002). Larval black fly (Diptera: Simuliidae) distribution and diversity along a floodplain river in the Alps (Tagliamento River, Italy). Internationale Vereinigung für theoretische und angewandte Limnologie: Verhandlungen, 28(2), 524-531.

Arscott, D., Tockner, K., van der Nat, D., & Ward, J. V. (2002). Aquatic Habitat Dynamics along a Braided Alpine River Ecosystem (Tagliamento River, Northeast Italy). Ecosystems, 5(8), 802-814.

Burgherr, P., Hieber, M., Klein, B., Monaghan, M. T., Robinson, C. T., & Tockner, K. (2002). Biodiversity of Zoobenthos in Alpine Steams: The Val Roseg. EAWAG news, 54e, 22-23.

Claret, C., Ward, J. V., & Tockner, K. (2002). Temperature heterogeneity of interstitial water in island-associated water bodies of a dynamic flood plain. Internationale Vereinigung für theoretische und angewandte Limnologie: Verhandlungen, 28(1), 345-351.

Malard, F., Hofmann, A., Tockner, K., & Uehlinger, U. (2002). Iron concentration in the water of glacial river system. Internationale Vereinigung für theoretische und angewandte Limnologie: Verhandlungen, 28(1), 134-139.

Malard, F., Tockner, K., Dole-Olivier, M., & Ward, J. V. (2002). A landscape perspective of surface-subsurface hydrological exchanges in river corridors. Freshwater Biology, 47(4), 621-640.

Robinson, C. T., Tockner, K., & Burgherr, P. (2002). Seasonal patterns in macroinvertebrate drift and seston transport in streams of an alpine glacial flood plain. Freshwater Biology, 47(5), 985-993.

Robinson, C. T., Tockner, K., & Ward, J. V. (2002). The fauna of dynamic riverine landscapes. Freshwater Biology, 47(4), 661-677.

Tockner, K. (2002). Ausgedehnte Wildflusslandschaft durch Hochwasserschutz bedroht. Garten + Landschaft, 12, 42.

Tockner, K., Malard, F., Uehlinger, U., & Ward, J. V. (2002). Nutrients and organic matter in a glacial river-floodplain system (Val Roseg, Switzerland). Limnology and Oceanography, 47(1), 266-277. Tockner, K., Paetzold, A., & Karaus, U. (2002). Leben in der Flussdynamik zwischen Trockenfall und Hochwasser. In Bayerische Akademie der Wissenschaften (Ed.), Rundgespräche der Kommission für Ökologie (Vol. 24, pp. 37-46). Pfeil.

Tockner, K., & Peter, A. (2002). Totholz spielt im Ökosystem der Gewässer eine wichtige Rolle. Kommunalmagazin, 19(10), 31.

Tockner, K., & Stanford, J. A. (2002). Riverine flood plains: present state and future trends. Environmental Conservation, 29(3), 308-330.

Tockner, K., Uehlinger, U., & Malard, F. (2002). Habitat Dynamics in the Val Roseg Flood Plain. EAWAG news, 54e, 14-15.

Tockner, K., Ward, J. V., Edwards, P. J., & Kollmann, J. (2002). Riverine landscapes: an introduction. Freshwater Biology, 47(4), 497-500.

Uehlinger, U., Malard, F., & Tockner, K. (2002). Val Roseg: A Glacial Flood Plain in the Swiss Alps. EAWAG news, 54e, 12-13.

Uehlinger, U., Tockner, K., & Burgherr, P. (2002). Vielfalt im Gebirgsbach. Resultate aus dem Val-Roseg-Projekt. Hotspot, 6, 9.

Uehlinger, U., Tockner, K., & Malard, F. (2002). Ecological Windows in Glacial Stream Ecosystems. EAWAG news, 54e, 20-21

van der Nat, D., Schmidt, A. P., Tockner, K., Edwards, P. J., & Ward, J. V. (2002).

Inundation Dynamics in Braided Floodplains: Tagliamento River, Northeast Italy. Ecosystems, 5(7), 636-647.

van der Nat, D., Tockner, K., Edwards, P. J., & Ward, J. V. (2002). Quantification of large woody debris in large floodplain rivers: an area-based approach using differential GPS and GIS. Internationale Vereinigung für theoretische und angewandte Limnologie: Verhandlungen, 28(1), 332-335.

Ward, J. V., Malard, F., & Tockner, K. (2002). Landscape ecology: a framework for integrating pattern and process in river corridors. Landscape Ecology, 17(1), 35-45.

Ward, J. V., Robinson, C. T., & Tockner, K. (2002). Applicability of ecological theory to riverine ecosystems. Internationale Vereinigung für theoretische und angewandte Limnologie: Verhandlungen, 28(1), 443-450.

Ward, J. V., Tockner, K., Arscott, D. B., & Claret, C. (2002). Riverine landscape diversity. Freshwater Biology, 47(4), 517-539


Arscott, D. B., Tockner, K., & Ward, J. V. (2001). Thermal heterogeneity along a braided floodplain river (Tagliamento River, northeastern Italy). Canadian Journal of Fisheries and Aquatic Sciences, 58(12), 2359-2373.

Gurnell, A. M., Petts, G. E., Hannah, D. M., Smith, B. P. G., Edwards, P. J., Kollmann, J., … Tockner, K. (2001). Riparian vegetation and island formation along the gravel-bed Fiume Tagliamento, Italy. Earth Surface Processes and Landforms, 26(1), 31-62.;2Y

Klaus, I., Baumgartner, C., & Tockner, K. (2001). Die Wildflusslandschaft des Tagliamento (Italien, Friaul) als Lebensraum einer artenreichen Amphibiengesellschaft. Zeitschrift für Feldherpetologie, 8, 21-29.

Tockner, K., Ward, J. V., Edwards, P. J., Kollmann, J., Gurnell, A. M., & Petts, G. E. (2001). Der Tagliamento (Norditalien): Eine Wildflussaue als Modellökosystem für den Alpenraum. Laufener Seminarbeiträge, 3, 25-34 

Ward, J. V., & Tockner, K. (2001). Biodiversity: towards a unifying theme for river ecology. Freshwater Biology, 46(6), 807-819.

Ward, J. V., Tockner, K., Uehlinger, U., & Malard, F. (2001). Understanding natural patterns and processes in river corridors as the basis for effective river restoration. Regulated Rivers: Research & Management, 17(4-5), 311-323.


Arscott, D. B., Tockner, K., & Ward, J. V. (2000). Aquatic habitat diversity along the corridor of an alpine floodplain river (Fiume Tagliamento, Italy). Archiv für Hydrobiologie, 149(4), 679-704.

Gurnell, A. M., Petts, G. E., Hannah, D. M., Smith, B. P. G., Edwards, P. J., Kollmann, J., … Tockner, K. (2000). Wood storage within the active zone of a large European gravel-bed river. Geomorphology, 34(1-2), 55-72.

Gurnell, A. M., Petts, G. E., Harris, N., Ward, J. V., Tockner, K., Edwards, P. J., & Kollmann, J. (2000). Large wood retention in river channels: the case of the Fiume Tagliamento, Italy. Earth Surface Processes and Landforms, 25(3), 255275.<255::AIDESP56>3.0.CO;2H

Klein, B., & Tockner, K. (2000). Biodiversity in springbrooks of a glacial flood plain (Val Roseg, Switzerland). Internationale Vereinigung für theoretische und angewandte Limnologie: Verhandlungen, 27(2), 704-710.

Malard, F., Tockner, K., & Ward, J. V. (2000). Physico-chemical heterogeneity in a glacial riverscape. Landscape Ecology, 15(8), 679-695.

Petts, G. E., Gurnell, A. M., Gerrard, A. J., Hannah, D. M., Hansford, B., … Tockner, K., Smith, B. P. G. (2000). Longitudinal variations in exposed riverine sediments: a context for the ecology of the Fiume Tagliamento, Italy. Aquatic Conservation: Marine and Freshwater Ecosystems, 10(4), 249-266.<249::AIDAQC410>3.0.CO;2R

Tockner, K. (2000). Invertebrates in Freshwater Wetlands of North America. Freshwater Biology, 45(1), 103-104.

Tockner, K. (2000). Schweizer Forschung in italienischer Wildflusslandschaft: Der Tagliamento als Modell für Flussrevitalisierungen. CH-Forschung, 7, 1-2.

Tockner, K., Baumgartner, C., Schiemer, F., & Ward, J. V. (2000) Biodiversity of a Danubian floodplain: structural, functional and compositional aspects. In B. Gopal, W. J. Junk, & J. A. Davis (Eds.), Biodiversity in Wetlands: Assessment, Function and Conservation (Vol. 1, pp. 141-159). Backhuys.

Tockner, K., Malard, F., & Ward, J. V. (2000). An extension of the flood pulse concept. Hydrological Processes, 14(16-17), 2861-2883.<2861::AIDHYP124>3.0.CO;2F

Ward, J. V., Tockner, K., Edwards, P., Kollmann, J., Gurnell, A., Petts, G., … Rossaro, B. (2000). Potential role of island dynamics in river ecosystems. Internationale  Vereinigung für theoretische und angewandte Limnologie: Verhandlungen, 27(5), 2582-2585.




Edwards, P. J., Kollmann, J., Tockner, K., & Ward, J. V. (1999). The role of island dynamics in the maintenance of biodiversity in an Alpine river system. Bulletin of the Geobotanical Institute ETH, 65, 73-86 

Kollmann, J., Vieli, M., Edwards, P. J., Tockner, K., & Ward, J. V. (1999). Interactions between vegetation development and island formation in the Alpine river Tagliamento. Applied Vegetation Science, 2(1), 25-36.

Malard, F., Tockner, K., & Ward, J. V. (1999). Shifting Dominance of Subcatchment Water Sources and Flow Paths in a Glacial Floodplain, Val Roseg, Switzerland. Arctic, Antarctic, and Alpine Research, 31(2), 135-150.

Schiemer, F., Baumgartner, C., & Tockner, K. (1999). Restoration of floodplain rivers: the ‘Danube restoration project’. Regulated Rivers: Research & Management, 15(1-3),  231-244.<231::AIDRRR548>3.0.CO;25

Tockner, K., Pennetzdorfer, D., Reiner, N., Schiemer, F., & Ward, J. V. (1999). Hydrological connectivity, and the exchange of organic matter and nutrients in a dynamic river-floodplain system (Danube, Austria). Freshwater Biology, 41(3), 521-535.

Tockner, K., Schiemer, F., Baumgartner, C., Kum, G., Weigand, E., Zweimüller, I., & Ward, J. V. (1999). The Danube restoration project: species diversity patterns across connectivity gradients in the floodplain system. Regulated Rivers: Research & Management, 15(1-3), 245-258.<245::AIDRRR540>3.0.CO;2G

Tockner, K., & Ward, J. V. (1999). Biodiversity along riparian corridors. Large Rivers, 11(3), 293-310.

Ward, J. V., Malard, F., & Tockner, K. (1999). Landscape ecology in integrates pattern and process in river corridors. In J. A. Wiens & M. R. Moss (Eds.), Issues in Landscape Ecology (pp. 97-102). International Association for Landscape Ecology.

Ward, J. V., Malard, F., Tockner, K., & Uehlinger, U. (1999). Influence of ground water on surface water conditions in a glacial flood plain of the Swiss Alps. Hydrological Processes, 13(3), 277-293.<277::AIDHYP738>3.0.CO;2N

Ward, J. V., Tockner, K., Edwards, P., Kollmann, J., Bretschko, G., Gurnell, A., … Rossaro, B. (1999). A reference river system for the Alps: the ‘Fiume Tagliamento’. Regulated Rivers: Research & Management, 15(1-3), 63-75.<63::AIDRRR538>3.0.CO;2F

Ward, J. V., Tockner, K., & Schiemer, F. (1999). Biodiversity of floodplain river ecosystems: ecotones and connectivity. Regulated Rivers: Research &  Management, 15(1-3), 125-139.<125::AIDRRR523>3.0.CO;2E


Tockner, K., Schiemer, F., & Ward, J. V. (1998). Conservation by restoration: the management concept for a river-floodplain system on the Danube River in Austria. Aquatic Conservation: Marine and Freshwater Ecosystems, 8(1), 71-86.<71::AIDAQC265>3.0.CO;2D

Ward, J. V., Burgherr, P., Gessner, M. O. G., Malard, F., Robinson, C. T., Tockner, K., … Zah, R. (1998). The Val Roseg project: habitat heterogeneity and connectivity gradients in a glacial flood-plain system. In K. Kovar, U. Tappeiner, N. E. Peters, & R. G. Craig (Eds.), Hydrology, Water Resources and Ecology in Headwaters (pp. 425-432). International Association of Hydrological Sciences.


Tockner, K., Malard, F., Burgherr, P., Robinson, C. T., Uehlinger, U., Zah, R., & Ward, J. V. (1997). Physico-chemical characterization of channel types in a glacial floodplain ecosystem (Val Roseg, Switzerland). Archiv für Hydrobiologie, 140(4), 433-463.

Tockner, K., & Schiemer, F. (1997). Ecological aspects of the restoration strategy for a river-floodplain system on the Danube River in Austria. Global Ecology and Biogeography Letters, 6(3/4), 321-329.

Tockner, K., & Waringer, J. A. (1997). Measuring Drift during a Receding Flood: Results from an Austrian Mountain Brook (Ritrodat-Lunz). Internationale Revue der gesamten Hydrobiologie und Hydrographie, 82(1), 1-13.


Tockner, K. (1996). Colonization experiments for biomonitoring riparian communities of a large regulated river, the Danube (Austria). Large Rivers, 113(1-4), 433-442.

Tockner, K., & Bretschko, G. (1996). Spatial distribution of particulate organic matter (POM) and benthic invertebrates in a river-floodplain transect (Danube, Austria): importance of hydrological connectivity. Large Rivers, 11(1), 11-27.

Ozeane im Klimawandel
Prof. Dr.  Angelika Brandt
Head of Marine Zoology division, Senckenberg Research Institute and Natural History Museum Frankfurt


Die Tiefsee ist der größte Lebensraum der Welt, jedoch gehört sie zu den am wenigsten erforschten Regionen. Unsere bisherigen Erkenntnisse dokumentieren jedoch eine sehr hohe Artenvielfalt am Meeresboden der Tiefsee. Ich versuche daher mit meiner Arbeitsgruppe die Artenvielfalt der Tiefsee zu erfassen, die Verbreitung der Organismen zu analysieren sowie ihre Ökologie und Evolution zu erforschen.

In meiner Arbeitsgruppe wird hauptsächlich an Krebsen (insbesondere Meeresasseln, Isopoda) als Modellorganismen gearbeitet, da diese Tiergruppe in der Tiefsee sehr häufig ist und gute Vergleichsuntersuchungen vorliegen. Wir analysieren die stammesgeschichtliche Herkunft und Besiedlungsgeschichte der Isopoden in der Tiefsee (wie z. B. Sub- und Emergenzphänomne im Südpolarmer), wir versuchen zu analysieren welches in der Tiefsee die treibenden Faktoren für die hohe Diversität sind und bearbeiten Artbildung, Konkurrenz und Koexistenz in Raum und Zeit.

Unsere aktuellen Projekte thematisieren die Besiedlungsmuster in der Tiefsee sowie ihre ökologischen und evolutionsbiologischen Hintergründe sowie die mögliche zukünftige Entwicklung der Biodiversität.


Aktuelle Forschungsprojekte

  • Kuril Kamchatka Biodiversity Studies II (KuramBio II)
  • Beneficial – Biogeography of the northwest Pacific fauna. A benchmark study for estimations of alien invasions into the Arctic Ocean in times of rapid climate chance.
  • Larsen-C PEARL – The influence of Larsen-C ice-cover on macrobenthic peracarid crustacean assemblages on the Antarctic shelf. (Comparisons of the Larsen-C assemblages with those from the seasonally ice-covered Filchner Trough and the ice free South Orkney Islands)
  • Deep-MiPoll – The role of hadal zones in the long-term fate of marine microplastics: Identification of microplastics in the deep sea of the Kuril-Kamchatka Trench, Northwest Pacific.
  • Ausschluss der bodenberührenden Fischerei in marinen Schutzgebieten am Sylter Außenriff, Borkum Riffgrund und Doggerbank Entenschnabel: Zustandsbeschreibung der Bodensedimente und benthopelagischen Habitate. Teilprojekt 4: Raum-Zeitliche Variabilität der Endo- und Epifaunagemeinschaften in den ausgewiesenen Meeresschutzgebieten der Nordsee.


Themen für Abschlussarbeiten

Themen im Zusammenhang mit der Taxonomie und Systematik, Ökologie, Biogeographie und Evolution von Isopoda und/oder Marofaunaorganismen sowie den oben genannten Forschungsprojekten können im Rahmen von Bachelor- und Masterarbeiten sowie Praktika in unserer Sektion und Abteilung erforscht werden.

Beispiele für erfolgreich abgeschlossene und im Anschluss publizierte Bachelorarbeiten:

  • Kniesz K, Brandt A, Riehl T (2018) Peritrich ciliate epibionts on the new hadal isopod species Macrostylis marionae from the Puerto Rico Trench as an indicator for sex-specific behaviour. Deep-Sea Res Pt II 148:105–129.
  • Brandt, A., Alalykina, I., Brix, S., Brenke, N., Błażewicz, M., Golovan, O., Heitland, N., Hrinko, A.M., Jażdżewska, A.M., Jeskulke, K., Kamenev, G., Lavrenteva, A., Malyutina, M., Riehl, T., Lins, L. (2019): Depth zonation of Northwest Pacific deep-sea macrofauna. Progress in Oceanography 176 (2019), pp 1-10102131;

Beispiele für erfolgreich abgeschlossene und im Anschluss publizierte Masterarbeiten:

  • Bober S, Riehl T, Brandt A (2018) An organ of equilibrium in deep-sea isopods revealed: the statocyst of Macrostylidae (Crustacea, Peracarida, Janiroidea). Zoomorphology 137:71–82.

