Dr. Moritz Sonnewald
Scientific Associate

Research foci: Biodiversity, ecology and zoology of marine organisms, zoological monitoring projects, focus on North Sea and Red Sea epi- and endobenthos,

zoogeography of fish, in particular neozoans and their immigration routes

since 2018 scientific associate, section Ichthyology

2015-2018 project manager and coordinator Red Sea Biodiversity Surveys

2014-2015 project coordinator Red Sea Biodiversity Surveys

2012-2014 Research Associate in section Crustacea, working on the GBIF-D project (Germany, BMBF)

2012 PhD in Biology, Goethe University, Frankfurt am Main

Selected publications

Gries L, Paule J, Sonnewald M & Nilsson MA (2023). Eocene origin of anemone-carrying behaviour in polydectine crabs (Brachyura: Xanthidae). Zoological Journal of the Linnean Society, (May), 1–8. https://doi.org/10.1093/zoolinnean/zlad075

Winter S, …, Sonnewald M, … & Janke, A. (2023). A chromosome-scale reference genome assembly of the great sand eel, Hyperoplus lanceolatus. Journal of Heredity, 114(2), 189–194. https://doi.org/10.1093/jhered/esad003

Hahn SJ, Brandt A & Sonnewald M (2022). Annotated checklist and biodiversity analysis of benthic fauna at Sylt Outer Reef and Borkum Reef Ground (North Sea). Check List, 18(3), 593–628. https://doi.org/10.15560/18.3.593

Winter S, …, Sonnewald M, … & Janke, A. (2020). Chromosome-level genome assembly of a benthic associated Syngnathiformes species: the common dragonet, Callionymus lyra. Gigabyte, 2020, 1–10. https://doi.org/10.46471/gigabyte.6

Sonnewald M, & Al-Aidaroos, AM (2018). The Red Sea Biodiversity Project : Collection , understanding and preservation of the marine fauna of Saudi Arabia, (May), 4–5.

Al-Aidaroos AM, Kumar AAJ, Al-Haj AE, & Sonnewald M (2017). A new species of Schizophrys (Majoidea: Majidae: Majinae) from the Red Sea coast of Saudi Arabia, with description of its first zoea. Marine Biodiversity, 47(4), 1163–1169. https://doi.org/10.1007/s12526-017-0800-6

Bowler DE, …, Sonnewald M. et al. (2017) Cross-realm assessment of climate change impacts on species’ abundance trends. Nature Ecology and Evolution 1:67. DOI 10.1038/s41559-016-0067

Sonnewald M, Türkay M (2017) Composition of the Epibenthic Decapod Crustacean Megafauna of the German Exclusive Economic Zone: Comparison and Analysis of Past and Recent Surveys. Crustaceana 90: 1193–1210. DOI 10.1163/15685403-00003642
Schwinn, M, Sonnewald M, Türkay M (2014) Decapod Fauna of the Helgoland Trench (Crustacea). A Long Term Study in a Biodiversity Hotspot. Marine Biodiversity 44: 491-517. DOI 10.1007/s12526-014-0217-4

Sonnewald M, Türkay M (2012) Abundance analyses of mega-epibenthic species on the Dogger Bank (North Sea): diurnal rhythms and short-term effects caused by repeated trawling, observed at a permanent station. Journal of Sea Research 73:1-6. DOI 10.1016/j.seares.2012.05.015

Sonnewald M, Türkay M (2011) Environmental Influence on the Megaepifauna Communities of the Dogger Bank: A long-term Survey. Helgoland Marine Research 66(4):503-511. DOI 10.1007/s10152-011-0286-8

Sonnewald M, Türkay M (2011) The megaepifauna of the Dogger Bank (North Sea): species composition and faunal characteristics 1991–2008. Helgoland Marine Research 66(1): 63-75. DOI 10.1007/s10152-011-0247-2

Dr. Friedhelm Krupp
Curator, retired
Honorary Research Associate


Taxonomy, systematics and biogeography of Indo-Pacific fishes with a focus on the Arabian Seas Region

Taxonomy, systematics, phylogeny and biogeography of Middle Eastern freshwater fishes

Marine environmental research and management

Planning and management of marine protected areas


1987-1991, 2002-2011 and since 2017: Head of Fish Section, Senckenberg Research Institute, Frankfurt a. M., Germany (periods in between released for service in international organizations)