Beispiele für erfolgreich abgeschlossene und im Anschluss publizierte Doktorarbeiten:

  • Schmidt, C., Wolf, K.H.E., Lins, L., Martinez, P., Brandt, A., (2017 online, 2018): Abundance and community pattern of meiofauna in the Vema Fracture Zone and in the Puerto Rico trench. Deep-Sea Research II 148: 236-250.
  • Bober, S., Brix, S., Riehl, T., Schwentner, M., Brandt, A. (2018): Does the Mid Atlantic Ridge affect the distribution of benthic crustaceans across the Atlantic Ocean? Deep-Sea Research II 148: 91-104.
  • Kohlbach, D. et al., Brandt, A. (2018): Dependency of Antarctic zooplankton species on ice algae-produced carbon suggests sea ice-driven ecosystem during winter. Global Change Biology; 1-15. DOI: 10.1111/gcb.14392
  • Guggolz, T., Meißner, K., Schwentner, M., Brandt, A. (2019): Diversity and distribution of Laonice species (Annelida: Spionidae) in the equatorial North Atlantic and Puerto Rico Trench. Nature Scientific Reports: 2019, 9:9260 |

Beispiele für erfolgreiche PostDoktorand*innen in der Arbeitsgruppe:

  • Saeedi, H., Costello, M.J., Warren, D., Brandt, A. (2019): Latitudinal and bathymetrical species richness in the NW Pacific and Arctic Ocean. Nature Scientific Reports, 2019; 9:9303, 1-10,|
  • Saeedi, H., Simões, M., Brandt, A. (2019): Endemicity and community composition of marine species along the NW Pacific and adjacent Arctic Ocean. PROOCE 178, 1-11;
  • Saeedi, H., Simões, M., Brandt, A. (2020): Biodiversity and distribution patterns of deep-sea fauna along the temperate NW Pacific. Progress in Oceanography 183, 1-12;
  • Lins, L., Brandt, A. (2020): Comparability between box-corer and epibenthic-sledge data on higher taxon level: A case study based on deep-sea samples from the NW Pacific. PROOCE. Progress in Oceanography 182 (2020) 102273
  • Riehl, T., Lins, L., Brandt, A. (2017 online, 2018): The effects of depth, distance, and the Mid-Atlantic Ridge on genetic differentiation of abyssal and hadal isopods (Macrostylidae). Deep-Sea Research II 148: 74-90.
  • Riehl, T, Brandão, S.N., Brandt, A. (2019): Chapter 6. Conquering the Ocean Depths Over Three Geological Eras. In G. Poore and M. Thiel, editors. The Natural History of the Crustacea, Volume 8: Evolution and Biogeography. Oxford University Press, New York, USA. In Preparation to press.



Ich unterrichte ich im Fachbereich Biowissenschaften am Institut für Ökologie und Evolution an der Goethe-Universität Frankfurt.

Im Bachelorstudiengang Biowissenschaften betreue ich einen Kurs im Bachelormodul 6b „Diversität der Organismen: Zoologie“. Im Rahmen von Exkursionen, Vorlesungen und eines Praktikums behandle ich die Systematik und Anatomie ausgewählter Tierstämme und unterrichte über die Bedeutung naturwissenschaftlicher Sammlungen und Forschungsmuseen.

Für das Mastermodul Evo-11 „Integrative Biodiversitätsforschung in der Zoologie“ habe ich mit Herrn Dr. Riehl und Kolleg*innen einen interaktiven und sehr praxisorientierten Kursteil entworfen, der innerhalb von zwei Wochen alle wichtigen Arbeitsschritte der taxonomisch-systematischen Erforschung von Tiefseemakrofauna vermittelt. Die Teilnehmer sortieren echte Tiefseeproben, entdecken und bestimmen Organismen daraus und erlernen dann über die Praxis das taxonomische Beschreiben einer Art, das Charakterisieren und kodieren von morphologischen Merkmalen sowie das Erstellen von Bestimmungsschlüsseln.


Beschriebene Taxa

(Unterordnungen: 2; Familien: 2; Gattungen: 18; Arten: 132), siehe PDF.

Beschriebene Arten nach Angelika Brandt: 19, siehe PDF

Kurzer Lebenslauf

Prof. Dr. Angelika Brandt wurde in Minden Westfalen geboren und studierte Biologie und Pädagogik an der Universität Oldenburg (MSc 1987, PhD 1992). 1995 wurde sie ordentliche Professorin an der Universität Hamburg am Zoologischen Museum und Kuratorin der Abteilung Wirbellose II (Krustentiere und Polychaeta). Seit 2003 war sie entweder stellvertretende Direktorin oder Direktorin (2004-2009) des Zoologischen Museums der Universität Hamburg. Angelika Brandt arbeitet an der Makrofauna der Tiefsee und der Polarregionen und hat an 29 Expeditionen teilgenommen. Sie beschäftigt sich mit Systematik, Biodiversität, Biogeographie, Evolution und Ökologie der Tiefsee-Isopoda (Crustacea, Malacostraca). Die Abteilung für Marine Zoologie und die Abteilung Krustentiere des Senckenberg-Forschungsinstituts und Naturmuseums wird seit April 2017 von Angelika Brandt geleitet, die seit ihrer Mitgliedschaft im Stiftungsausschuss des Deutschen Zentrums für marine Biodiversität in Wilhelmshaven und Hamburg (DZMB) mit Senckenberg-zusammenarbeitet. Ihre Arbeit konzentriert sich auf die Systematik, Ökologie und Evolution von perakariden Krebstieren mit Schwerpunkt auf Isopoda aus Tiefsee- und Polarlebensräumen. Im Rahmen der Volkszählung des Meeresorganismen (CoML) war sie an den Antarktis- und Tiefseefeldprojekten CAML (Vielfalt des Meereslebens in der Antarktis) und CeDAMar (Vielfalt des Meereslebens im Abyssal) beteiligt. Ihre Arbeit dokumentierte die hohe Artenvielfalt von Tiefsee-Wirbellosen im Südpolarmeer. In den letzten Jahren konzentrierte sich ihr Forschungsinteresse auf die Identifizierung von Evolutionstreibern in der Tiefsee wie Ausbreitungsbarrieren sowie auf die Weiterentwicklung der Methoden zur Probenahme in der Tiefsee. Sie ist in der Tiefseeforschung tätig (derzeit im Nordwestpazifik und im Südpolarmeer), interessiert sich aber auch für die Artenvielfalt der Weltmeere. Senckenberg hat auch einen starken traditionellen marinen Fokus auf das Rote und Arabische Meer sowie die Nordsee und die Ostsee Meer.


Curriculum vitae – Prof. Dr. Angelika Brandt

Berufliche Beschäftigung und akademische Ausbildung

04.2017 –           Professorin für Marine Zoologie am Senckenberg Forschungsinstitut und Naturmuseum sowie an der Goethe-Universität Frankfurt

04.2017 –           Abteilungsleiterin „Marine Zoologie“ am Senckenberg Forschungsinstitut und Naturhistorisches Museum

12.1995 – 31.3.2017       Professor für Zoologie an der Universität Hamburg und Kurator für Krustentiere und Polychaeta

11.2009 – 31.3.2017       Stellvertretender Leiterin des Zoologischen Museums der Universität Hamburg.

09.2004 – 11.2009          Leiterin des Zoologischen Museums der Universität Hamburg

01.1989 – 09.2012          29 Expeditionen mit Forschungsschiffen in die Arktis, den Süden, den Pazifik und den Atlantik sowie das Japanische Meer und den Kuril-Kamtschatka-Graben und das angrenzende Abyssal

01.1992 – 11.1995          Wissenschaftlicher Mitarbeiter im Sonderforschungsbereich 313, Arbeit im Nordatlantik zum Partikelfluss und zur Gemeinschaftsstruktur vor dem Hintergrund vom Umweltveränderungen.

03.1987 – 12.1990          Wissenschaftlicher Mitarbeiter an der Universität Oldenburg (Doktorand)

06.1989 –  11.1989         Wissenschaftlicher Mitarbeiter an der Scripps Institution of Oceanography, La Jolla, USA


Betreute Wissenschaftliche Arbeiten:

37 Diploma-, 21 Bachelor-, 19 Master- und 28 Doktoranden (4 davon laufend)

– Die Unterstützung junger Forscherinnen ist mir wichtig. Die meisten meiner ehemaligen Studenten haben sich international etabliert: z.B.

– Dr. Katrin Linse (leitende Wissenschaftlerin, BAS, British Antarctic Survey, Cambridge, Großbritannien);

– Dr. Anne-Nina Lörz (leitende Wissenschaftlerin, NIWA-Forschungsinstitut, Wellington, Neuseeland), jetzt wieder in Deutschland am Zentrum für Naturkunde (CeNak)

– Dr. Stefanie Kaiser (leitende Wissenschaftlerin, NOC, Nationales Institut für Ozeanographie, Southampton, Großbritannien), jetzt wieder in Deutschland

 Dr. Saskia Brix (leitende Wissenschaftlerin, DZMB, Senckenberg)

– Prof. Dr. Jürgen Guerrero Kommritz (Professor, Universität Kolumbien, Bogota), jetzt in Santa Martha

– Vier meiner Studenten (Dr. Katrin Linse und Dr. Stefanie Kaiser und Sarah Schnurr MSc) erhielten 2001, 2010 und 2014 den BMBF-Preis „Annette-Barthelt-Preis für Meereswissenschaften“ und meine Mentee Dr. Agnes Weiner, die arbeitet an Foraminiferen im Jahr 2015.


Ausgewählte Auszeichnungen

  • Anerkennung als herausragende Antarktisforscherin (Wikipedia-Biografie)
  • Mitglied der Akademie der Wissenschaften und Literatur Mainz im Jahr 2012
  • Verleihung der SCAR-Medaille für herausragende Leistungen in den Polarwissenschaften im Juli 2008 in St. Petersburg
  • Die ANDEEP-Veröffentlichung (Nature, 2007) wurde vom TIME MAGAZINE als „eines der 10 wichtigsten wissenschaftlichen Ergebnisse des Jahres 2007“ bezeichnet
  • Von der National Geographic Society als Abenteurer des Jahres 2007 ausgezeichnet
  • Mitglied der Linnean Society seit 2003
  • Preise für Diplom und Doktorarbeit („Annette-Barthelt-Preis für Meereswissenschaften“)
  • Gastgeberin von zwei Post-Docs der Alexander von Humboldt-Stiftung


Ausgewählte Gremienarbeit und Leistungen

  • Expertenteamleiter des Teams „Clean Ocean“ für das UN-Jahrzehnt zum Auftakttreffen zu Meereswissenschaften und Nachhaltigkeit in Berlin 29.5.-2.6.2021
  • Referent für das Forschungsfeld „Systematik und Taxonomie“ in Senckenberg
  • Lenkungsausschuss von SOOS (Southern Ocean Observing System) (2011-2015)
  • Vizepräsident der Deutschen Gesellschaft für Polarforschung (DGP) (2010-2015)
  • Beirat des ESF ERICOM-RV Aurora Borealis Science (ESAP) Mitglied (2009 – 2012)
  • Vizepräsident der Deutschen Gesellschaft für Meeresforschung (DGM) (2009 -)
  • Deutsche Kommission für das Internationale Polarjahr (2003 -2009)
  • Redaktionen, z.B. Zeitschrift für marine Biodiversität (Springer) (2008 -)
  • Vorsitzender der Kooperation der Wissenschaftssammlungen Hamburg (VNSH) (2008 -2014)
  • Vorsitzender der NORe-Zusammenarbeit (Museen der Nord- und Ostseeregion) (2008 -2016)
  • Mitglied des Lenkungsausschusses vieler internationaler Programme (z. B. CeDAMar (Volkszählung der Vielfalt des abyssalen Meeresorganismen) und CAML (Volkszählung der antarktischen Meeresorganismen), sowie SOOS (Southern Ocean Observing System)
  • LA-SCAR (Wissenschaftlicher Ausschuss für Antarktisforschung) (2001-2009)
  • Mitglied des Lenkungsausschusses der SCAR-EBA (Evolution und Biodiversität antarktischer Organismen) in der LSSSG (Life Sciences Scientific Standing Group) (2004 – 2010)
  • Fachkommission der Umweltbehörde des Ministeriums für Naturschutz und nukleare Sicherheit (2006 – 2012)
  • Wissenschaftlicher Beirat – Deutsche Fischereiforschungsinstitution (2004 – 2008)
  • Beirat für die Schiffszeit von RV Meteor, Alkor usw. (2002 – 2012)
  • Beirat für die Schiffszeit von RV Polarstern (2007 – 2014)
  • Verschiedene Aufträge der Universität Hamburg (1995 – 2017)
  • 1999 Gründungsausschuss des DZMB (Deutsches Zentrum für marine Biodiversität)
  • 2002 Gründungsmitglied von CeDAMar (einem internationalen Konsortium für die Zählung der biologischen Vielfalt in der Tiefsee)
  • -16.5.2012 Bewertung des Zoologischen Museums von Kopenhagen (Wissenschaftlicher Beirat)
  • 2012 – Mitglied der Akademie der Wissenschaften und der Literatur Mainz

2014 –NORe wird Mitglied von Cetaf

2014/2015 – Jungfernexpedition des neuen deutschen Tiefsee-Forschungsschiffes Sonne.



Zusammenfassung der Veröffentlichung

– Publikationen begutachtet 273; Bücher: 8

– Journal Sonderbände hreausgegeben: 9 in (DSR II), +1 Zootaxa + 1 Journal of Marine Systems and Engineering + 1 Progress in Oceanography

– Unbegutachtete Artikel, kurze Forschungsarbeiten und Notizen: 118

– Online-Datenbanken: 6

– Wissenschaftliche Filme: 1


10 ausgewählte begutachtete Publikationen

Brandt, A., Alalykina, I., Brix, S., Brenke, N., Błażewicz, M., Golovan, O., Heitland, N., Hrinko, A.M., Jażdżewska, A.M., Jeskulke, K., Kamenev, G., Lavrenteva, A., Malyutina, M., Riehl, T., Lins, L. (2019): Depth zonation of Northwest Pacific deep-sea macrofauna. Progress in Oceanography 176 (2019), pp 1-10102131;

Saeedi, H., Costello, M.J., Warren, D., Brandt, A. (2019): Latitudinal and bathymetrical species richness in the NW Pacific and Arctic Ocean. Nature Scientific Reports, 2019; 9:9303, 1-10,|

Johannsen, N., Lins, L., Riehl, T., Brandt, A. (2019): Changes in species composition of Haploniscidae (Crustacea: Isopoda) across potential barriers to dispersal in the Northwest Pacific. Progress in Oc4eanography

Kohlbach, D. et al., Brandt, A. (2018): Dependency of Antarctic zooplankton species on ice algae-produced carbon suggests sea ice-driven ecosystem during winter. Global Change Biology; 1-15. DOI: 10.1111/gcb.14392

Bober, S., Brix, S., Riehl, T., Schwentner, M., Brandt, A. (2018): Does the Mid Atlantic Ridge affect the distribution of benthic crustaceans across the Atlantic Ocean? Deep-Sea Research II 148: 91-104.

Brandt, A., Linse, K., Ellingsen, K., Somerfield, P. (2016): Depth-related gradients in community structure and relatedness of bivalves and isopods in the Southern Ocean? Progress in Oceanography 144: 25–38, doi:

Kennicutt II M., Chown S.L., Cassano J., Liggett D., Massom R., Peck L., Rintoul S., Storey J., Vaughan D., Wilson T., Allison I., Ayton J., Badhe R., Baeseman J., Barrett P., Bell R., Bertler N., Bo S., Brandt A., Bromwich D., Cary C., Clark M., Convey P., Costa E.S., Cowan D., Deconto R., Dunbar R., Elfring C., Escutia C., Francis J., Fricker H., Fukuchi M., Gilbert N., Gutt J., Havermans C., Hik D., Hosie G., Jones C., Kim Y., Le Maho Y., Lee S.H., Leppe M., Leichenkov G., Li X., Lipenkov V., Lochte K., López-Martinez J., Lüdecke C., Lyons W., Marenssi S., Miller H., Morozova P., Naish T., Nayak S., Ravindra R., Retamales J., Ricci C., Rogan-Finnemore M., Ropert-Coudert Y., Samah A.A., Sanson L., Scambos T., Schloss I., Shiraishi K., Siegert M.J., Simões J., Sparrow M., Storey B., Wall D., Walsh J., Wilson G., Winther J.G., Xavier J., Yang H., Sutherland W.J. (2014). Six priorities for Antarctic science. Nature 512, 23-25. (2014): Six priorities for Antarctic science. Nature comment 521: 23-25, suppl. 7 pp.

Connolly, S.R., MacNeil, M.A., Caley, M.J., Cripps, E., Hisano, M., Thibaut, L.M.,  Bhattacharya, B.D., Benedetti-Cecchi, L., Brainard, R.E., Brandt, A., Bulleri, F., Kaiser, S., Knowlton, N., Kroncke, I., Linse, K., Maggi, E., O’Hara, T.D., Plaisance, L., Poore, G.C.B, Sarkar, S.K., Satpathy, K.K., Schückel, U., Sogin, L.M., Stocks, K.I., Williams, A., Wilson, R.S., Zettler, L.A. (2014): Commonness and rarity in the marine biosphere. Proceedings of the National Academy of Sciences 111: 8524-8529.

Constable, A.J., Melbourne-Thomas, J., Corney, S.P., Arrigo, K.R., Barbraud, C., Barnes, D.K.A., Bindoff, N.L., Boyd, P.W., Brandt, A., Costa, D.P., Davidson, A.T., Ducklow, H.W., Emmerson, L., Fukuchi, M., Gutt, J., Hindell, M.A., Hofman, H.E., Hosie, G.W., Iida, T., Jacob, S., Johnston, N.M., Kawaguchi, S., Koubbi, P., Lea, M.-A., Makhado, A., Massom, R.A., Meiners, K., Meredith, M.P., Murphy, E.J., Nicol, S., Richerson, K., Riddle, M.J., Rintoul, S.R., Smith Jr., W.O., Southwell, C., Stark, J.S., Sumner, M., Swadling, K.M., Takahashi, K.T., Trathan, P.N., Welsford, D.C., Weimerskirch, H., Westwood, K.J., Wienecke, B.C., Wolf-Gladrow, D., Wright, S.W., Xavier, J.C., Ziegler, P. (2014): Climate change and Southern Ocean ecosystems I: How changes in physical habitats directly affect marine biota. Global Change Biology, 1-22, doi: 10.1111/gcb.12623.

Brandt A, Gooday AJ, Brix SB, Brökeland W, Cedhagen T, Choudhury M, Cornelius N, Danis B, De Mesel I, Diaz RJ, Gillan DC, Ebbe B, Howe J, Janussen D, Kaiser S, Linse K, Malyutina M, Brandao S, Pawlowski J, Raupach M (2007). The Southern Ocean deep sea: first insights into biodiversity and biogeography. Nature 447, 307-311.


Editierte Wissenschaftliche Sonderbände

Brandt, A. & Hilbig, B. (2004): ANDEEP (ANtarctic benthic DEEP-sea biodiversity: colonization history and recent community patterns) – a tribute to Howard L. Sanders. Deep-Sea Research II, 51 (14-16): 1457-1919.

Brandt, A. & Ebbe, B. (2007): ANDEEP III ANtarctic benthic DEEP-sea biodiversity: colonisation history and recent community patterns. Deep-Sea Research II, 54(16-17): 1645-1904.

Brandt, A. & Ebbe, B. (2011): Southern Ocean biodiversity – from pelagic processes to deep-sea response. Deep-Sea Research II (19-20): 1945-2050.

Malyutina, M.  & Brandt, A. (2013): SoJaBio (Sea of Japan Biodiversity study). Deep-Sea Research II (guest editors) 86-87: Pages 1-238.

Schüller, M., Brandt, A., Ebbe, B. (2013): Diversity of Southern Ocean deep-sea benthos between cosmopolitanism and crytic speciation: new species from the ANDEEP expeditions. Zootaxa 3692: 1-258.

Brandt, A., Würzberg, L. (2014): Southern Ocean deep sea – a benthic view to pelagic processes. DSR II: 108: 1-112.

Brandt A., Malyutina, M.V. (eds.)(2015): The German-Russian deep-sea expedition KuramBio (Kurile Kamchatka Biodiversity Studies) to the abyssal area of the Kuril-Kamchatka Trench on board of the RV Sonne in 2012 following the footsteps of the legendary expeditions with RV Vityaz. Deep-Sea Research II 111: 1-405.

Brandt, A. (2016): “Research Tools and Methods for Marine Species Acquisition and Identification”. Journal of Marine Science and Engineering.

Brandt A., Kaiser, S., Riehl, T. (eds.) (2018): bathymetry of the Vema-Fracture Zone and Puerto Rico TRench and Abyssal AtlaNtic BiodiverSITy Study (Vema-TRANSIT). Deep-Sea Research II

Malyutina, M.V., Chernyshev, A.V., Brandt, A. (eds.) (2018): SokhoBio (Sea of Okhotsk Biodiversity Studies). Deep-Sea Research II 154: 1-382.

Brandt A., Brix, S., Malyutina, M.V., Riehl, T. (eds.) (2018-2020): Biodiversity and biogeography of the abyssal and hadal Kuril-Kamchatka Trench and adjacent NW Pacific deep-sea regions. Progress in Oceanography.


Expeditionen gesamt 29, hier gelistet nur ab 2012

2012      ANT XXVIII/3 Expedition RV „Polarstern“ (SYSTCO II) Southern Polar Front (1.2012-3.2012)

2012      Expedition KuramBio (Kurile-Kachatka Biodiversity Studies) RV „Sonne“ (7.2012-9.2012) (expedition leader A. Brandt)

2014      Expedition Vema-TRANSIT “Bathymetry of the Vema-Fracture Zone and Puerto Rico TRench and Abyssal AtlaNtic BiodiverSITy Study (TRANSIT)” RV „Sonne“ (15.12.2014-16.1.2015)

2015      Expedition RV „Akademik Lavrentyev“ Sea of Ochotsk (6.-7.2015)

2016      Expedition RV „James Clarke Ross“(JR15005), to the South Orkney Islands (25.2.-25.3.2016)

2016      Expedition KuramBio II (Kurile-Kamchatka Biodiversity Studies) RV „Sonne“ to the Kuril-Kamchatka Trench 16.8.-26.9.2016 (expedition leader A. Brandt)

2018 Expedition with RV “James Clarke Ross” (JR17003a) to Larsen-C Ice shelf, western Weddell Sea (21.2.-15.3.2018)

2019 Expedition with FS “Polarstern” (PS118) to the Larsen-C ice shelf, western Weddell Sea (9.2.-10.4.2019)

Alle Publikationen

Martin Mittelbach
Dr.  Martin Mittelbach
Administrative Director of Senckenberg – Leibniz Institution for Biodiversity and Earth System Research
Administrativer Direktor der Senckenberg Gesellschaft für Naturforschung
Prof. Dr. Andreas Mulch
Director of Senckenberg Research Institute and Natural History Museum Frankfurt

The Mulch lab at the Senckenberg Biodiversity and Climate Research Centre and Goethe University Frankfurt uses state-of-the-art isotope geochemical approaches including novel applications of clumped isotope thermometry and molecular biomarker analysis to track changes in temperature and precipitation based on geologic materials such as paleosoils, fossil teeth or minerals in the Earth’s crust that store information about near surface meteoric waters in active deformation zones of our planet.

Collectively, these data permit reconstructions of the change in elevation of mountain belts through time and their role in guiding atmospheric moisture transport over continents or provide cornerstones in establishing the interactions of biotic and abiotic drivers of speciation and evolution in the highly diverse mountainous regions of e.g. the Andes and the Himalayas.

Research interests

  • Stable isotope paleoaltimetry
  • Interactions of mountains, climate and biodiversity
  • Paleoclimate reconstructions

Google Scholar

2015 – Present Director, Senckenberg Research Institute and Natural History Museum Frankfurt
2013 – 2020 Vice-Director General, Senckenberg Gesellschaft für Naturforschung
2010 – Present Professor (W3), Goethe-Universität Frankfurt & Senckenberg Biodiversity and Climate Research Centre
2014 – 2015 Cox Visiting Professor, School of Earth Sciences, Stanford University, USA
2010 – 2017 Vice-Director, Senckenberg Biodiversity and Climate Research Centre
2007 – 2010 Director, Institute of Geology, Leibniz Universität Hannover
2006 – 2010 Professor (W2, tenured), Leibniz Universität Hannover
2005 – 2006 Research Associate, Stanford University, USA
2004 – 2005 US-NSF Research Associate, University of Minnesota, Minneapolis, USA
2004 Dr. ès sci. in Mineralogy/Geochemistry, Universite de Lausanne, Switzerland
1999 Diploma in Geology (Tectonics), Justus-Liebig Universität Gießen


*advised or **co-advised student; underlined = advised postdoctoral scholar


Licht, A., Dupont-Nivet, G., Meijer, N., Caves-Rugenstein, J., Schauer, A., Fiebig, J., Mulch, A., Hoorn, C., Barbolini, N., Guo, Z. (2020.) Decline of soil respiration in northeastern Tibet through the transition into the Oligocene icehouse. Paleogeography, Paleoclimatology, Paleoecology, 560, 110016.

 Methner, K., Campani, M., Fiebig, J., *Löffler, N., Kempf, O., Mulch, A. (2020) Middle Miocene long-term continental temperature change in and out of pace with marine climate records. nature Scientific Reports 10: 7989.

Vasiliev, I., Feurdean, A., Reichart, G-J., Mulch, A. (2020) Late Miocene intensification of continentality in eastern Eurasia (Black Sea basin). Int J Earth Sciences

Pingel, H., Strecker, M.R., Mulch, A., Alonso, R.A., Cottle, J., Rohrmann, A. (2020) Late Cenozoic topographic evolution of the Eastern Cordillera and Puna Plateau margin (NW Argentina). Earth and Planetary Science Letters

Botsyun, S., Ehlers, T.A., Mutz, S.G., Methner, K., *Krsnik, E., Mulch, A. (2020) Opportunities and challenges for paleoaltimetry in “small” orogens: Insights from the European Alps. Geophys Res Letters, 47, e2019GL086046.

Meijers, M.J.M., Brocard, G., Whitney, D.L., Mulch, A. (2020) Paleoenvironmental conditions and drainage evolution of the central Anatolian lake system (Turkey) during late Miocene to Pliocene surface uplift. Geosphere, 16.



*Ardenghi, N., Mulch, A., Koutsodendris, A., Pross, J., Kahmen, A., Niedermeyer, E. (2019) Temperature and moisture variability in the Eastern Mediterranean region during Marine Isotope Stages 11–10 based on biomarker analysis of the Tenaghi Philippon peat deposit. Quaternary Science Reviews, 225, 105977.

Methner, K., Lenz, O., Riegel, W., Wilde, V., Mulch, A. (2019) Palaeoenvironmental response of mid-latitudinal wetlands to PETM climate change (Schöningen lignite deposits, Germany). Climate of the Past, 15, 1-15.

Heineke, C., Hetzel, R., Nilius, N.-P., Zwingmann, H., Todd, A., Mulch, A., Wölfler, A., Glotzbach, C., Akal, C., Dunkl, I., Raven, M., Hampel, A. (2019) Detachment faulting in a bivergent core complex constrained by fault gouge dating and low-temperature thermochronology. Journal of Structural Geology.

Fiebig, J., Bajnaj, D., *Löffler, N., Methner, K., *Krsnik, E., Mulch, A., Hofmann, S. (2019) Combined high-precision D47 and D48 analysis of carbonates. Chemical Geology, 522, 186-191.

Huang, S.1, Meijers, M.J.M.1, Eyres, A., Mulch, A., Fritz, S.A. (2019) Unravelling the history of biodiversity in mountain ranges through integrating geology and biogeography. Journal of Biogeography. Alexander von Humboldt Special Volume. 1= joint 1st authors

Dusseaux, C., Gébelin, A., Boulvais, P., Gardien, V., Grimes, S., Mulch, A. (2019) Meteoric fluid-rock interaction in Variscan shear zones. Terra Nova 2019;00:1–7.

*Löffler, N., Fiebig, J., Mulch, A., Tütken, T., Schmidt, B.C., Bajnai, D., Conrad, A.C., Wacker, U., Böttcher, M.E. (2019) Refining the temperature dependence of the oxygen and clumped isotopic compositions of structurally bound carbonate in apatite. Geochimica et Cosmochimica Acta, 253, 19-38.

Lynch, E.A., Mulch, A., Yonkee, A., van der Pluijm, B. (2019) Hydrogen isotopes in authigenic clay minerals suggest presence of surface fluids in the evolving Sevier fold-thrust belt of ID-WY. Earth and Planetary Science Letters, 513, 29-39.

Page, M., Licht, A., Dupont-Nivet, G., Meijer, N., Barbolini, N., Hoorn, C., Schauer, A., Huntington, K., Bajnai, D., Fiebig, J., Mulch, A., Guo, Z. (2019) Synchronous cooling and decline in monsoonal rainfall in NE Tibet during the fall into the Oligocene Icehouse. Geology

Schwartz, T., Methner, K., Mulch, A., Graham, S.A., Chamberlain, C.P. (2019) Paleogene topographic and climatic evolution of the Northern Rocky Mountains from integrated sedimentary and isotopic data. Geological Society of Americal Bulletin.

Vasiliev, I., Reichart, G-J., Krijgsman, W., Mulch, A. (2019) Black Sea rivers capture drastic change in catchment-wide mean annual temperature and soil pH during the Miocene-to-Pliocene transition. Global and Planetary Change, 172, 428-439.



Lüdecke, T., Kullmer, O., Wacker, U., Sandrock, O., Fiebig, J., Schrenk, F., Mulch, A. (2018) Dietary versatility of early Pleistocene hominins. Proceedings of the National Academy of Sciences, 115, 52, 13330-13335. Paper featured as „Hominins had flexible diets“ (2019) in nature Human Behaviour

Antonelli, A., Kissling, W.D., Flantua, S.G.A., Bermudez, M.A., Mulch, A., Muellner-Riehl, A.N.M., Kreft, H., Lindner, H.P., Badgley, C., Fjeldsa, J., Fritz, S.A., Rahbek, C., Herman, F., Hooghiemstra, H., Hoorn, C. (2018) Geological and climatic determinants of mountain biodiversity. nature geoscience, 11, 718-725.

Meijers, M.J.M., Brocard, G.Y., Cosca, M.A., *Lüdecke, T., Teyssier, C., Whitney, D.L., Mulch, A. (2018) Rapid Late Miocene Surface Uplift of the Central Anatolian Plateau Margin. Earth and Planetary Science Letters, 497, 29-41.

 Meijers, M.J.M., Peynircioglu, A., Cosca, M.A., Brocard, G.Y., Whitney, D.L., Langereis, C., Mulch, A. (2018) Climate stability in Central Anatolia during the Messinian Salinity Crisis. Paleo­geography, Paleoclimatology, Paleoecology, 498, 53-67.

Boles, A. Mulch, A., v.d. Pluijm, B. (2018) Near-surface clay authigenesis in exhumed fault rock of the Alpine Fault Zone (New Zealand); O-H-Ar isotopic, XRD and chemical analysis of illite and chlorite. Journal of Structural Geology, 111, 27-41.

Mulch, A., Chamberlain, C.P. (2018) Stable isotope paleoaltimetry: Paleotopography as a key element in the evolution of landscapes and life. in: Mountains, Climate and Biodiversity, Hoorn, C., Perrigo, A. & Antonelli, A. (eds.) Wiley, 81-94.

Mosbrugger, V., Favre, A., Muellner-Riehl, A., Päckert, M., Mulch, A. (2018) Cenozoic evolution of Geo-Biodiversity in the Tibeto-Himalayan region. in: Mountains, Climate and Biodiversity, Hoorn, C., Perrigo, A. & Antonelli, A. (eds.) Wiley, 429-448.

**Hellwig, A., Voigt, S., Mulch, A., Bartenstein, A., Pross, J., Gerdes, A., Voigt, T. (2018) Late Oligocene-early Miocene increase in seasonality of precipitation in Central Asia recorded in terrestrial sequences from the Ili Basin (SE Kazakhstan), Sedimentology, 65, 2, 517-539.

Linnemann, U., Su T., Kunzmann, L., Spicer, R.A., Spicer, T.E.V., Zieger, J., Hofmann, M., Moraweck, K., Gärtner, A., Ding, W., Zhang, S.-T., Gerdes, A., Marko, L., Mulch, A., Mosbrugger, V., Zhou, Z.-K. (2018) New U/Pb dates show a Paleogene Origin for the modern Asian biodiversity hotspots, Geology, 46, 1, 3-6.



*Ardenghi, N., Mulch, A., Pross, J., Niedermeyer, E.M. (2017) Leaf wax n-alkane extraction: An optimised procedure. Organic Geochemistry. 113, 283-292.

*Schemmel, F., Niedermeyer, E.M., Koutsodendris, A., Pross, J., Fiebig, J., Mulch, A., (2017) Paleohydrological changes in the Eastern Mediterranean region during the early Holocene recorded in plant wax n-alkane distributions and d13CTOC – new data from Tenaghi Philippon, NE Greece. Organic Geochemistry, 110, 100-109.

Licht, A., Coster, P., Ocakoglu, F., Campbell, C., Metais, G., Mulch, A., Taylor, M., Kappelman, J., Beard, K.C. (2017) Tectono-stratigraphy of the Orhaniye Basin, Turkey: Implications for collision chronology and Paleogene biogeography of central Anatolia, Journal of Asian Earth Sciences, 143, 45-58.

Gébelin , A., Jessup, M., Teyssier, C., Cosca, M.A., Law, R.D., Brunel, M., Mulch, A. (2017) Infiltration of meteoric water in the South Tibetan Detachment (Mt. Everest, Himalaya): When and Why? Tectonics, 36, doi:10.1002/2016TC004399.

Barnosky, A.D., Hadly, E.A., Gonzalez, P., Head, J., Polly, P.D., Lawing, A.M., Eronen, J.T., … Mulch, A., … Zhang, Z. (2017) Merging paleobiology with conservation biology to guide the future of terrestrial ecosystems. Science, 355, eaah4787.