2006-2008 Coordinator of the “Middle Eastern Biodiversity Network”, funded by the German Academic Exchange Service (DAAD)

2008-2010 Senckenberg Biodiversity and Climate Research Centre (BiK-F), Frankfurt a. M.: team leader tropical marine ecosystems

2011-2017: Director of the Qatar Museum of Nature and Science, based in Doha, Qatar

2012-2015: UNESCO World Heritage Committee, Member of Qatari Delegation, specializing in Natural World Heritage

2002 World Bank, Washington D.C.: Evaluator of marine protected area projects in the Indian Ocean, funded by the Global Environment Facility (GEF)

1998-2002: United Nations Development Program (UNDP), New York: Chief Technical Advisor of the “Strategic Action Programme for the Red Sea and Gulf of Aden”, based in Jeddah, Saudi Arabia

1998 Environmental Authority, Dubai Municipality, UAE: planning marine protected areas in Dubai

1997-1999 with UNDP/UNOPS-GEF project “Conservation and Sustainable Use of Biodiversity of Socotra Archipelago, Yemen”, team leader marine surveys

1996-1998 UNDP-GEF Chief Technical Consultant of the project “Strategic Action Programme for the Red Sea and Gulf of Aden”, based in Jeddah, Saudi Arabia

1991-1995 Manager of the project “Establishment of a Marine Habitat and Wildlife Sanctuary for the Gulf Region”, supervising a team of 80 scientists; activities included assessment of the environmental effects of the Gulf War oil spill, development of environmental restoration methods, and the establishment and management of a large marine protected area; funded by the EU; based in Jubail, Saudi Arabia

1986 Institute for Ancient Middle Eastern Archaeology, University of Berlin, Germany; planned and managed zoological component of interdisciplinary archaeological project in north-eastern Syria

1982-1985 research assistant, University of Tübingen, Germany; zoological research in interdisciplinary project “Tübingen Atlas of the Middle East” (DFG Sonderforschungsbereich 19)


1985: Dr. rer. nat. in zoology, Johannes-Gutenberg-University, Mainz, Germany

1982: Diploma in biology, University of Mainz, Germany

1980: Studies of Arabic language and literature, University of Jordan, Amman, Jordan

1975-1982: Student of biological and geological sciences, and Middle Eastern studies; University of Mainz, Germany; scholarship holder of German National Merit Foundation

About 160 publications, co-editor of 25 book volumes, editorial board member of five scientific journals

Since 1984: co-editor/Editor-in-Chief of the continuous book series “Fauna of Arabia“

2006-2009: scientific editor of “Aqua, International Journal of Ichthyology”



Zajonz, U., Lavergne, E., Bogorodsky, S.V., Krupp, F., Saeed, F.N. & Aideed, M.S. (in press). Coastal fish diversity of the Socotra Archipelago, Yemen. Zootaxa.

Buchanan, J.R., Ralph, G.M., Krupp, F., Harwell, H., Abdallah, M., Abdulqader, E., Al-Husaini, M., Bishoph, J.M., Burt, J.A., Choat, J.H., Collette, B.B., Feary, D.A., Hartmann, S.A., Iwatsuki, Y., Kaymaram, F., Larson, H.K., Matsuura, K., Motomura, H., Munroe, T., Russell, B., Smith-Vaniz, W., Williams, J. & Carpenter, K.E. (2019). Regional extinction risks for marine bony fishes occurring in the Persian/Arabian Gulf. Biological Conservation 230: 10-19; doi: 10.1016/j.biocon.2018.11.027

Krupp, F. (2018). In Memoriam Prof. Dr. Wolfgang Klausewitz (1922-2018). Aqua, International Journal of Ichthyology 24 (3): 139-140.

Alwan, N.H., Esmaeili, H.R. & Krupp, F. (2016). Molecular phylogeny and zoogeography of the Capoeta damascina species complex (Pisces: Teleostei: Cyprinidae). PLoS ONE 11(6): 1 – 25; e0156434. doi: 10.1371/journal.pone.0156434.

Buchanan, J.R., Krupp, F., Burt, J.A., Feary, D.A., Ralph, G.M., Carpenter, K.E. (2016). Living on the edge: Vulnerability of coral-dependent fishes in the Gulf. Marine Pollution Bulletin. doi: 10.1016/j.marpolbul.2015.11.033

Lavergne, E., Zajonz, U., Krupp, F., Naseeb, F. & Aideed, M.S. (2016). Diversity and composition of estuarine and lagoonal fish assemblages of Socotra Island, Yemen. Journal of Fish Biology 88: 2004–2026; doi: 10.1111/jfb.12964.

Zajonz, U., Lavergne, E., Klaus, R., Krupp. F., Sheikh, M.A. & Naseeb F.S. (2016). The coastal fishes and fisheries of the Socotra Archipelago, Yemen. Marine Pollution Bulletin 105 (2) 660-675. doi: 10.1016/j.marpolbul.2015.11.025.

Krupp, F., Almuftah, A.M.H., Carpenter, K.E., Buchanan, J.R. & Ralph, G.M. (2015). How do the Arabian Gulf’s fish assemblages respond to multiple stressors? Qatar University Life Science Symposium_QULSS 2015. Global Changes: The Arabian Gulf Ecosystem. Q-Science.com. doi: 10.5339/qproc.2015.qulss2015.8

Borkenhagen, K. & Krupp, F. (2013). Taxonomic revision of the genus Carasobarbus Karaman, 1971 (Actinopterygii, Cyprinidae). ZooKeys 339: 1–53. doi: 10.3897/zookeys.339.4903

Randall, J.E., Bogorodsky, S.V., Krupp, F., Rose, J.M. & Fricke, R. (2013). Epinephelus geoffroyi (Klunzinger, 1870) (Pisces: Serranidae), a valid species of grouper endemic to the Red Sea and Gulf of Aden. Zootaxa 3641 (5): 524-532.

Krupp, F., Al-Jumaily, M., Bariche, M., Khalaf, M., Malek, M., Streit, B. (2009). The Middle Eastern Biodiversity Network: Generating and sharing knowledge for ecosystem management and conservation. ZooKeys 31: 3-15.

Krupp, F. & Budd, K. (2009): A new species of the genus Garra (Teleostei: Cyprinidae) from Oman. Aqua, International Journal of Ichthyology 15(2): 117-120.

Krupp, F., Musselman, L.J., Kotb, M.M.A. & Weidig, I. (2009). The effects of climate change on biodiversity: pressing issues and research priorities. BioRisk – Biodiversity and Ecosystem Risk Assessment 3: 1-4.

Krupp, F. & Zajonz, U. (2009). Erforschung und Management der biologischen Vielfalt des Sokotra Archipels. Jemen-Report 40(2): 49-54.

Krupp, F., Zajonz, U. & Khalaf, M.A. (2009). A new species of the deepwater cardinalfish genus Epigonus (Perciformes: Epigonidae) from the Gulf of Aqaba, Red Sea. Aqua, International Journal of Ichthyology 15(4): 223-227.

Khalaf, M. & Krupp, F. (2008): A new species of the genus Symphysanodon (Perciformes: Symphysanodontidae) from the Gulf of Aqaba, Red Sea. Aqua, International Journal of Ichthyology 14(2): 85-88.

Krupp, F. & Schneider, W. (2008): Die Fischfauna des Nahr al-Khabur, Nordost-Syrien. In: Kühne, H. (ed.): Umwelt und Subsistenz der assyrischen Stadt Dur-Katlimmu am unteren Khabur 8: 41-51. Harrassowitz-Verlag, Wiesbaden.

Krupp, F., Zajonz, U. (2008). Korallenriffe im Klimawandel. Natur und Museum 138 (7/8): 163-177.

Abuzinada, A.H., Barth, H.-J., Krupp, F., Böer, B. & Abdelsalaam T., eds (2007). Protecting the Gulf’s Marine Ecosystems from Pollution. VIII + 279 pp. Birkhäuser Verlag, Switzerland.

Jones, D.A., Hayes, M., Krupp, F., Sabatini, G., Watt, I. & Weishar, L. (2007). The impact of the Gulf War (1990-91) oil release upon the intertidal Gulf coast line of Saudi Arabia and subsequent recovery. In: Abuzinada, A.H., Barth, H.-J., Krupp, F., Böer, B. & Abdelsalaam T. (eds): Protecting the Gulf’s Marine Ecosystems from Pollution:237-254. Birkhäuser Verlag, Switzerland.