Badgley, C., Smiley, T.M., Davis, E.B., DeSantis, L.R.G., Fox, D.L., Hopkins, S.B., Jezkova, T., Matocq, M.D., Matzke, N., McGuire, J.L., Mulch, A., Riddle, B.R., Roth, L., Samuels, J.X., Strömberg, C.A.E., Terry, R., Yanites, B.J. (2017) Biodiversity and topographic complexity: Modern and geohistorical perspectives, Trends in Ecology and Evolution, 32, 211-226.

Panait, A., Diaconu, A., Galka, M., Hutchinson, S.M., Hickler, T., Lamentowicz, M., Mulch, A., Tantau, I., Werner, C., Feurdean, A. (2017.) Hydrological conditions and carbon accumulation rates reconstructed from a raised mountain bog in the Carpathians: a multi-proxy approach. Catena 152, 57-68.



**Rohrmann, A., Sachse, D., Mulch, A., **Pingel, H., Tofelde, S., Alonso, R.N., Strecker, M.R. (2016 Miocene orographic uplift forces rapid hydrological change in the southern central Andes. nature Scientific Reports 6, 35678.

Mancktelow, N., Zwingmann, H., Mulch, A. (2016) K-Ar dating of fault gouge from the Naxos detachment (Cyclades, Greece). Tectonics, 35, 2334-2344.

Haines, S., Lynch, E., Mulch, A., Valley, J.W., van der Pluijm, B. (2016) Meteoric fluid infiltration in crustal-scale normal fault systems as indicated by d18O and d2H geochemistry, and Ar dating of neoformed clays in brittle fault rocks. Lithosphere, 8, 587-600.

Fritz, S.A., Eronen, J.T., Schnitzler, J., Hof, C. Janis, C.M., Mulch, A., Böhning-Gaese, K., Graham, C.H. (2016) Humans dissolve a 20-million-year link between productivity and mammalian diversity. Proceedings of the National Academy of Sciences, doi/10.1073/pnas.1602145113

Nieto-Moreno, V., **Rohrmann, A., van der Meer, M.T.J., Sinninghe Damsté, J.S., Sachse, D., Tofelde, S., Niedermeyer, E.N., Strecker, M.R., Mulch, A. (2016) Elevation-dependent changes in n-alkane dD and soil GDGTs across the South Central Andes. Earth and Planetary Science Letters 453, 234-242.

*Lüdecke, T., Mulch, A., Kullmer, O., Sandrock, O., Thiemeyer, H., Fiebig, J., Schrenk, F. (2016) Stable isotope dietary reconstructions of herbivore enamel reveal heterogeneous wooded savanna ecosystems in the Plio-Pleistocene Malawi Rift. Palaeogeography, Palaeoclimatology, Palaeoecology. 459, 170-181.

*Methner, K., Mulch, A., Fiebig, J., Wacker, U., Gerdes, A., Graham, S.A., Chamberlain, C.P., (2016) Rapid Middle Eocene temperature change in western North America. Earth and Planetary Science Letters, 450, 132-139.

Niedermeyer, E.N., Forrest, M., Beckmann, B., Sessions, A.L., Mulch, A., Schefuß, E. (2016) The stable hydrogen isotopic composition of sedimentary plant waxes as quantitative proxy for rainfall in the West African Sahel. Geochimica Cosmochimica Acta, 184, 55-70.

Lagomarsino, L.P., Condamine, F.L., Antonelli, A., Mulch, A., Davis, C.C. (2016) The abiotic and biotic drivers of rapid diversification in Andean bellflowers (Campanulaceae). New Phytologist 210, 1430-1442.

Meijers, M.J.M., Strauss, B., Özkaptan, M., Feinberg, J.M., Mulch, A., Whitney, D.L., Kaymakci, N. (2016) Age and paleoenvironmental reconstruction of partially remagnetized lacustrine sedimentary rocks (Oligocene Aktoprak basin, central Anatolia, Turkey). Geochem. Geophys. Geosyst. 17, doi:10.1002/2015GC006209.

*Methner, K., Fiebig, J., Wacker, U., Umhoefer, P., Chamberlain, C.P., Mulch, A. (2016) Eo-Oligocene proto-Cascades topography revealed by clumped (D47) and oxygen isotope (d18O) geochemistry (Chumstick Basin, WA, USA). Tectonics, 35, doi: 10.1002/2015TC003984.

**Pingel, H., Mulch, A., Alonso, R.A., Cottle, J., Hynek, S.A., Poletti, J., **Rohrmann, A., Schmitt, A.K., Stockli, D.F., Strecker, M.R. (2016) Surface uplift and convective rainfall along the southern Central Andes (Angastaco Basin, NW Argentina). Earth and Planetary Science Letters, 440, 33-42.

*Schemmel, F., Niedermeier, E.M., Schwab, V.F., Gleixner, G., Pross, J., Mulch, A. (2016) Plant wax dD values record changing Eastern Mediterranean atmospheric circulation patterns during the 8.2 kyr B.P. climatic event. Quaternary Science Reviews, 133, 96-107.

Mulch, A. (2016) Stable isotope paleoaltimetry and the evolution of landscapes and life. Earth and Planetary Science Letters, Invited Frontiers Article 433, 180-191.

*Lüdecke, T., Schrenk, F., Thiemeyer, H., Kullmer, O., Bromage, T.G., Sandrock, O., Fiebig, J., Mulch, A. (2016) Persistent C3 vegetation accompanied Plio-Pleistocene hominin evolution in the Malawi Rift (Chiwondo Beds, Malawi). Journal of Human Evolution, 90, 163-175.

 Olaka, L., Wilke, F.D.H., Olago, D., Odada, E., Mulch, A., Musolff, A. (2016) Groundwater fluoride enrichment in an active rift setting: Central Kenya Rift case study. Science of the Total Environment, 545-546, 641-653.

Mix, H., Ibarra, D.E., Mulch, A., Graham, S.A., Chamberlain, C.P. (2016) A hot and high Eocene Sierra Nevada. Geol. Soc. America Bulletin, doi:10.1130/B31294.1

Macaulay, E.A., Sobel, E.R., Mikolaichuk, A., Wack, M., Gilder, S., Mulch, A., Fortuna, A.B., Hynek, S. (2016) The sedimentary record of the Issyk Kul basin, Kyrgyzstan: climatic and tectonic inferences. Basin Research, 28, 57-80.



Fanara, S., Botcharnikov, R.E., Palladino, D.M., Adams, F., Buddensiek, J., Mulch, A., Behrens, H. (2015) Volatiles in magmas related to Campanian ignimbrite eruption: Experiments vs. natural findings. American Mineralogist, 100, 10, 2284-2297.

Caves, J.K., Winnick, M.J., Graham, S.A., Sjostrom, D.J., Mulch, A. and Chamberlain, C.P. (2015) Role of Westerlies in Central Asia climate over the Cenozoic. Earth and Planetary Science Letters, 428, 33-43.

 Boles, A., van der Pluijm, B., Mulch, A., Mutlu, H., Uysal, I. T., Warr, L. (2015) Hydrogen and 40Ar/39Ar isotope evidence for multiple and protracted paleofluid flow events within the long-lived North Anatolian Keirogen (Turkey). Geochem. Geophys. Geosyst. DOI 10.1002/2015GC005810

Eronen, J.T., Janis, C.M., Chamberlain, C.P., Mulch, A. (2015) Mountain uplift explains differences in Palaeogene patterns of mammalian evolution and extinction between North America and Europe. Trans. Royal. Soc London B. 282: 20150136

Feurdean, A., Galka, M., Kuske, E., Tantau, I., Lamentowicz, M., Florescu, G., Hutchinson, S.M., Liakka, J., Mulch, A., Hickler, T. (2015) Last Millennium hydro-climate variability in Central Eastern Europe (Northern Carpathians, Romania). The Holocene, 25, 1179-1192. doi: 10.1177/ 0959683615580197

McFadden, R., Mulch, A., Teyssier, C., Heizler, M. (2015) Eocene extension and meteoric fluid flow in the Wildhorse Detachment, Pioneer metamorphic core complex, Idaho. Lithosphere. doi: 10.1130/L429.1

Mancktelow, N., Zwingmann, H., Campani, M., Fügenschuh, B., Mulch, A. (2015) Timing and conditions of brittle faulting on the Silltal-Brenner Fault Zone, Eastern Alps (Austria). Swiss Journal of Geosciences, 108, 305-326.

Vespasiano, G., Apollaro, C., De Rosa, R., Muto, F., Fiebig, J., Mulch, A., Marini, L. (2015) The Small Spring Method (SSM) for the definition of stable isotope – elevation relationships in Northern Calabria (Southern Italy). Applied Geochemistry, 63, 333-346.

*Methner, K., Mulch, A., Teyssier, C., Wells, M., Cosca, M., **Gottardi, R., Gébelin, A., Chamberlain, C.P. (2015) Eocene and Miocene extension, meteoric fluid infiltration and core complex formation in the Great Basin (Raft River Mountains, Utah). Tectonics 34, doi:10.1002/2014TC003766

Mulch, A., Chamberlain, C.P., Cosca, M.A., Teyssier, C., *Methner, K., Hren, M.T., Graham, S.A. (2015) Rapid change in western North American high-elevation rainfall patterns during the Middle Eocene Climatic Optimum (MECO). American Journal of Science, 315, 317-336. doi: 10.2475/04.2015.02

**Gottardi, R., Mulch, A., Teyssier, C., Valley, J. W., **Quilichini, A., Vennemann, T.W. (2015). Strain and Permeability gradients traced by stable isotope exchange in the Raft River detachment shear zone, Utah. Journal of Structural Geology, 71, 41-57. doi: 10.1016/j.jsg.2014.10.005

**Quilichini, A., Siebenaller, L., Nachlas, W.O., Teyssier, C., Vennemann, T., Heizler, M.T., Mulch, A. (2015) Infiltration of meteoric fluids in an extensional detachment shear zone (Kettle dome, WA, USA): How quartz dynamic recrystallization relates to fluid-rock interaction. Journal of Structural Geology, 71, 71-85. doi: 10.1016/j.jsg.2014.11.008

Gébelin, A., Teyssier, C., Heizler, M., Mulch, A. (2015) Meteoric water circulation in a rolling-hinge detachment system (northern Snake Range core complex, Nevada). Geol. Soc. America Bulletin, 127, 149-161. doi: 10.1130/B31063.1.



**Rohrmann, A., Strecker, M.R., Bookhagen, B., Mulch, A., Sachse, D., Pingel, H., Alonso, R.N., Schildgen, T.F., Montero, C. (2014) Can stable isotopes ride out the storm? The role of convection for water isotopes in models, records, and paleoaltimetry studies in the central Andes. Earth and Planetary Science Letters, 407, 187-195.

**Pingel, H., Alonso, R.A., Mulch, A., **Rohrmann, A., Sudo, M., and Strecker, M.R. (2014) Pliocene orographic barrier uplift in the southern Central Andes. Geology, 42, 691-694.

Nachlas, W.O., Whitney, D.L., Teyssier, C.T., Bagley, B., Mulch, A. (2014) Titanium concentration in quartz as a record of multiple deformation mechanisms in an extensional shear zone. Geochem. Geophys. Geosyst., 15, 1374–1397, doi:10.1002/2013GC005200.



Mix, H., Winnick, M.J., Mulch, A., Chamberlain, C.P. (2013) Grassland expansion as an instrument of hydrologic change in Neogene western North America. Earth and Planetary Science Letters, 377-378, 73-83.

Kent-Corson, M.L., Barnosky, A. D., Mulch, A., Carrasco, M.A., Chamberlain, C.P. (2013) Possible regional tectonic controls on mammalian evolution in western North America. Palaeogeography, Palaeoclimatology, Palaeoecology, 387, 17-26.

Feng, R., Poulsen, C.J., Werner, M., Chamberlain, C.P., Mix, H., Mulch, A. (2013) Evolution of Early Cenozoic topography, climate and stable isotopes in precipitation in the North American Cordillera. American Journal of Science, 313, 613-648.

Regnery, J., Püttmann, W., Koutsodendris, A., Mulch, A., Pross, J. (2013) Comparing the paleoclimatic significance of higher land plant biomarker concentrations and pollen data: A case study on lake sediments from the Holsteinian interglacial. Organic Geochemistry, 61, 73-84.

Gébelin, A., Mulch, A., Teyssier, C., Jessup, M.J., Law, R.D., Brunel, M. (2013) The Miocene elevation of Mount Everest. Geology, 41, 7, 799-802.

Hoorn, C., Mosbrugger, V., Mulch, A., Antonelli, A (2013) Biodiversity from mountain building. Nature geoscience, 6, 154.

Hetzel, R., Zwingmann, H., Mulch, A., Gessner, K., Akal, C., Hampel, A., Güngör, T., Petschick, R. Mikes, T., Wedin, F. (2013) Spatio-temporal evolution of brittle normal faulting and fluid infiltration in detachment fault systems – a case study from the Menderes Massif, western Turkey. Tectonics, 32, 1-13. doi:10.1002/tect.20031

*Lüdecke, T., Mikes, T., Rojay, B., Cosca, M., Mulch, A. (2013) Oligo-Miocene paleoenvironment and paleohydrology of Central Anatolian lake basins. Turkish Journal of Earth Sciences (VAMP Special Volume); 22, 793-819. DOI: 10.3906/yer-1207-11

*Schemmel, F., Mikes, T., Rojay, B., Mulch, A. (2013) The impact of topography on isotopes in precipitation across the Central Anatolian Plateau (Turkey). Am. Journal of Science, 313, 61-80.

Mazzini, I., Hudackova, N., Joniak, P., Kovacova, M., Mikes, T., Mulch, A., Rojay, B., Lucifora, S., Esu, D., Soulie-Märsche, I. Palaeoenvironmental And Chronological Constraints On The Tuğlu Formation (Çankırı Basin, Central Anatolia, Turkey) (2013) Turkish Journal of Earth Sciences (VAMP Special Volume) 22, 747-777. DOI: 10.3906/yer-1207-10



Campani, M., Mulch, A., Kempf, O., Schlunegger, F., Mancktelow, N. (2012) Miocene paleotopography of the Central Alps. Earth and Planetary Science Letters, 337-338, 174-185.

Gebelin, A., Mulch, A., Teyssier, C., Chamberlain, C.P., Heizler, M. (2012) Coupled basin-detachment systems as paleoaltimetry archives of the western North American Cordillera, Earth and Planetary Science Letters, 335-336, 36-47.

Chamberlain, C.P., Mix, H.T., Mulch, A., Hren, M.T., Kent-Corson, M.L., Davis, S.J., Horton, T.W., Graham, S.A. (2012) The Cenozoic Climatic and Topographic Evolution of the Western North American Cordillera, American Journal of Science, 312, 213-262. DOI 10.2475/04.2011.00



Königer, P., Marshall, J.D., Link, T., Mulch, A. (2011) An inexpensive, fast, and reliable method for vacuum extraction of soil and plant water for stable isotope analyses by mass spectrometry. Rap. Comm. Mass Spect., 25, 3041-3048.

 Gebelin, A., Mulch, A., Teyssier, C., Heizler, M., Vennemann, T.W., Seaton, N.C.A. (2011) Oligo-Miocene extensional tectonics and fluid flow across the Northern Snake Range detachment system, Nevada. Tectonics, 30, TC5010 DOI 10.1029/2010TC002797.

Gottardi, R., Teyssier, C., Mulch, A., Vennemann, T. W., Wells, M. L. (2011) Preservation of an extreme geotherm in the Raft River detachment shear zone. Geology, 39, 759-762.

Mix, H., Mulch, A., Kent-Corson, M.L., Chamberlain, C.P. (2011) Cenozoic migration of topography in the North American Cordillera, Geology, 39, 87-90.



Cecil M.R. Ducea, M., Reiners, P., Gehrels, G., Mulch, A., Allen, C., Campbell, I. (2010) Provenance of Eocene river sediments from the central northern Sierra Nevada and implications for paleotopography. Tectonics, 29, TC6010.

Ballato, P., Mulch, A., Landgraf A., Strecker M.R., Dalconi M.C., Friedrich A., Tabatabaei S.H. (2010) Middle to late Miocene Middle Eastern climate from stable oxygen and carbon isotope data, southern Alborz mountains, N Iran. Earth and Planetary Science Letters, 300, 125-138.

**Kent-Corson, M.L., Mulch, A., Graham, S.A., Carroll, A.C., Ritts, B.D., Chamberlain, C.P, (2010) Diachronous isotopic and sedimentary responses to topographic change as indicators of mid-Eocene hydrologic reorganization in the western United States. Basin Research 22, 6, 829-845.

Mulch A., Uba, C., Strecker, M.R., Schönberg, R., Chamberlain, C.P. (2010) Late Miocene climate variability and surface elevation in the central Andes. Earth and Planetary Science Letters, 290, 173-182.

Doebbert A., Carroll, A.R., Mulch, A., Chetel, L., Chamberlain, C.P. (2010) Geomorphic Controls On Lacustrine Isotopic Compositions: Evidence From The Laney Member, Green River Formation (Wyoming). Geological Society of America Bulletin, 122, 236-252.



Davis, S.J., Mulch A., Carroll, A.R., Horton T.W., Chamberlain, C.P. (2009) Paleogene landscape evolution of the central North American Cordillera: Developing topography and hydrology in the Laramide foreland. Geological Society of America Bulletin, 121, 100-116.