Krupp, F. & Abuzinada, A. H. (2007). Impact of oil pollution and increased sea surface temperatures on marine ecosystems and biota in the Gulf. In: Abuzinada, A.H., Barth, H.-J., Krupp, F., Böer, B. & Abdelsalaam T. (eds): Protecting the Gulf’s Marine Ecosystems from Pollution: 45-56. Birkhäuser Verlag, Switzerland.

Krupp, F., Al-Muftah, A. Jones, D.A. & Hoolihan, J. (2006). Marine and coastal ecosystem management requirements in the Arabian Peninsula with special regard to water resources. Policy Perspectives for Ecosystem and Water Management in the Arabia Peninsula. UNESCO, Doha & United Nations University, Hamilton, Ontario, pp. 73-87.

Krupp, F., Apel, M., Hamoud, A., Schneider, W. & Zajonz, U. (2006): Zoological survey in the Red Sea coastal zone of Yemen. Fauna of Arabia 21: 11-32.

Kotb, M., M. Abdulaziz, Z. Al-Agwan, K. Alshaikh, A., H. Al-Yami, A. Banajah L. Devantier, M. Eisinger, M. Eltayeb, M. Hassan, G. Heiss, S. Howe, J. Kemp, R. Klaus, F. Krupp, N. Mohamed, A. Rouphael, J. Turner & U. Zajonz (2004): Status of Coral Reefs in the Red Sea and Gulf of Aden in 2004. In C. Wilkinson (Ed.): Status of Coral Reefs of the World: 2004. Vol. 1: 137-154. Australian Institute of Marine Science.

Krupp, F. (2004). Droht Haien die weltweite Ausrottung? Natur und Museum. 134(2): 33-44

Gladstone, W., Krupp, F. & Younis, M. (2003). Development and management of a network of marine protected areas in the Red Sea and Gulf of Aden region. Ocean and Coastal Management 46: 741-761.

Hariri, K.I., Nichols, P., Krupp, F., Mishrigi, S., Barrania, A., Ali, A.F. & Kedidi, S.M. (2002). Status of the Living Marine Resources in the Red Sea and Gulf of Aden Region and their Management. 134 pp. The World Bank. Washington D.C.

Krupp, F. (2002). Marine protected areas.- In: The Gulf Ecosystem, Health and Sustainability. N.Y. Khan, M. Munawar & A.R.G. Price (eds.). Ecovision World Monograph Series 14: 447-473. Backhuys Publishers, Leiden

Krupp, F., Apel, M. & Zajonz, U. (2002). Meeresbiologische Untersuchungen im Sokotra-Archipel (Jemen).- Natur und Museum 132(8). 273-294.

Price, A.R.G., Jones, D.A. & Krupp, F. (2002). Biodiversity. In: The Gulf Ecosystem, Health and Sustainability. N.Y. Khan, M. Munawar & A.R.G. Price (eds.). KISR, Kuwait: 105-123.

Krupp, F., Almarri, M., Zajonz, U., Carpenter, K., Almatar, S. & Zetzsche, H. (2000.). Twelve new records of fishes from the Gulf. Fauna of Arabia 18: 323-335.

Gladstone, W.; Tawfiq, N.; Nasr, D.; Andersen, I.; Cheung, C.; Drammeh, H., Krupp, F. & Lintner, S. (1999). Sustainable use of renewable resources and conservation in the Red Sea and Gulf of Aden: issues, needs and strategic actions. Ocean and Coastal Management. 42: 671-697.

Carpenter, K.; Krupp, F.; Jones, D.J. & Zajonz, U. (1997). FAO Species Identification Guide for Fishery Purposes. The Living Marine Resources of Kuwait, Eastern Saudi Arabia, Bahrain, Qatar and the United Arab Emirates. 311 pp. FAO, Rome.

Compagno, L.J.V., Krupp, F. & Carpenter, K.E. (1996). A new weasel shark of the genus Paragaleus from the northwestern Indian Ocean and the Arabian Gulf (Carcharhiniformes: Hemigaleidae).- Fauna of Saudi Arabia, 15: 391-401.

Krupp, F. & Al-Marri, M. (1996). Fishes and fish assemblages of the Jubail Marine Wildlife Sanctuary. In: Krupp, F., Abuzinada, A.H. & Nader, I.A., eds: A Marine Wildlife Sanctuary for the Arabian Gulf – Environmental Research and Conservation Following the 1991 Gulf War Oil Spill: 339-350.