Garzione, C.N., Hoke, G.D., Libarkin, J.C., Withers, S., MacFadden B., Eiler, J., Mulch, A. (2008) The Rise of the Andes. Science, 320, 1304-1307.

Mulch, A., Sarna-Wojcicki, A.M., Perkins, M.E., Chamberlain, C.P. (2008) A Miocene to Pleistocene climate and elevation record of the Sierra Nevada, California. Proceedings of the National Academy of Sciences 105, 19, 6819-6824.

Agosta, F., Mulch, A., Chamberlain, C.P., Aydin, A. (2008) Geochemical traces of CO2-rich fluid flow along normal faults in central Italy. Geophysical Journal International 174, 758-770.



Mulch, A. and Chamberlain, C.P. (2007) Stable Isotope Paleoaltimetry in Orogenic Belts – The silicate record in surface and crustal geological archives. Reviews in Mineralogy and Geochemistry, 66, 89-118.

Mulch, A., Teyssier, C., Cosca, M.A., and Chamberlain, C.P. (2007) Stable isotope paleoaltimetry of Eocene Core Complexes in the North American Cordillera. Tectonics, 26, TC4001, doi:10.1029/ 2006TC001995.

Person, M., Mulch, A., Teyssier, C., Gao, Y. (2007) Isotope Transport and Exchange within Metamorphic Core Complexes. American Journal of Science 307, 555-589.

Monjoie, P., Bussy, F., Schaltegger, U., Mulch, A., Lapierre, H., Pfeifer, H.R. (2007) Contrasting magma types and timing of intrusion in the Permian layered mafic complex of Mont Collon (Western Alps,Valais, Switzerland): evidence from U/Pb zircon and 40Ar/39Ar amphibole dating. Swiss Journal of Geosciences 100, 125–135.



Mulch, A., Graham, S.A., and Chamberlain, C.P. (2006) Hydrogen Isotopes in Eocene River Gravels and Paleoelevation of the Sierra Nevada. Science 313, 87-89.*elected to be among the „Top 100 Science papers in 2006“ (Discover v. 28, 01, 2007)

Mulch, A. & Chamberlain, C.P. (2006) The rise and growth of Tibet. Nature 439, p. 670.

**Kent-Corson, M.L., Sherman, L.S., Mulch, A., and Chamberlain, C.P., (2006) Cenozoic topographic and climatic response to changing tectonic boundary conditions in western North America. Earth and Planetary Science Letters 252, 453-466.

Mulch, A., Teyssier, C., Cosca, M.A., and Vennemann, T. (2006) Thermomechanical analysis of strain localization in a detachment zone. Journal of Geophysical Research, 111, B12405, doi:10.1029/2005JB004032



Mulch, A., Cosca, M.A., Fiebig, J., and Andresen, A. (2005) Time scales of mylonitic deformation and meteoric fluid infiltration during extensional detachment faulting: an integrated in situ 40Ar/39Ar geochronology and stable isotope study of the Porsgrunn-Kristiansand Shear Zone (Southern Norway). Earth and Planetary Science Letters, 233, 375-390.


2004 – 2001

Mulch, A., Teyssier, C., Cosca, M.A., Vanderhaeghe, O., and Vennemann, T. (2004) Reconstructing paleoelevation in eroded orogens. Geology. 32, 6, 525-528. *paper was highlighted in Science 305, 19 and Geology Today 20, 5, 118-119.

Mulch, A. and Cosca, M.A. (2004) Recrystallization or cooling ages? – In situ UV-laser 40Ar/39Ar geochronology of muscovite in mylonitic rocks. Journal of the Geological Society London 161, 573-582.

Mulch, A., Cosca, M.A., and Handy, M. R. (2002) In-situ UV-laser 40Ar/39Ar geochronology of a micaceous mylonite – an example of defect-enhanced argon loss. Contributions to Mineralogy and Petrology 142, 738-752.

Mulch, A., Rosenau, M. R., Dörr, W., and Handy, M. R. (2002) The age and structure of dikes along the tectonic contact of the Ivrea-Verbano and Strona-Ceneri Zones (southern Alps, Northern Italy, Switzerland). Swiss Bulletin of Mineralogy and Petrology 82, 55-76

Handy, M.R., Mulch, A., Rosenau, M.R., and Rosenberg, C.L. (2001) The role of fault zones and melts as agents of weakening, hardening and differentiation of the continental crust: a synthesis. in: Holdsworth R.E., Strachan, R.A., Magloughlin, J.F., and Knipe, R.J. (eds.): The Nature and Tectonic Significance of Fault Zone Weakening. Geological Society London, Special Publications 186, 305-332.

Karsten Wesche
Prof. Dr. Karsten Wesche
Head of Botany division, Senckenberg Museum of Natural History Görlitz
Abteilungsleiter Botanik, Senckenberg Museum für Naturkunde Görlitz

Born 1970 in Lüchow, Lower Saxony, Germany

1990 to 1996: university studies; subject biology (botany, conservation biology, ecology) and related fields (soil science, geography) in Marburg, Gießen, and Aberdeen (UK). Diploma thesis on tropical Sal forests in southern Nepal

Aug. 1996 to Jan. 2000: PhD. project at the Fac. of Geography, Univ. of Marburg. Fieldwork on Mt. Elgon (Uganda/Kenya). Thesis: “The high-altitude environment of Mt. Elgon (Uganda/Kenya). Climate, vegetation and the impact of fire”

June 2000 to April 2008: Post-doctoral researcher/scientific assistant at the Institute of Biology – Geobotany and Botanical Garden, Univ. Halle-Wittenberg

July 2007: Habilitation / venia legendi for Botany, thesis: “Plant survival in southern Mongolian desert steppes – Ecology of communities, interactions and populations” (submitted Oct. 2006)

May 2008 to Dec. 2009: Scientific assistant / project coordinator for “BioChange – Germany”, Dept. of Ecology and Ecosystems Research, Georg August University Göttingen

Jan 2010 onwards: Head of Botany Division, Senckenberg Museum for Natural History, Görlitz

July 2016 onwards: Professor, Chair Biodiversity of Plants, International Institute Zittau, Technical University Dresden

Fieldwork in a number of countries including: Switzerland, Mongolia, China, Nepal, Russia, Uganda, Kenya, Tanzania, Ethiopia, Cameroon, Bolivia, Costa Rica (and Germany of course)

Funding member German Centre for Integrative Biodiversity Research (iDiv) Halle-Jena-Leipzig. Subject editor Flora, member of the editorial boards of Journal of Vegetation Science / Applied Vegetation Science and Phytocoenologia, Co-editor Rothmaler-Exkursionsflora.

Publications are ordered chronologically (peer-reviewed first, others follow)


Khurelpurev, O., Herklotz, V., Harpke, D., Batlai, O., Wesche, K., Ritz, C.M. (2021) Contrasting effects of local environment and grazing pressure on genetic diversity and structure of Artemisia frigida. Conservation Genetics.

Shukherdorj Baasanmunkh, Batlai Oyuntsetseg, Chuluunkhuyag Oyundari, Khurelpurev Oyundelger, Magsar Urgamal, Davaajav Darikhand, Nergui Soninkhishig, Dashzeveg Nyambayar, Khurelbaatar Khaliunaa, Zagarjav Tsegmed, Kechaykin, A. A., Shmakov, A. I., Erst, A. S., Friesen, N., Ritz, C. M., Wesche, K., Hyeok Jae Choi. (2021) The vascular plant diversity of Dzungarian Gobi in western Mongolia, with an annotated checklist. – Phytotaxa 501: 1-55.

Lorenz, J., Heinrich, R., Schneider, A., Schwager, M., Herklotz, V., Wesche, K., Ritz, C.M. (2021) Invasive populations of Spiraea tomentosa (Rosaceae) are genetically diverse but shrinking under increasing canopy cover. Plant Biology.


Valencia, E., de Bello, F., Galland, T., Adler, P.B., Lepš, J., E-Vojtkó, A., van Klink, R., Carmona, C.P., Danihelka, J., Dengler, J., Eldridge, D.J., Estiarte, M., García-González, R., Garnier, E., Gómez‐García, D., Harrison, S.P., Herben, T., Ibáñez, R., Jentsch, A., Juergens, N., Kertész, M., Klumpp, K., Louault, F., Marrs, R.H., Ogaya, R., Ónodi, G., Pakeman, R.J., Pardo, I., Pärtel, M., Peco, B., Peñuelas, J., Pywell, R.F., Rueda, M., Schmidt, W., Schmiedel, U., Schuetz, M., Skálová, H., Šmilauer, P., Šmilauerová, M., Smit, C., Song, M., Stock, M., Val, J., Vandvik, V., Ward, D., Wesche, K., Wiser, S.K., Woodcock, B.A., Young, T.P., Yu, F.-H., Zobel, M., & Götzenberger, L. 2020. Synchrony matters more than species richness in plant community stability at a global scale. – Proceedings of the National Academy of Sciences, 201920405

Valencia, E. de Bello, F., Lepš, J.,  …  Wesche, K, … Zobel, M., Götzenberger, L. 2020. Directional trends in species composition over time can lead to a widespread overemphasis of year‐to‐year asynchrony. – Journal of Vegetation Science:

Török, P., Neuffer, B., Heilmeier, H., Bernhardt, K. G., Wesche, K. 2020. Climate, landscape history and management drive Eurasian steppe biodiversity. – Flora, 151685

Müller, F., Ritz, C, Welk, E., Wesche, K. in press. Rothmaler Exkursionsflora von Deutschland. Grundband. 22., grundlegend überarbeite Auflage. Springer Spektrum, Berlin, Heidelberg, ca. 1100 S.

Eichenberg, D., Bernhardt-Römermann, M., Bowler, D., Bruelheide, H., Conze K., Dauber, J., Dengler, J., Engels D., Fartmann, T., Frank D., Geske C., Grescho V., Harter, D., Henle, K., Hofmann, S., Jandt, U., Jansen, F., Kamp, J., Kautzner A., König-Ries, B., Krämer R., Krüß, A., Kühl, H., von B., Ludwig, M., Lueg H., May R., Musche, M., Opitz, A., Ronnenberg, K., Schacherer, A., Schäffler, L., Schiffers, K., Schulte, U., Schwarz J., Sperle T., Stab S., Stöck, M., Theves, F., Trockur, B., Wesche, K., Wessel M., Winter, M., Wirth, C., Bonn, A. in press. Langfristige Biodiversitätsveränderungen in Deutschland erkennen – mit Hilfe der Vergangenheit in die Zukunft schauen. – Natur und Landschaft.

Bruelheide, H., Jansen, F., Jandt, U., Bernhardt-Römermann, M., Bonn, A., Bowler, D., Dengler, J., Eichenberg, D., Grescho, V., Harter, D., Jugelt, M., Kellner, S., Ludwig, M., Wesche, K., Lütt, S. 2020. Using incomplete floristic monitoring data from habitat mapping programmes to detect species trends. – Diversity and Distributions: online first.

Oyundelger, K., Ritz, C.M., Munkhzul, O., Lang, B., Ahlborn, J., Oyuntsetseg, B., Römermann, C., Wesche, K. 2020. Climate and land use affect genetic structure of Stipa glareosa P. A. Smirn. in Mongolia. – Flora 266: 151572.

Borsch, T., Stevens, A.-D., Häffner, E., … Wesche, K., Yurkov, A., Zizka, G. 2020. A complete digitization of German herbaria is possible, sensible and should be started now. – Research Ideas and Outcomes 6: e50675.

Treiber, J., von Wehrden, H., Zimmermann, H., Welk, E., Jäger, E.J., Ronnenberg, K., Wesche, K. 2020. Linking large-scale and small-scale distribution patterns of steppe plant species — An example using fourth-corner analysis. – Flora 263: 151553.

Jäschke, Y., Heberling, G., Wesche, K. 2020. Environmental controls override grazing effects on plant functional traits in Tibetan rangelands. – Functional Ecology: 34:747–760

Ahlborn, J., von Wehrden, H., Lang, B., Römermann, C., Oyunbileg, M., Oyuntsetseg, B., Wesche, K. 2020. Climate – grazing interactions in Mongolian rangelands: Effects of grazing change along a large-scale environmental gradient. – Journal of Arid Environments 173: 104043.

Lang, B., Ahlborn, J., Munkhzul, Oyunbileg, Geiger, A., von Wehrden, H., Wesche, K., Oyuntsetseg, Batlai, Römermann, C.  2020. Grazing effects on intraspecific trait variability vary with changing precipitation patterns in Mongolian rangelands. – Ecology and Evolution: online first.

Schuch, S., Ludwig, H., Wesche, K. 2020. Erfassungsmethoden für ein Insektenmonitoring – Eine Materialsammlung. – BfN-Skripten 565: 83 S.


Baasanmunkh, S., Shiga, T., Batlai, O., Wesche, K., Ritz, C.M., Khaliunaa, K., Kim, J.Y., Jo, H.J., Batkhuu, N.-O., Chung, G.Y., Choi, H.J. 2019 Contribution to the knowledge on the flora of Numrug Strictly Protected Area and some parts of East Mongolia.  –  Journal of Asia-Pacific Biodiversity: 12: 284-301.

Bruelheide, H., Dengler, J., Jimenéz-Alvaro, B. et al. 2019. sPlot – a new tool for global vegetation analyses – Journal of Vegetation Science: 30: 161-186.

Schuch, S., Meyer, S., Bock, J., van Klink, R., Wesche, K. 2019. Drastischer Biomassenverlust bei Zikaden verschiedener Grasländer innerhalb von fünf bis sechs Jahrzehnten. – Natur und Landschaft 94: 141-145.

Diekmann, M., Andres, C., Becker, T., Bennie, J., Blüml, V., Bullock, J.M., Culmsee, H., Fanigliulo, M., Hahn, A., Heinken, T., Leuschner, C., Luka, S., Meißner, J., Müller, J., Newton, A., Peppler-Lisbach, C., Rosenthal, G., van den Berg, L.J.L., Vergeer, P., Wesche, K. 2019. Patterns of long-term vegetation change vary between different types of semi-natural grasslands in Western and Central Europe. – Journal of Vegetation Science 30: 187-202.

Lv Chaoyan, Schlecht, E., Goenster-Jordan, S., Buerkert, S., Zhang Ximing, Wesche, K. 2019. Vegetation responses to fixed stocking densities in highly variable montane pastures in the Chinese Altay. – Rangeland Ecology and Management 72: 812–821.

Miehe, G., Schleuss, P.-M., Seeber, E., Jäschke, Yun, Lehnert, L., Biermann, T., Babel, W., Ingrisch, J., Gerken, T., Willinghöfer, S., Hafner, S., Unteregelsbacher, S., Liu, Shibin Holzapfel, M.,  Coners, H., Miehe, S., Leuschner, C., Guggenberger, G., Kuzyakov,Y., Xu, Xingliang, Spielvogel, S., Foken, T., Graf, H.-F., Ma, Yaoming, Yang, Yongping,  Zhang, Shuren, Noltie, H. J. Opgenoorth, L., Schmidt, J., Braendle, M., Lai, Zhongping,  Mosbrugger, V., Chen Fahu, Liu, Jianquan,  Wesche, K. 2019. The Kobresia pygmaea ecosystem of the Tibetan highlands – origin, functioning and degradation of the world’s largest pastoral alpine ecosystem. – Science of the Total Environment 648: 754-771.

Lang, B., Geiger, A., Oyunbileg, M., Ahlborn, J., von Wehrden, H., Wesche, K., Oyuntsetseg, B., Römermann, C. 2019. Intraspecific trait variation patterns along a precipitation gradient in Mongolian rangelands. – Flora 254, 135-146.

Lashchinsky, N., Korolyuk, A., Wesche, K. (2019). Vegetation Patterns and Ecological Gradients: From Forest to Dry Steppes. In: Guggenberger (G.) KULUNDA: Climate Smart Agriculture, pp. 33-48. Springer, Heidelberg

Wesche, K., Korolyuk, A., Lashchinsky, N., Silanteva, M.M., Rosche, C. and Hensen, I. 2019. The Kulunda steppe as part of the Eurasian steppe belt. In: Guggenberger (G.) KULUNDA: Climate Smart Agriculture, pp. 33-48. Springer, Heidelberg

IPBES Global Assessment (2019): K. Wesche contributing author on Temperate Grassland.

Dressler, S., Ritz, C., Wesche, K. 2019. Vom Kräuterbuch zum Informationsknoten – die heimische Flora in Forschung und Anwendung. In: Senckenberg 2018. Senckenberg Gesellschaft für Naturforschung, Frankfurt am Main, pp. 30-33

Xylander, W.E.R., Ansorge, H., Balkenhol, B., Burkhardt, U., Christian, A., Damm, U., Düker, Ch., Gebauer, P., Hohberg, K., Hutchinson, J.M.C., Lehmitz, R., Otte, V., Reise, H., Ritz, C., Russell, D. J., Schlitt, B., Schulz, H.-J., Seifert, B., Tietz, O., Voigtländer, K., Wesche, K. 2019. Obituary Prof. Dr. Wolfram Dunger. – Soil Organisms 91.

Franke, R., Gebauer, P, Ritz C., Wesche, K. 2019. Zum 70. Geburtstag von Dr. Friedrich Wilhelm Sander – Berichte der Naturforschenden Gesellschaft der Oberlausitz 27: 225-226.

Pfeiffer, M., C. Dulamsuren, Y., Wesche K. 2019. Grasslands and shrublands of Mongolia. – Encyclopedia of the World’s Biomes. Pp. 14.

Müller, F., Ritz, C, Welk, E., Wesche, K. 2019. Kommentare und Ergänzungen zur Neubearbeitung der 22. Auflage der Rothmaler Exkursionsflora von Deutschland Grundband 2019: Eine Vorbemerkung. – Schlechtendalia 35: 41.  

Wesche, K. in press. Teil E: Subzonobiom VII (rIll) der zentralasiatischen Wüsten. In: Breckle, S.W. Ökologie der Erde. Bd. 3. Spezielle Ökologie der gemäßigten Zone. Borntraeger. 


Bruelheide, H., Dengler, J., Purschke, O., Lenoir, J., Jiménez-Alfaro, B., Hennekens, S.M., Botta-Dukát, Z., Chytrý, M., Field, R., Jansen, F., Kattge, J., Pillar, V.D., Schrodt, F., Mahecha, M.D., Peet, R.K., Sandel, B., van Bodegom, P., Altman, J., Alvarez Davila, E., Arfin Khan, M.A.S., Attorre, F., Aubin, I., Baraloto, C., Barroso, J.G., Bauters, M., Bergmeier, E., Biurrun, I., Bjorkman, A.D., Blonder, B., Čarni, A., Cayuela, L., Černý, T., Cornelissen, J.H.C., Craven, D., Dainese, M., Derroire, G., De Sanctis, M., Díaz, S., Doležal, J., Farfan-Rios, W., Feldpausch, T.R., Fenton, N.J., Garnier, E., Guerin, G.R., Gutiérrez, A.G., Haider, S., Hattab, T., Henry, G., Hérault, B., Higuchi, P., Hölzel, N., Homeier, J., Jentsch, A., Jürgens, N., Kącki, Z., Karger, D.N., Kessler, M., Kleyer, M., Knollová, I., Korolyuk, A.Y., Kühn, I., Laughlin, D.C., Lens, F., Loos, J., Louault, F., Lyubenova, M.I., Malhi, Y., Marcenò, C., Mencuccini, M., Müller, J.V., Munzinger, J., Myers-Smith, I.H., Neill, D.A., Niinemets, Ü., Orwin, K.H., Ozinga, W.A., Penuelas, J., Pérez-Haase, A., Petřík, P., Phillips, O.L., Pärtel, M., Reich, P.B., Römermann, C., Rodrigues, A.V., Sabatini, F.M., Sardans, J., Schmidt, M., Seidler, G., Silva Espejo, J.E., Silveira, M., Smyth, A., Sporbert, M., Svenning, J., Tang, Z., Thomas, R., Tsiripidis, I., Vassilev, K., Violle, C., Virtanen, R., Weiher, E., Welk, E., Wesche, K., Winter, M., Wirth, C., Jandt, U. 2018. Global trait–environment relationships of plant communities. – Nature Ecology and Evolution 2: 1906-1917.

Ashastina, K., Kuzmina, S., Rudaya, N., Troeva, E., Schoch, W.H., Römermann, C., Reinecke, J., Otte, V., Savvinov, G., Wesche, K., Kienast, F. 2018. Woodlands and steppes: Pleistocene vegetation in Yakutia’s most continental part recorded in the Batagay permafrost sequence. – Quaternary Science Reviews 196: 38-61.

Wang, Yun, Lehnert, L., Holzapfel, M., Schultz, R., Heberling, G., Görzen, E., Meyer, H., Seeber, E., Pinkert, S., Ritz, M., Fu Yao, Ansorge, H., Bendix, J., Seifert, B., Miehe, G., Long Ruijun, Yang Yong-Ping, Wesche, K. 2018. Multiple indicators yield diverging results on grazing degradation and climate controls across Tibetan pastures. – Ecological Indicators 93: 1199-1208.

Dengler, J., Wagner, V., Dembicz, I., … Wesche, K. & Biurrun, I. 2018. GrassPlot – a database of multi-scale plant diversity in Palaearctic grasslands. – Phytocoenologia 48: 331–347.

Tserendulam, Ts., Wesche, K., Bayarsaikhan, U., Oyuntsetseg, B. 2018. The vascular plant flora of Hustai National Park, Mongolia: Composition, life forms, ecological groups and geographical elements. – Feddes Repertorium 129: 137-160.

Monaco, E. T., Borries, C., Nikolei, J., Chalise, M. K., Ganzhorn, J. U., Wesche, K., König, A. 2018. The function of geophagy in Nepal gray langurs: Sodium acquisition rather than detoxification or prevention of acidosis. – American Journal of Physical Anthropology 165: 1-10.

Schneider. J. V., Negraschis, V., Habersetzer, J., Rabenstein, R., Wesenberg, J., Wesche, K., Zizka, G. 2018. Taxonomic diversity masks leaf vein–climate relationships: lessons from herbarium collections across a latitudinal rainfall gradient in West Africa – Botany Letters: online first.

Schneider. J. V., Rabenstein, R., Wesenberg, J., Wesche, K., Zizka, G., Habersetzer, J. 2018. Improved non-destructive 2D and 3D X-ray imaging of leaf venation. – Plant Methods 14: 7-15.

Rosche, C., Hensen, I., Silantieva, M. M., Stolz, J., Gröning, S., Wesche, K. 2018. Spatio-environmental determinants of the genetic structures of co-occurring species in a highly fragmented steppe. – Basic and Applied Ecology 28: 48-59.

IPBES Regional Assessment – Asia Pacific (3rd order draft): K. Wesche contributing author on Temperate Grasslands.

Pfeiffer, M., C. Dulamsuren, Y. Jäschke, Wesche K. 2018. Grasslands of China and Mongolia: Spatial Extent, Land Use and Conservation. Pp. 167-196 in: Squires, V.R., Dengler, J., Feng, H., Hua, L. (eds.) . CRC Press, Boca Raton, US. ISBN 9781498796262.


Hirsch, H., Brunet, J.,  Zalapa, J.E., von Wehrden, H., Hartmann, M., Kleindienst, C., Schlautmann, B., Kosman, E., Wesche, K., Renison, D. & Hensen, I. 2017. Intra- and interspecific hybridization in invasive Siberian elm. – Biological Invasions 19: 1889-1904.

Schneider, J.V., Habersetzer, J., Rabenstein, R., Wesenberg, J., Wesche, K., Zizka, G. 2017. Water supply and demand remain coordinated during breakdown of the global scaling relationship between leaf size and major vein density. – New Phytologist 214: 473-486.

Reinecke, J., Troeva, E., Wesche, K. 2017. Extrazonal steppes of northern Siberia – Phytosociological classification and ecological characterization. – Phytocoenologia: 47: 167–196.

Dressler, S., Gregor, T., Hellwig, F., Korsch, H., Wesche, K., Wesenberg, J., Ritz, C. 2017. Comprehensive  and reliable: a new virtual herbarium of critical plant taxa in Germany. – Plant Systematics and Evolution 303: 1109–1113. DOI 10.1007/s00606-017-1419-6

Al-Gharaibeh, M.,  Hamasha, H.,  Rosche, C., Lachmuth, S., Wesche, K., Hensen, I. 2017. Environmental gradients shape the genetic structure of two medicinal Salvia species in Jordan. – Plant Biology 19: 227–238.

Paule, J., Gregor, T., , Schmitt, M., Gerstner, E.-M., Dersch, G., Dressler, S., Wesche, K., Zizka, G. 2017. Chromosome numbers of the flora of Germany – a new online database of georeferenced chromosome counts and flow cytometric ploidy estimates. – Plant Systematics and Evolution 303:1123–1129.

Wagner, Chytrý, M., Zelený, D., Wehrden, H., Brinkert, A., Danihelka, J., Hölzel, N., Kamp, J. H., Lustyk, P., Merunková, K., Palpurina, S., Preislerová, Z., Wesche, K.  2017. Plant species vary their soil pH niche in dry grasslands across Eurasia. – Oikos 126: 660–670.

Palpurina, S., Wagner, V., von Wehrden, H., Hajék, M., Horsak, M., Brinkert, A., Hölzel, N., Wesche, K., Kamp, J., … Chytry, M. 2017. The relationship between soil pH and plant species richness vanishes with increasing aridity across Eurasian dry grasslands. – Global Ecology and Biogeography 26: 425-434.

Wang Yun, Heberling, G., Görzen, E., Miehe, G., Seeber, E., Wesche, K. 2017. Combined effects of livestock grazing and environment on plant species composition and soil condition across Tibetan grasslands. – Applied Vegetation Science 20: 327-339.

Wesenberg, J., Gebauer, P., Ritz, C.M., Wesche, K. 2017. Virtuelles Herbarium der Lausitz. – Senckenberg Gesellschaft für Naturforschung, online resource.

Dressler, S., Gregor, T., Hellwig, F.H., Korsch, H., Wesche, K., Wesenberg, J., Ritz, C.M. 2017. Bestimmungskritische Taxa der deutschen Flora. – Herbarium Senckenbergianum Frankfurt/Main, Görlitz & Herbarium Haussknecht Jena .

Dedek, M., Wesche, K. 2017 Die Segetalflora der Oberlausitz im Wandel – heutige Situation im Vergleich zu historischen Daten von Max Militzer. – Berichte der Naturforschenden Gesellschaft der Oberlausitz 25: 83-106.

Wesche, K. & Ritz, C. (Hrsg.) 2017. Von der Heide ins Gebirge – vielfältige Kulturlandschaft im Osten Deutschlands. Tuexenia, Beiheft 10. pp. 1-116. – darin:

Wesche, K., Beck, A., Ritz, C.M. 2017. Riesengebirge: Mitteleuropas nördlichster Hochgebirgskamm. – Tuexenia, Beiheft 10. pp. 95-112. 

Culmsee, H., Wesche, K. 2017. Das Biosphärenreservat Oberlausitzer Heide- und Teichlandschaft: Vielfalt in Biodiversität und Landnutzung. – Tuexenia, Beiheft 10. pp. 25-48.

Damm, U., Dressler, S., Hoppenrath, M., Kienast, F., Otte, V., Paule, J., Printzen, C., Ritz, C., Wesche, K., Wesenberg, J., Zizka, G. 2017. Botanik und Mykologie bei Senckenberg seit 1817 – Herbarium Senckenbergianum Frankfurt/Main, Görlitz, Weimar, Wilhelmshaven. – Senckenberg Gesellschaft für Naturforschung, Frankfurt am Main.

Damm, U., Dressler, S., Hoppenrath, M., Kienast, F., Otte, V., Paule, J., Printzen, C., Ritz, C., Wesche, K., Wesenberg, J., Zizka, G. 2017. Botany and mycology at Senckenberg since 1817 – Herbarium Senckenbergianum Frankfurt/Main, Görlitz, Weimar, Wilhelmshaven. – Senckenberg Gesellschaft für Naturforschung, Frankfurt am Main.

Jäger, E., Müller, F., Ritz, C, Welk, E., Wesche, K. 2017. Rothmaler Exkursionsflora von Deutschland. Gefäßpflanzen: Atlasband. 13. Auflage. Springer Spektrum, Berlin, Heidelberg, 822 S.


Hirsch, H., Hensen, I., Wesche, K., Renison, D., Luti, R., Wypior, C., Hartmann, M., von Wehrden, H. 2016. Evolution of an invasive tree: non-native Siberian elms outperform native provenances. – AoB Plants 8: plw071 (

Heinicke, S., Hensen, I., Rosche, C., Hanselmann, D., Gudkova, P. D.; Silanteva, M. M.; Wesche, K. 2016. Fragmentation and environmental constraints influence genetic diversity and germination of Stipa pennata in natural steppes – Flora 224: 42–49.

Wiehle, M., Vornam, B., Wesche, K.,  Goenster S., Buerkert, A. 2016. Population structure and genetic diversity of Populus laurifolia in fragmented riparian gallery forests of the Mongolian Altai Mountains. – Flora 224: 112–122.

Lehnert, L., Wesche, K., Trachte, K., Reudenbach, C., Bendix, J. 2016. Climate variability rather than overstocking  causes recent degradation of Tibetan pastures. – Scientific Reports 6: 24367 DOI: 10.1038/srep24367.

Reichel, K., Richter, F.,  Eichel, L., Kącki, Z.,  Wesche, K. Welk, E., Neinhuis, C.,  Ritz, C. 2016. Genetic diversity in the locally declining Laserpitium prutenicum L. and the more common Selinum carvifolia (L.) L. – a “silent goodbye”?” – Conservation Genetics 17:847-860.

Kadereit, J. W., Albach, D. C., Ehrendorfer, F., Galbany-Casals, M., Garcia-Jacas, N., Gehrke, B., Kadereit. G., Kilian, N., Klein, J. T., Koch, M. A., Kropf, M., Oberprieler,  C., Pirie, M. D., Ritz, C. M., Röser, M., Spalik, K., Susanna, A., Weigend, M., Welk., E., Wesche, K., Li-Bing Zhang & Dillenberger, M. (2016). Generic circumscriptions in the German flora. – Willdenowia 46:39-91.

Törok, P., Ambarli, D., Kamp, J., Wesche, K., Dengler, J. 2016. Step(pe) up! Raising the profile of the Palaearctic natural grasslands. – Biodiversity and Conservation 25: 2187–2195.

Wesche, K., Ambarli, D., Kamp, J., Törok, P., Treiber, J., Dengler, J. 2016. The Palaearctic steppe biome: a new synthesis – Biodiversity and Conservation 25:2197–2231.

Wang, Yun, Wesche, K. 2016. Vegetation and soil responses to livestock grazing in Central Asian grasslands: a review of Chinese literature. – Biodiversity and Conservation 25: 2401–2420.

Karger, D. N., Cord, A., Kessler, M., Kreft, H., Kühn, I., Pompe, S., Sandel, B., Sarmento-Cabral, J., Smith, A., Svenning, J. C., Tuomisto, H., Weigelt, P., Wesche, K. 2016. Probabilistic species pools in ecology and biogeography. – Global Ecology and Biogeography 25: 489–501.

Müller, F., Ritz, C, Welk, E., Wesche, K. 2016. Rothmaler Exkursionsflora von Deutschland. Kritischer Ergänzungsband. Springer Spektrum, Berlin, Heidelberg, 221. S.

Wesche, K., Otte, V., Damm, U., Ritz, C., Wesenberg, J., Boyle, H., Gebauer, P. 2016. Die botanisch-mykologischen Sammlungen in Görlitz – zentrale Anlaufstellen für die berufliche und ehrenamtliche Pflanzen- und Pilzkunde in der Oberlausitz. – Berichte der Naturforschenden Gesellschaft der Oberlausitz 24: 37-50.

Reinecke, J., Troeva, E., Wesche, K. 2016. Was von der Mammutsteppe übrig blieb. Beweidete Steppeninseln in Nordsibirien sind Relikte eiszeitlicher Vergangenheit – Natur, Forschung, Museum 146: 275-283.


Seeber E., Miehe G., Hensen I., Yang Y., Wesche K. 2015. Mixed reproduction strategy and polyploidy facilitate dominance of Kobresia pygmaea on the Tibetan Plateau. – Journal of Plant Ecology 9: 87-99.

Meyer, S., Bergmeier, E., Becker, T., Wesche, K., Krause, B. & Leuschner, C. 2015. Detecting long-term losses at the plant community level – arable fields in Germany revisited. – Applied Vegetation Science 18, 432-442.

Meyer, S., Hans, J., Wesche, K., Leuschner, C., Albach, D. 2015. Landscape complexity has limited effects on genetic structure of two arable weed species, Adonis aestivalis and Consolida regalis. – Weed Research 55: 406-415.

Krause, B., Culmsee, H., Wesche, K., Leuschner, C. 2015. Historical and recent fragmentation of temperate floodplain grasslands: Do patch size and distance affect plant species richness of wet meadow communities?” – Folia Geobotanica: 50:253-266.


Babel, T. Biermann, H. Coners, E. Falge, E. Seeber, J. Ingrisch, P.-M. Schleuß, T. Gerken, J. Leonbacher, T. Leipold, S. Willinghöfer, K. Schützenmeister, O. Shibistova, L. Becker, S. Hafner, S. Spielvogel, X. Li, X. Xu, Y. Sun, L. Zhang, Y. Yang, Y. Ma, K. Wesche, H.-F. Graf, C. Leuschner, G. Guggenberger, Y. Kuzyakov, G. Miehe, Foken, T. 2014. Pasture degradation modifies the water and carbon cycles of the Tibetan highlands. – Biogeosciences: discussion forum.

Von Wehrden, H., Wesche, K., Oyundari, Chuluunkhuyag, Fust, P. 2014. Trends in cashmere production and declines of large wild mammals are correlated yet not causally linked – a reply to Berger et al. 2013 – Conservation Biology 29: 286–289.

Seeber, E., Winterfeld, G., Hensen, I., Sharbel, T. F., Durka, W., Liu Jianquan, Yang Yongping, Wesche, K. 2014. Ploidy levels in the alpine sedge Kobresia pygmaea (Cyperaceae) and related species – combined application of chromosome counts, new microsatellite markers and flow cytometry – Botanical Journal of the Linnean Society 176: 22-35.

Krause, B., Wesche, K., Culmsee, H., Leuschner, C. 2014. Diversitätsverluste und floristischer Wandel im Grünland seit 1950 – Natur und Landschaft 89: 399-404.

Meyer, S., Wesche, K., Brütting, T., Krause, B., Hensen, I., Leuschner, C. 2014. Diversitätsverluste und floristischer Wandel im Ackerland seit 1950 – Natur und Landschaft 89: 392-397.