Fischer, W., Krupp, F., Schneider, W., Sommer, C. Carpenter, K.E. & Niem, V.H. eds (1995). Guia FAO para la identificación de especies para los fines de la pesca. Pacífico centro-oriental. 3 vols, 1813 pp. FAO, Rome.

Coad, B.W. & Krupp, F. (1994). Capoeta aculeata (Valenciennes in Cuv. & Val., 1844), a valid species of cyprinid fish from Iran (Teleostei: Cyprinidae). Zoology in the Middle East 10: 63-72.

Krupp, F. & Müller, T. (1994). The status of fish populations in the northern Arabian Gulf two years after the 1991 Gulf War oil spill. Courier Forschungsinstitut Senckenberg 166: 67-75.

Krupp, F. & Jones, D.A. (1993). The creation of a marine sanctuary after the Gulf War oil spill. Marine Pollution Bulletin 27: 315-323.

Klausewitz, W., Krupp, F. & Köhler, C. (1992). Sektion Ichthyologie I. 175 Jahre SNG, Jubiläumsband II: 42-50.

Krupp, F. (1992). A new species of Nemacheilus (Pisces: Balitoridae) from the Van Gölü drainage basin in eastern Turkey. Mitteilungen aus dem hamburgischen zoologischen Museum und Institut 89: 291-294.

Krupp, F. (1992). Two new species of cyprinid fishes from the Mediterranean coastal drainage basin of Syria (Pisces: Osteichthyes: Cyprinidae). Senckenbergiana biologica 72 (1/3): 19-25.

Krupp, F. (1992). The karst springs of Ra’s al-Ain – a freshwater community in the Mesopotamian desert. Aqua geographica 1(1): 26-33.

Krupp, F., Al-Hassan, L.A.J. & Ziegler, T. (1992). A possible natural hybrid of Acanthobrama marmid and Alburnus mossulensis from Haur al-Hammar, southern Iraq (Pisces: Osteichthyes: Cyprinidae). Senckenbergiana biologica 72 (4/6): 219-223.

Krupp, F. & Moubayed, J. (1992). A new species of Cobitis Linnaeus 1758 from the Orontes and Litani drainage basins of Syria and Lebanon (Pisces: Osteichthyes: Cobitidae). Senckenbergiana biologica 72 (1/3): 13-18.

Krupp, F. (1991). Die Fauna des Persisch/Arabischen Golfes und seiner Küsten. Natur und Museum 121 (4): 97-110.

Krupp, F. & Paulus, T. (1991). First record of the coral reef fish Pseudanthias fasciatus (Kamohara, 1954) from the Red Sea (Perciformes: Serranidae). Fauna of Saudi Arabia 12: 388-392.

Krupp, F. & Paulus, T. (1991). Territoriality and courtship behaviour in the coral reef fish Pseudanthias heemstrai (Pisces: Serranidae). Revue française d’Aquariologie 18: 43-46.

Krupp, F. & Schneider, W. (1991). Two new species of Nemacheilus Bleeker 1863 from the Orontes River drainage basin of Lebanon, Syria and Turkey (Pisces: Osteichthyes: Balitoridae). Senckenbergiana biologica 71 (1990) (1/3): 23-34.

Krupp, F. & Schneider, W. (1991). Bestandserfassung der rezenten Fauna im Bereich des Nahr al-Habur. In: Kühne, H. (ed.). Die rezente Umwelt von Tall Seh Hamad und Daten zur Umweltrekonstruktion der assyrischen Stadt Dur-Katlimmu 1: 69-85.

Krupp, F. & Schneider, W. (1991). Artdetermination und Bewertung ökologischer und erdgeschichtlicher Faktoren bei der Darstellung zoologischer Verbreitungsmuster. In: Röllig, W. (ed.). Von der Quelle zur Karte. VCH Verlag, Weinheim: 65-76.

Krupp, F. & Debelius, H. (1990). The hybrid of Centropyge multifasciatus x Holacanthus venustus from the Philippines and notes on aberrant colour forms of Centropyge multispinis from the Maldives and the Red Sea. Revue française d’Aquariologie 17 (2): 53-56.

Krupp, F. & Schneider, W. (1989). The fishes of the Jordan River drainage basin and Azraq Oasis. Fauna of Saudi Arabia 10: 347-416.

Schuhmacher, H., Krupp, F. & Randall, J.E. (1989). Pseudanthias heemstrai, a new species of anthiine fish (Perciformes: Serranidae) from the Gulf of Aqaba, Red Sea. Fauna of Saudi Arabia 10: 338-346.