Wesche, K., Schuch, S., Bock, J. 2014. Diversitätsverlust und faunistischer Wandel in ausgewählten Insektengruppen des Grünlandes seit 1950 – Natur und Landschaft 89: 417-421.

Rueff, H., Sternberg, T., Batbuyan Bajav, Middleton, N., Ganzorig, G, Wesche, K. 2014. Pastoral mobility must be preserved to ensure Mongolian rangeland sustainability. Univ. of Oxford, Senckenberg.


Meyer, S., Wesche, K., Krause, B., Leuschner, C. 2013. Dramatic impoverishment of arable plant communities since the 1950s/60s – a large scale analysis across geological substrate groups. – Diversity and Distributions 19: 1175-1187.

Durka, W., Nossol, C., Ruprecht, E., Welk, E., Wagner, V., Wesche, K., Hensen, I. 2013. Extreme genetic depauperation and differentiation of both populations and species in Eurasian feather grasses (Stipa) – Plant Systematics and Evolution 299: 259-269.

Brütting, C., Hensen, I., Wesche, K. 2013. Ex situ cultivation reduces genetic diversity of arable plants. – Plant Biology 15: 505-513.

Leuschner, C., Wesche, K., Meyer, S., Krause, B., Steffen, K., Becker, T., Culmsee, H. 2013. Veränderungen und Verarmung in der Offenlandvegetation Norddeutschlands seit den 1950er Jahren: Wiederholungsaufnahmen in Äckern, Grünland und Fließgewässern – Berichte der Reinhold-Tüxen-Gesellschaft 25: 166-210.

Meyer, S. Wesche, K., Krause, B., Leuschner, C. 2013. Veränderungen der Segetalflora der letzten Jahrzehnte und mögliche Konsequenzen für Agrarvögel – Julius Kühn Archiv 442: 64-78.

Dressler, S., Gregor, T., Otte, V., Paule, J., Ritz, C., Wesche, K., Wesenberg, J. 2013. Forschungen zur heimischen Flora. – Natur, Forschung, Museum 143: 254-255.

Ansorge, H., Wesche, K. 2013. Im Land der Gräser und wilden Pferde. Schlaglichter der Biologischen Forschung in der Mongolei. – Natur, Forschung, Museum 143: 46-51.

Jäger, E.J., Müller, F., Ritz, C.M., Welk, E. & Wesche, K. 2013. Rothmaler – Exkursionsflora von Deutschland. Gefäßpflanzen: Atlasband, 20 edn. Spektrum Akademischer Verlag, Heidelberg, 822 p.

Wesche, K. 2013. Wie verbreitet ist Degradation in tibetischen Weideländern? – Natur, Forschung, Museum 143: 18-23.

Biermann , T with contributions from Seeber, E., Schleuß, P., Willinghöfer, S., Leonbacher,J., Schützenmeister, K., Steingräber, L., Babel, W. Coners, H. Foken, T. Guggenberger, G., Kuzyakov, Y. Leuschner, C. Miehe, G., Wesche K. 2013. Tibet Plateau Atmosphere-Ecology-Glaciology Cluster Joint Kobresia Ecosystem Experiment: Documentation of the 2nd Intensive Observation Period Summer 2012 in KEMA, Tibet. Arbeitsergebnisse Nr. 54, Eigenverlag: Universität Bayreuth, Abt. Mikrometeorologie, Bayreuth, ISSN 1614-8916.

Miehe, G., Miehe, S., Bach, K., Wesche, K., Seeber, E., Behrendes, L., Kaiser, K., Reudenbach, C., Nölling, J., Hanspach, J. Herrmann, M., Ma Yaoming, Mosbrugger, V. 2013. Resilience of vulnerability? Vegetation patterns of the central Tibetan pastoral ecotone. – In: Morales Prieto, M. J., Diaz, J. T. Steppe Ecosystems. Biological diversity, management and restoration. Nova publisher, New York, p. 111-152.


Schuch, S., Bock, J., Krause, B., Wesche, K. Schaefer, M. 2012. Long-term population trends in three grassland insect groups: a comparative analysis of the years 1951 and 2009. – Journal of Applied Entomology 136: 321-331.

von Wehrden, H., Hanspach, J., Kaczensky, P., Fischer, J. & Wesche, K. 2012. A global assessment of the non-equilibrium concept in rangelands. – Ecological Applications 22: 393-399.

Zimmermann, H., von Wehrden, H., Renison, D., Wesche, K., Welk, E., Damascos, M. A., Hensen, I. 2012. Shrub management is the principal driver of differing population sizes between native and invasive populations of Rosa rubiginosa L. – Biological Invasions 14: 2141-2157.

Wagner, V., Treiber, J., Danihelka, J., Ruprecht, E., Wesche, K. 2012. Declining genetic diversity and increasing genetic isolation towards the range periphery of a Eurasian feather grass: A study along a 3000 km longitudinal gradient – International Journal of Plant Sciences 173: 802-811.

Ruppert, J. Holm, A., Miehe, S., Muldavin, E., Snyman, H.A., Wesche, K., Linstädter, A. 2012. Meta-analysis of rain-use efficiency confirms indicative value for degradation and supports non-linear response along precipitation gradients in drylands. – Journal of Vegetation Science: 23: 1035-1050.

Brütting, C., Meyer, S., Kühne, P., Hensen, I., Wesche, K. 2012. Spatial genetic structure and low diversity of the rare arable plant Bupleurum rotundifolium L. in Central Europe – Agriculture, Ecosystems and Environment 161: 70-77.

Hirsch, H., Wypior, C., von Wehrden, H., Wesche, K., Renison, D.,Hensen, I. 2012. Germination performance of native and non-native Ulmus pumila populations. – NeoBiota 15: 53-68.

Wesche, K.,  Treiber, J. 2012. Abiotic and biotic determinants of steppe productivity and performance – a view from Central Asia, In: Werger, M.J.A. & van Staalduinen, M. (eds.) Eurasian Steppes. Ecological problems and livelihoods in a changing world, pp. 3-43. Springer, Heidelberg.

Hensen, I., Cierjacks, K, Hirsch, H., Kessler, M., Romoleroux, K, Renison, D., Wesche, K. 2012. Historic and recent fragmentation coupled with altitude affect the genetic population structure of one of the world’s highest tropical tree line species – Global Ecology and Biogeography 21: 455-464.

Brütting, C., Wesche, K., Meyer, S., Hensen, I. 2012. Genetic diversity of six arable plants in correlation to their Red List status – Biodiversity and Conservation 21: 745-761.

Wesche, K., Krause, B., Culmsee, H., Leuschner, C. 2012. Fifty years of change in Central European grassland vegetation: large losses in species richness and animal-pollinated plants. – Biological Conservation 150: 76-85.

Schuch, S., Wesche, K., Schaefer, M. 2012. Long-term decline in the abundance of leafhoppers and planthoppers (Auchenorrhyncha) in Central European protected dry grasslands. – Biological Conservation: 149: 75-83.

Kohlbrecher, C., Wesche,  K., Hilbig,  W., Leuschner Ch., Meyer St. 2012. Veränderungen in der Segetalflora am Kyffhäusergebirge in den letzten 50 Jahren (1961-2011). – Landschaftspflege und Naturschutz in Thüringen 49: 1-9.


Hafner, S., Unteregelsbacher, S., Seeber, E., Lena, B., Xu, Xingliang, Li, Xiaogang, Guggenberger, G., Miehe, G., Kuzyakov, Y. 2011. Effect of grazing on carbon stocks and assimilate partitioning in a Tibetan montane pasture revealed by 13CO2 pulse labeling. – Global Change Biology 18.: 528–538.

Otte, V., Wesche, K., Dressler, S., Zizka, G., Hoppenrath, M., Kienast, F. 2011. The New Herbarium Senckenbergianum: Old institutions under a new common roof. – Taxon 60: 617-618.

Wagner, V. von Wehrden, H., Wesche, K., Fedulin, A., Sidorova, T., Hensen, I. 2011. Similar performance in peripheral and central populations of Stipa capillata L. despite a different climate and soil pH. – Ecography: 34: 498-506.

Hirsch, H., Zimmermann, H., Ritz, C., Wissemann, V., von Wehrden, H., Renison, D., Wesche, K., Welk, E., Hensen, I. 2011. Tracking the origin of invasive Rosa rubiginosa populations in Argentina. – International Journal of Plant Sciences 172: 530-540.

Krause, B., Culmsee, H., Wesche, K., Bergmeier, E., Leuschner, C. 2011. Habitat loss of floodplain meadows in north Germany since the 1950s. – Biodiversity and Conservation 20: 2347–2364.

Schuch, S., Bock, J., Leuschner, C., Schaefer, M., Wesche, K. 2011. Minor changes in orthopteran assemblages of Central European dry grasslands during the last 40 years. – Journal of Insect Conservation 15: 811-822.

Miehe, G., Miehe, S., Bach, K., Kluge, J., Wesche, K., Yongping, Yang; Jianquan, Liu 2011. Ecological stability during LGM and mid-Holocene in an arid alpine biome indifferent to environmental changes? The Alpine Steppes of the Tibetan highlands – Quaternary Research 76: 243-252.

Miehe, G., Miehe, S., Bach, K., Nölling, J., Hanspach, J., Reudenbach, C., Kaiser, K., Wesche, K., Mosbrugger, V., Yongping, Yang, Yaoming, Ma. 2011. Plant communities of central Tibetan pastures in the Alpine Steppe / Kobresia pygmaea ecotone. – Journal of Arid Environments 75: 711-723.

Miehe, G.; Miehe, S.; Bach, K.; Kluge, J.; Yongping, Yang; Duo, La; Co, Sonam; Wesche, K. 2011. Alpine Steppe Plant Communities of the Tibetan highlands. – Applied Vegetation Science: 14: 547-560.

Sun, D.S., Wesche, K., Chen, D.D., Zhang, S.H., Wu, G.L., Du, G.Z. & Comerford, N.B. 2011. Grazing depresses soil carbon storage through changing plant biomass and composition in a Tibetan alpine meadow. – Plant, Soil and Environment 57: 271-278.

Wesche, K., von Wehrden, H. 2011. Surveying Southern Mongolia: application of multivariate classification methods in drylands with low diversity and long floristic gradients. – Applied Vegetation Science 14: 561-570.

Ronnenberg, K., Hensen I., Wesche K. 2011. Water availability unequally affects seed production and seed viability of Stipa krylovii Roshev. in Mongolian drylands. – Basic and Applied Ecology 12: 141-151.

Wesche, K., Walther, D., von Wehrden, H., Hensen, I. 2011. Trees in the desert: Reproduction and genetic structure of fragmented Ulmus pumila forests in Mongolian drylands. – Flora 206: 91-99.

Ronnenberg, K. & Wesche, K. 2011. Effects of fertilization and irrigation on Stipa steppes in dry southern Mongolia. – Plant and Soil 340: 239-251.

Biermann T, Leipold T. with contributions from Babel W, Becker L, Coners H, Foken T, Guggenberger G, He S, Ingrisch J, Kuzyakov Y, Leuschner C, Miehe G, Richards K, Seeber E, Wesche K. 2011. Joint Kobresia Ecosystem Experiment: Documentation of the first Intensive Observation Period Summer 2010 in Kema, Tibet. Arbeitsergebnisse Nr. 44, Eigenverlag: Universität Bayreuth, Abt. Mikrometeorologie, Bayreuth, ISSN 1614-8916.

Yoseph Assefa, Wesche, K. & Masresha Fetene. 2011. The status of the ericaceous vegetation on the southern slopes of the Bale Mountains. In: Tanskanen, H., Nelson, A. & William, S. (eds.) Conservation of the Bale Mountains. Walia (Special Issue), pp. 158-170, Addis Ababa.      pdf


Hirsch, H.,  von Wehrden, H. , Wesche, K. , Renison, D., Hensen, I. 2010. Investigations on the invasion success of Ulmus pumila L. in North America and Argentina. – Newsletter of the Native Plant Society of New Mexico 35: 14.

Zimmermann, H., Ritz, C. M., Hirsch, H. , Renison, D., Wesche, K., Hensen, I. 2010.  Highly reduced genetic diversity of Rosa rubiginosa L. populations in the invasive range. – International Journal of Plant Sciences 171: 435-446.

Wagner, V. von Wehrden, H., Wesche, K., Fedulin, A., Sidorova, T., Hensen, I. 2010. Similar performance in peripheral and central populations of Stipa capillata L. despite a different climate and soil pH. – Ecography: online first.

Meyer, S., Wesche, K., Leuscher, C., van Elsen, T., Metzner, J. 2010. A new conservation strategy for arable weed vegetation in Germany – the project “100 fields for biodiversity”. – Plant Breeding und Seed Science 61: 25-34.

von Wehrden, H.,  Hanspach, J., Ronnenberg, K. & Wesche, K. 2010. The inter-annual climatic variability in Central Asia – a contribution to the discussion on the importance of environmental stochasticity in drylands. – Journal of Arid Environments 74: 1212-1215.

Meyer, S., Wesche, K., Metzner, J., van Elsen, T., Leuscher, C. 2010. Are current agri-environment schemes suitable for long-term conservation of arable plants? – A short review of different conservation strategies from Germany and short remarks on the new project “100 fields for diversity”. – Aspects of Applied Biology 100: 287-294.

Bläß, C., Ronnenberg, K., Tackenberg, O., Hensen, I., Wesche, K. 2010. Relative importance of different seed dispersal types in dry Mongolian rangelands. – Journal of Arid Environments: 74: 991-997.

Wesche, K., Ronnenberg, K. 2010. Effects of NPK-fertilisation in arid southern Mongolian steppes. – Plant Ecology 207: 93-105.

Wesche, K., Ronnenberg, K., Retzer, V., Miehe, G. 2010. Effects of herbivore exclusion in southern Mongolian desert steppes. – Acta Oecologica 36: 234-241.

Hensen, I., Kilian, C., Wagner, V., Durka, W., Pusch, J., Wesche, K. 2010. Low genetic variability and strong differentiation among isolated populations of the rare steppe grass Stipa capillata L. in central Europe. – Plant Biology 12: 526-536.

Wesche, K., Miehe, G. Kaiser, K. 2010. Wald oder Weide? Zum Einfluss der nomadischen Viehhaltung auf die Grasländer Tibets. In: Herrmann, B. (ed.) Beiträge zum Göttinger Umwelthistorischen Kolloquium 2009 – 2010: 89-108.

Meyer, S., Wesche, K., Leuschner, C., van Elsen, T., Metzner, J. 2010. Schutzbemühungen für die Segetalflora in Deutschland – Das Projekt “100 Äcker für die Vielfalt”. – Treffpunkt Biologische Vielfalt IX – BFN-Skripten 265: 59-64.


(older publications by K. Wesche)


Wesche, K., Krause, B., Culmsee, H., Leuschner, C. 2009. Veränderungen in der Flächen-Ausdehnung und Artenzusammensetzung des Feuchtgrünlandes in Norddeutschland seit den 1950er Jahren. – Berichte der Reinhold-Tüxen-Gesellschaft 21: 196-210.

von Wehrden, H., Hanspach, J., Bruelheide, H., Wesche, K. 2009. Pluralism and diversity – trends in the use and application of ordination methods 1990-2007. – Journal of Vegetation Science 20: 695-705.    pdf

Seltmann, P., Hensen, I., Renison, D., Ploch, S., Dueñas, J. R., Cocucci, A., Wesche, K. 2009. Biparental inbreeding depression, genetic relatedness and progeny vigor in a wind-pollinated treeline species in Argentina – Plant Ecology 205: 155-164.

von Wehrden, H., Zimmermann, H., Hanspach, J., Ronnenberg, K., Wesche, K. 2009. Predicting plant species and communities by using GIS and Landsat data in a southern Mongolian mountain range. – Folia Geobotanica 44: 211-225.

von Wehrden, H., Wesche, K., Miehe, G. 2009. Plant communities of the southern Mongolian Gobi – Phytocoenologia 39: 331-376.

Schiebold, S., Hensen, I., Wesche, K., Röser, M. 2009. Extensive clonality of the endemic Calamagrostis pseudopurpurea ex O.R. Heine in central Germany revealed by RAPD markers. – Plant Biology 11: 473-482.


Bläß, C., Ronnenberg, K., Hensen, I., Wesche, K. 2008. Grazing impact on seed production in southern Mongolia. – Mongolian Journal of Biological Sciences 6: 3-9.

Esfeld, K., Hensen, I., Wesche, K., Jakob, S. S., Tischew, S., Blattner, F. R. 2008. Molecular data indicate multiple independent colonizations of former lignite mining areas in Eastern Germany by Epipactis palustris (Orchidaceae) – Biodiversity and Conservation 17: 2441-2453.

Wesche, K., Cierjacks, A., Yoseph Assefa, Wagner, S., Masresha Fetene, Hensen, I. 2008. Recruitment of trees at tropical-alpine treelines: Polylepis in South America and Erica species in Africa. – Plant Ecology and Diversity 1: 35-46.

Marcora, P., Hensen, I., Renison, D., Seltmann, P., Wesche, K. 2008. Performance of the tree line species Polylepis australis along its entire altitudinal range in Central Argentina. – Diversity and Distributions 14: 630-636.

Ronnenberg K., Wesche K., Hensen, I. 2008. Germination ecology of Central Asian Stipa spp: differences among species, seed provenances, and the importance of field studies. – Plant Ecology 196: 269-280.