Krupp, F. (1988). Freshwater fishes of the Wadi Batha drainage. Journal of Oman Studies, special report no. 3: 401-404.

Krupp, F. & Schneider, W. (1988). Die Süßwasserfauna des Vorderen Orients – Anpassungsstrategien und Besiedlungsgeschichte einer zoogeographischen Übergangszone. Natur und Museum 118 (7): 193-213.

Mir, S., Al-Absy, A. & Krupp, F. (1988). A new natural intergeneric cyprinid hybrid from the Jordan River drainage, with a key to the large barbine cyprinids of the southern Levant. Journal of Fish Biology 32: 931-936.

Krupp, F. (1987). Tiefenwasser- und Tiefseefische aus dem Roten Meer. XV. The occurrence of Cynoglossus acutirostris Norman 1939 in the Red Sea (Pisces: Cynoglossidae)- Senckenbergiana maritima 19 (3/4): 249-259.

Krupp, F. (1987). Freshwater ichthyogeography of the Levant. In: Krupp, F., Schneider, W. & Kinzelbach, R. (eds). Proceedings of the Symposium on the Fauna and Zoogeography of the Middle East, Mainz 1985. Beihefte zum Tübinger Atlas des Vorderen Orients, A 28: 229-237.

Krupp, F. (1985). A new species of Chondrostoma from the Orontes river drainage basin of Turkey and Syria (Pisces: Osteichthyes: Cyprinidae). Senckenbergiana biologica 66 (1/3): 27-33.

Krupp, F. (1985). Barbus chantrei (Sauvage 1882), a valid species of cyprinid fish from the northern Levant (Pisces: Osteichthyes: Cyprinidae). Senckenbergiana biologica 66 (1/3): 17-25.

Krupp, F. (1985). Barbus continii Vinciguerra 1926, a possible natural hybrid of Barbus canis and Barbus longiceps (Pisces: Osteichthyes: Cyprinidae). Senckenbergiana biologica 66 (1/3): 9-15.

Krupp, F. (1985). Rehabilitation of Barbus lorteti Sauvage, 1882, and comments on the validity of the generic names Bertinius Fang, 1943, and Bertinichthys Whitley, 1953 (Pisces: Cyprinidae).- Hydrobiologia 120: 63-68.

Villwock, W., Scholl, A. & Krupp, F. (1983). Zur Taxonomie, Verbreitung und Speziation des Formenkreises Aphanius dispar (Rüppell, 1828) und Beschreibung von Aphanius sirhani n.sp. (Pisces: Cyprinodontidae). Mitteilungen aus dem hamburgischen zoologischen Museum und Institut 80: 251-277.

Coad, B.W. & Krupp, F. (1983). Redescription of Barilius mesopotamicus Berg 1932, a poorly known cyprinid fish from the Tigris-Euphrates basin. Cybium 7 (1): 47-56.

Krupp, F. (1983). Freshwater fishes of Saudi Arabia and adjacent regions of the Arabian peninsula. Fauna of Saudi Arabia 5: 568-636.

Krupp, F. (1982). Garra tibanica ghorensis subsp. nov. (Pisces: Cyprinidae), an African element in the cyprinid fauna of the Levant.- Hydrobiologia 88: 319-324.

Susanne Dorow
Technische Assistentin
Jennifer Steppler
Technische Assistenz
Uwe Zajonz
Researcher, Affiliated to Research Group 'Quantitative Biogeography'
PhD candidate

Former projects

Project group leader: F2 (Stakeholder dialogues and resource use concepts)
Project leader: F2.4F2.5B1.14
Researcher/collaborator: A4.2A3.3

Research interests and expertise

Basic research
Community ecology of tropical marine ecosystems in climate change, especially of fish communities of biogene reefs and coastal estuaries 
Marine biogeography
Taxonomy, systematics and biogeography of coastal fishes of the Indo-West-Pacific, focusing on the circum-Arabian Seas
Taxonomy and ecology of the deep-water and deep-sea fishes of the Red Sea and Gulf of Aden

Transfer and applied research
Tropical marine ecosystem functions and services under climate change 
Integrated Coastal Zone Management and marine conservation and resource use management at the interface to human development cooperation
Marine and coastal environmental research and consulting at the technical and policy level for institutional and private organizations and stakeholders 
Project development and management


– Transdisciplinary work group „Tropical Marine Ecosystems“ (TMS)  
– Field research station Socotra 
– Recent article on Socotra in National Geographic
– Soqotra Symposium 2012

Short CV

Since 2008 Researcher and project leader with BiK-F
2008-2010 Member of the start-up team and of the Scientific Coordination Office (WKB) of BiK-F
2008  Member of the project formulation task force for the LOEWE centre BiK-F
2003-2008 Free lance environmental consultant and project leader to UNDP, World Bank, governmental authorities and the private sector, e.g.: 
– 2005-2007 Project leader „Coastal Zone Management Pilot Project in the Yemen Gulf of Aden“ (World Bank-GEF)
– 2004-2005 Project leader „Protection of Marine Ecosystems of the Yemen Red Sea Coast (UNDP-GEF)
– 2003 Deputy project leader „Socotra Conservation and Development Project“ (UNDP-GEF-Netherlands)
1997-2001 Project coordination assistant for SFN and ichthyologist of „Marine Habitat, Biodiversity and Fisheries Surveys“ in the framework of the UNDP-GEF Project „Conservation and Sustainable Use of Biodiversity of Socotra Archipelago“
1995-2002 Research assistant at SFN, Section Ichthyology, temporarily acting Head of Section
1995 Diploma in Biology (Dipl.-Biol.) with the subjects Zoology, Aquatic Ecology, Entomology and Soil Science
1988-1995 Studies of Biology at the Technical University Darmstadt

Publications and reports

Dipl.-Biol.  Florian Wicker

Forschungsschwerpunkte:  Systematik, Phylogenie und Zoogeographie der Gattungen Cyprinion Heckel, 1843 und Garra Hamilton, 1822 aus der Familie der Karpfenfische

Dr. Tilman J. Alpermann

Forschungsschwerpunkte: Phylogeographie, Populationsgenetik und Taxonomie der Fische der Arabischen Meere und des Indopazifik

Hata, H., Lavoué, S., Bogorodsky, S.V., Alpermann, T.J. & Motomura, H. (2023) A new Stolephorus (Teleostei: Clupeiformes: Engraulidae: Engraulinae) from the Red Sea. Ichthyology & Herpetology, 111 (2): 191–203. https://doi.org/10.1643/i2022064

Bogorodsky, S.V., Kovačić, M., Mal, A.O. & Alpermann, T.J. (2023) A new species of Fusigobius (Teleostei: Gobiidae) from the Red Sea and Gulf of Aden. Zootaxa, 5256 (2): 101–124. https://doi.org/10.11646/zootaxa.5256.2.1

Robitzch, V., Saenz-Agudelo, P., Alpermann, T.J., Frédérich, B. & Berumen, M.L. (2023) Contrasting genetic diversity and structure between endemic and widespread damselfishes are related to differing adaptive strategies. Journal of Biogeography, 50 (2): 380–392. https://doi.org/10.1111/jbi.14540

Thieme, P., Bogorodsky, S.V., Alpermann, T.J., Whitfield, A.K., Freitas, R. & Durand J.-D. (2022) Contributions to the taxonomy of the mugilid genus Chelon Artedi (Teleostei: Mugilidae), with a major review of the status of C. persicus Senou, Randall & Okiyama, 1995. Zootaxa, 5188 (1): 001–042. https://doi.org/10.11646/zootaxa.5188.1.1

Russell, B.C, Bogorodsky, S.V., Mal,, A.O., Bineesh, K.K. & Alpermann, T.J. (2022) The taxonomic identity of the monocle bream Scolopsis vosmeri species complex (Perciformes: Nemipteridae), with comments on molecular phylogenetic relationships within the genus Scolopsis. Zootaxa, 5105 (4): 501–538. https://doi.org/10.11646/zootaxa.5105.4.3

Fraser, T.H., Bogorodsky, S.V., Mal, A.O. & Alpermann, T.J. (2021) Review of the cardinalfishes of the genus Cercamia (Percomorpha: Apogonidae) of the Red Sea and Indian Ocean with descriptions of three new species. Zootaxa, 5039 (3): 363–394. https://doi.org/10.11646/zootaxa.5039.3.3

Gon, O., Bogorodsky, S.V., Mal, A.O. & Alpermann, T.J. (2020) A new species of the cardinalfish genus Apogon (Teleostei, Apogonidae) from the southern Red Sea and Indian Ocean with comments on phylogenetic relationships within the Apogonini. Zootaxa, 4896 (4): 485–504. https://doi.org/10.11646/zootaxa.4896.4.2