Hertel, D., Wesche, K. 2008. Tropical moist Polylepis stands at the treeline in East Bolivia: the effect of elevation on stand microclimate, above- and below-ground structure, and regeneration. – Trees 22: 303-315.     pdf

Cierjacks, A., Salgado, S., Wesche, K., Hensen, I. 2008. Post-fire population dynamics of two tree species in high-altitude Polylepis (Rosaceae) forests of central Ecuador – Biotropica 40: 176-182.

Wesche, K., Yoseph Assefa, von Wehrden, H. 2008. Temperate Grassland Region: Equatorial Africa (high altitude). In: Peart, B. Life in a Working Landscape: Towards a Conservation Strategy for the World’s Temperate Grasslands – Compendium of Regional Templates on the Status of Temperate Grasslands. Conservation and Protection. IUCN / WCPA. Temperate Grasslands Conservation Initiative, Vancouver: 5-21. 

Wesche, K., Retzer, V. 2008. Die Bedeutung von Niederschlag und Beweidung für süd-mongolische Wüstensteppen – ein Beitrag zur Diskussion um die Nicht-Gleichgewichtstheorie. – Hamburger Beiträge zur Physischen Geographie und Landschaftsökologie 18: 165-181.


von Wehrden, H., Wesche, K. 2007. Relationships between climate, productivity and vegetation in southern Mongolian drylands. – Basic and Applied Dryland Research 1: 100-120.

Ronnenberg K., Pietsch M., Wesche K., Hensen I. 2007. Seed germination of five mountain steppe species of Central Asia. – Journal of Arid Environments 71: 404-410.

Miehe, G., Miehe, S., Opgenoorth, L., Cermak, J., Schlütz, F., Jäger, E., Samiya, R., Wesche, K. 2007. Mountain forest islands and Holocene forest retreat in Central Asian deserts. A case study from the South Gobi Altay, Mongolia. – Palaeogeography, Palaeoclimatology, Palaeoecology 250: 155-166.       pdf

Cierjacks, A., Rühr, N., Wesche, K., Hensen, I. 2007. Effects of altitude and livestock on the regeneration of two tree line forming Polylepis species in Ecuador. – Plant Ecology 194: 207-221.     pdf

Wesche, K., Nadrowski, K., Retzer, V. 2007. Habitat engineering under dry conditions: The impact of pikas (Ochotona pallasi) on vegetation and site condition in southern Mongolian steppes. – Journal of Vegetation Science 18: 665-674.    pdf

Cierjacks, A., Wesche, K., Hensen, I. 2007. Impact of sowing, canopy cover and litter on seedling dynamics of Polylepis species at upper treelines in Central Ecuador. – Journal of Tropical Ecology 23: 309-318.

Cierjacks, A., Wesche, K., Hensen, I. 2007. High natural recruitment in forest edges indicates potential lateral expansion of Polylepis fragments in central Ecuador. – Forest Ecology and Management 242: 477-486.

Hensen, I., Wesche, K. 2007. Genetic structure of the rare Poa badensis (Poaceae) in central Germany. – Nova Hedwigia, Beiheft 131: 177-186.

Leyer, I., Wesche, K. 2007. Multivariate Statistik in der Ökologie. Springer-Lehrbuch. Springer Verlag, Berlin, Heidelberg, New York: 221 Seiten.

Stubbe, M., Heidecke, D., Wesche, K., Stubbe, A. 2007. Springmaus, Amurfalke, Biber & Co – Biodiversitätsforschung in Zentralasien – Grundlagen des Internationalen Naturschutzes. – Scientia Halensis 1/07:26-27.

von Wehrden H., Wesche K. 2007. Mapping the vegetation of southern Mongolian protected areas: application of GIS and remote sensing techniques. – Arid Ecosystems 13: 130-139.

Hilbig, W., Wesche, K., Jäger, E. J. 2007. Die Forschungen der Mitarbeiter und Absolventen des Institutes für Geobotanik der Martin-Luther-Universität Halle-Wittenberg in der Mongolei in Zusammenarbeit mit ihren mongolischen Fachkollegen. Erforschung der Biologischen Ressourcen der Mongolei Band 10: 551-568. Martin-Luther-University Halle-Wittenberg, Halle.

von Wehrden, H., Wesche, K. 2007. Mapping Khulan habitats – a GIS-based approach. Erforschung der Biologischen Ressourcen der Mongolei Band 10: 31-44. Martin-Luther-University Halle-Wittenberg, Halle.


von Wehrden H., Wesche K., Tungalag, R. 2006. Plant communities of the Great Gobi B Strictly Protected Area in south-western Mongolia. – Mongolian Journal of Biological Sciences 4: 3-17.

von Wehrden H., Hilbig W., Wesche, K. 2006. Plant communities of the Great Gobi A Special Protected Area in the Mongolian Transaltay Gobi. – Feddes Repertorium 117: 526-570.

Wesche, K., Hensen, I., Undrakh, R. 2006. Range-wide genetic analysis provides evidence for natural isolation among populations of the Central Asian endemic Potentilla ikonnikovii Juz. (Rosaceae). – Plant Species Biology 21: 155-163.

Buhk, C., Götzenberger, L., Wesche, K., Sánchez Gómez, P., Hensen, I. 2006. Post-fire regeneration in Mediterranean pine forest with historically low fire frequency. – Acta Oecologica 30: 288-298.

Wesche, K., Hensen, I., Undrakh, R. 2006. Genetic structure of Galitzkya macrocarpa and G. potaninii, two closely related endemics of central Asian high mountain ranges. – Annals of Botany 98: 1025-1034.     pdf

Wesche, K., Pietsch, M., Ronnenberg, K., Undrakh, R., Hensen, I. 2006. Germination of fresh and frost-treated seeds from dry Central Asian steppes. – Seed Science Research 16: 123-136.

von Wehrden, H., Wesche, K., Reudenbach, C., Miehe G. 2006. A LANDSAT 7-based vegetation map of the deserts and desert steppes in the Gobi Gurvan Saikhan NP, southern Mongolia (scale 1: 200 000). – Erdkunde 60: 261-272.  pdf

Hensen, I., Wesche, K. 2006. Population size affects plant fitness in the rare Dictamnus albus in Central Germany. – Biodiversity and Conservation 15: 2249-2261.

Wesche, K. 2006. Is afroalpine plant biodiversity negatively affected by high-altitude fires? In: Spehn, E., Liberman, M., Körner, C. (Hrsg.). Land Use Change and Mountain Biodiversity. CRC Press, Boca Raton FL, USA: 39-49.

Bläß, K., Ronnenberg, K., Wesche, K. 2006. Exozoochorous seed dispersal: seed retention under experimental natural conditions in Mongolian steppes. In: Shmakov, A. I., Kamelin, R. V., Terekhina, T. A., Dyachenko, S. A., Smirnov, S. V., Kuzev, M. G., German, D. A. Problems of Botany of South Siberia and Mongolia. “Azbuka” Press, Barnaul: 329-331.

Ronnenberg, K., Wesche, K. 2006. The effects of NPK-fertilization and irrigation on Stipa krylovii and S. gobica in southern Mongolian desert steppes. In: Shmakov, A. I., Kamelin, R. V., Terekhina, T. A., Dyachenko, S. A., Smirnov, S. V., Kuzev, M. G., German, D. A. Problems of Botany of South Siberia and Mongolia. “Azbuka” Press, Barnaul: 332-334.



Wesche, K., Jäger, E., von Wehrden, H., Undrakh, R. 2005. The distribution of some rare plants in the Gobi-Altay. – Mongolian Journal of Biological Sciences 3: 3-11.

Dittbrenner, A., Hensen, I., Wesche, K. 2005. Genetic structure and RAPD diversity of the rapidly declining Angelica palustris (Apiaceae) in Eastern Germany in relation to population size and seed production. – Plant Species Biology 20: 191-200.

Wesche, K., Miehe, S., Miehe, G. 2005. Plant communities of the Gobi Gurvan Sayhan National Park (South Gobi Aimag, Mongolia). – Candollea 60: 149-205.

Wesche, K., Partzsch, M., Krebes, S., Hensen, I. 2005. Effects of habitat size, age, and nutrients in fragmented grassland communities. – Folia Geobotanica 40: 341-356.

Stumpp, M., Wesche, K., Retzer, V., Miehe, G. 2005. Impact of grazing livestock and distance from water points on soil fertility in southern Mongolia. – Mountain Research and Development 25: 244-251.    pdf

Sommerwerk, A., Partzsch, M., Wesche, K., Hensen, I. 2005. Standortökologie und phytozönologische Bindung der beiden Adventivarten Gypsophila perfoliata und G. scorzonerifolia (Caryophyllaceae) in Zentraldeutschland. – Hercynia 38: 263-286.

Wesche, K., Ronnenberg, K., Hensen, I. 2005. Lack of sexual reproduction in dry mountain steppe populations of the clonal shrub Juniperus sabina L. in southern Mongolia. – Journal of Arid Environments 63: 390-405.

Hensen, I., Oberprieler, C., Wesche, K. 2005. Genetic structure, population size, and seed production of Pulsatilla vulgaris Mill. (Ranunculaceae) in Central Germany. – Flora 200: 3-14.

Ronnenberg, K., Wesche, K. 2005. Clonal versus sexual reproduction – examples for complementary regeneration strategies in a dry ecosystem. In: Dorjsuren, Ch., Dorofeyuk, N.I., Gunin, P.D., Drobyshev, Yu. I., Bazha, S.N., Vasilieva, L.F. (Hrsg.) Ecosystems of Mongolia and frontier areas of adjacent countries: natural resources, biodiversity and ecological prospects. – Publishing House “Bembi San”, Ulaanbaatar: 143-145.

von Wehrden, H., Wesche, K. 2005. Mapping the vegetation of southern Mongolian protected areas: application of GIS and remote sensing techniques. In: Dorjsuren, Ch., Dorofeyuk, N.I., Gunin, P.D., Drobyshev, Yu. I., Bazha, S.N., Vasilieva, L.F. (Hrsg.) Ecosystems of Mongolia and frontier areas of adjacent countries: natural resources, biodiversity and ecological prospects. – Publishing House “Bembi San”, Ulaanbaatar: 232-234.

Wesche, K. 2005. Enclosure studies indicate non-equilibrium dynamics in southern Mongolian rangelands In: Dorjsuren, Ch., Dorofeyuk, N.I., Gunin, P.D., Drobyshev, Yu. I., Bazha, S.N., Vasilieva, L.F. (Hrsg.) Ecosystems of Mongolia and frontier areas of adjacent countries: natural resources, biodiversity and ecological prospects. – Publishing House “Bembi San”, Ulaanbaatar: 198-200.

Leyer, I., Wesche, K. 2005. Ordinationsmethoden zur Analyse ökologischer Daten. In: Grözinger, G., Matiaske, W. (Hrsg.) Deutschland regional – Sozialwissenschaftliche Daten im Forschungsverbund. – Rainer Hamp Verlag, München: 157-170.

Wesche, K., Stubbe, A., Samjaa, R., Retzer, V., Miehe, G., Stubbe, M. 2005. Herausgabe Erforschung der Biologischen Ressourcen der Mongolei 9. – Martin-Luther-Universität Halle-Wittenberg, Halle: 1-494. ISSN 0440-1298 Darin:

Wesche, K., Retzer, V. 2005. Is degradation a major problem in semi-desert environments of the Gobi region in southern Mongolia? In: Wesche, K., Stubbe, A., Samjaa, R., Retzer, V., Miehe, G., Stubbe, M. (Hrsg.) Erforschung der Biologischen Ressourcen der Mongolei Band 9: 133-146.

Miehe, G., Samjaa, R., Wesche, K. 2005. Preface: Ecosystem research in the arid environments of Central Asia: Results, challenges, perspectives. In: Wesche, K., Stubbe, A., Samjaa, R., Retzer, V., Miehe, G., Stubbe, M. (Hrsg.) Erforschung der Biologischen Ressourcen der Mongolei Band 9: 1-3.



Wesche, K., Ronnenberg, K. 2004. Phytosociological affinities and habitat preferences of Juniperus sabina L. and Artemisia santolinifolia Turcz. ex Bess. in mountain sites of the south-eastern Gobi Altay, Mongolia. – Feddes Repertorium 115: 585-600.

Schulze, H., Hensen, I., Wesche, K., Stordeur, R. 2004. Die Verbreitung epigäischer Flechten und ihre Beziehungen zu ausgewählten Phanerogamengesellschaften im NSG Harslebener Berge und Steinholz (Sachsen-Anhalt). – Hercynia 37: 73-85.

Wesche, K., Kessler, M. 2004. Die humide tropisch-alpine Stufe – weltweite Gemeinsamkeiten. In: Breckle, S.W., Walter, H. (Hrsg.). Ökologie der Erde. Bd. 2. Spezielle Ökologie der tropischen und subtropischen Zonen. – Spektrum, Heidelberg: 129-156.

Götzenberger, L., Ohl, C., Hensen, I., Sánchez Gómez, P., Wesche, K. 2003. Postfire regeneration of a thermomediterranean shrubland area in south-eastern Spain. – Anales de Biología 25:21-28.    pdf

Wesche, K. 2003. The importance of occasional droughts for afroalpine landscape ecology. – Journal of Tropical Ecology 19: 197-208.

Wesche, K. 2002. Structure and dynamics of Erica forest at tropical-African treelines. – Verhandlungen der Reinhold-Tüxen Gesellschaft 14: 145-159.

Ochyra, R., Wesche, K., Miehe, G., Miehe, S. 2002. New records of pleurocarpous mosses for Africa and Uganda. – Journal of Bryology 24: 251-258.

Wesche, K. 2002. The high-altitude environment of Mt. Elgon (Uganda/Kenya) – Climate, vegetation and the impact of fire. – Ecotropical Monographs 2: 1-253.  

Wesche, K. 2001. Hochgebirgslebensräume am Mt. Elgon (Uganda/Kenia) – Klima, Vegetation und die Bedeutung von Feuern. – Jahrbuch der Marburger Geographischen Gesellschaft 2001: 172-178.

Wesche, K. 2001. The high altitude vegetation of Mt. Elgon/Kenya as influenced by climate and fire. – Proceedings of the Workshop for Integrated Natural Resources Manage­ment Planning – Mt. Elgon, Kitale – IUCN, Kenya: 22-29. 

Wesche, K., Miehe, G., Kaeppeli, M. 2000. The significance of fire for the ericaceous belt of the East African Mountains. – Mountain Research and Development 20: 340-347.     pdf

Wesche, K., Clausnitzer, V., Miehe, S., Miehe, G. 2000. Habitat conditions in afroalpine communities – examples from East Africa. In: Breckle, S.-W., Schweizer, B., Arndt, U. (Hrsg.) Results of worldwide ecological studies – Proceedings of the 1st symposium of the A.F.W. Schimper-Foundation est. by H. & E. Walter – Hohenheim, October 1998. – G. Heimbach, Stuttgart: 335-346. 

Wesche, K. 1997. A classification of a tropical Shorea robusta forest stand in southern Nepal. – Phytocoenologia 27: 103-118.

Clausnitzer, V., Wesche, K. 1996. Odonata records from Nepal (around Annapurna, Chitwan District, Royal Bardia National Park). – Opuscula Zoologica Fluminensia 147: 1-8.

Wesche, K. 1995. Struktur und Nutzung eines Sal-Waldes im Süden Nepals. – GTZ/TÖB, Eschborn. 58 Seiten. 

Wesche, K. 1995. Structure and use of a Sal-forest in southern Nepal. – GTZ/TÖB, Eschborn. 35 Seiten.



Schmidt, L. Jahn, A. 2015. Die Blumen des Jahres – besprochen von K. Wesche in Berichte der Naturforschenden Gesellschaft der Oberlausitz 2015.

Huneck, S. 2010. 10000 Kilometer unterwegs im Herzen Asiens – Expeditionsberichte aus der Mongolei. – besprochen von K. Wesche in Hercynia 2012.

Latif, M. 2009. Klimawandel und Klimadynamik. – besprochen von K. Wesche in Hercynia.

Wittig, R., Streit, R. 2004. Ökologie. – besprochen von K. Wesche in Hercynia.

Kriechbaum, M. 2002. Flora, Vegetation und Landnutzung des Muktinath-Tales (Mustang, Nepal) als Beziehungsmuster von naturräumlicher Ausstattung und menschlicher Gestaltung im Zentralhimalaya. – besprochen von K. Wesche in Feddes Repertorium.

Succow, M., Joosten, H. (Hrsg.) 2002. Landschaftsökologische Moorkunde. – besprochen von K. Wesche in Feddes Repertorium.

Munk, K. 2001. Grundstudium Biologie – Botanik. – besprochen von K. Wesche in Wissen­schaft Online.

Riedel, W., Lange, H. (Hrsg.) 2001. Landschaftsplanung. – besprochen von K. Wesche in Wissenschaft Online.

Baumann, H., Müller, T. 2001. Farbatlas geschützte und gefährdete Pflanzen – besprochen von K. Wesche in Hercynia.

Porembski, S., Barthlott, W. (Eds.) 2000. Inselbergs. Biotic Diversity of Isolated Rock Out­crops in Tropical and Temperate Regions. – besprochen von K. Wesche in Flora.

Osmaston, H., Tukahirwa, J., Basalirwa, C., Nyakaana, J. (editors) 1998. The Rwenzori Mountains National Park, Uganda. – besprochen von K. Wesche in Mountain Re­search and Development.

Fischer, E. 1996. Die Vegetation des Parc National de Kahuzi-Biéga, Sud-Kivu, Zaire. – be­sprochen von K. Wesche in Mountain Research and Development.