Matsuura, K., Bogorodsky, S.V., Mal, A.O. & Alpermann T.J. (2020) Canthigaster aziz, a new deep dwelling toby fish (Tetraodontiformes: Tetraodontidae) from the Red Sea. Zootaxa, 4834 (1): 081–095. https://doi.org/10.11646/zootaxa.4834.1.5

McCosker, J. E., Bogorodsky, S.V., Mal, O. A. & Alpermann, T.J. (2020) Description of a new snake eel Ophichthus olivaceus (Teleostei: Anguilliformes, Ophichthidae) from the Red Sea. Zootaxa, 4750 (1): 31-48. https://doi.org/10.11646/zootaxa.4750.1.2

Smith, D. G., Bogorodsky, S. V., Mal, A. O. & Alpermann, T. J. (2019) Review of the moray eels (Anguilliformes: Muraenidae) of the Red Sea, with description of a new species. Zootaxa, 4704: 1–87. https://doi.org/10.11646/zootaxa.4704.1.1

Kovačić, M., Bogorodsky, S.V., Mal, A.O. & Alpermann, T.J. (2018) Redescription of the genus Koumansetta (Teleostei: Gobiidae), with description of a new species from the Red Sea. Zootaxa, 4459 (3): 453–481. https://doi.org/10.11646/zootaxa.4459.3.3

Kovačić, M, Bogorodsky, S.V, Mal, A.O. & Alpermann, T.J. (2018) First record of Valenciennea parva (Teleostei: Perciformes, Gobiidae) for the Red Sea, with a key to Red Sea species of the genus. Marine Biodiversity, 48: 1229–1236. https://doi.org/10.1007/s12526-017-0719-y

Bogorodsky, S.V., Smith-Vaniz, W., Mal, A.O. & Alpermann, T.J. (2017) Review of Carangoides (Perciformes: Carangidae) from the Red Sea, with a new record of Imposter Trevally Carangoides talamparoides Bleeker, 1852. Marine Biodiversity, 47: 1251–1271. https://doi.org/10.1007/s12526-017-0776-2

Marceniuk, A.P., Bogorodsky, S.V., Mal, A.O. & Alpermann, T.J. (2017) Redescription of the blacktip sea catfish Plicofollis dussumieri (Valenciennes) (Siluriformes: Ariidae), with a new record from the Red Sea and notes on the diversity and distribution of Plicofollis spp. Marine Biodiversity, 47: 1239–1250. https://doi.org/10.1007/s12526-017-0760-x

Bogorodsky, S.V., Iwatsuki, Y., Amir, S., Mal, A.O. & Alpermann, T.J. (2017) Morphological and molecular divergence between Crenidens crenidens (Forsskål, 1775) and C. indicus Day, 1873 (Perciformes: Sparidae) and notes on a Red Sea endemic lineage of C. crenidens. Marine Biodiversity, 47: 1273–1285. https://doi.org/10.1007/s12526-017-0764-6

Last, P.R., Bogorodsky, S.V. & Alpermann, T.J. (2016) Maculabatis ambigua sp. nov., a new whipray (Myliobatiformes: Dasyatidae) from the Western Indian Ocean. Zootaxa, 4154(1): 66–78. https://doi.org/10.11646/zootaxa.4154.1.4

Sala-Pérez, M., Alpermann, T.J., Krock, B. & Tillmann, U. (2016) Growth and bioactive secondary metabolites of arctic Protoceratium reticulatum (Dinophyceae). Harmful Algae, 55: 85–96. https://doi.org/10.1016/j.hal.2016.02.004

Tillmann, U., Krock, B., Alpermann, T.J. & Cembella, A. (2016) Bioactive compounds of marine dinoflagellate isolates from western Greenland and their phylogenetic association within the genus Alexandrium. Harmful Algae, 51: 67–80. https://doi.org/10.1016/j.hal.2015.11.004

Bogorodsky, S. V., Alpermann, T.J. & Mal, A.O. (2016) Redescription of Cheilinus quinquecinctus Rüppell, 1835 (Pisces: Perciformes, Labridae), a valid endemic Red Sea wrasse. Zootaxa, 4158 (4): 451–472.

Last, P.R., Bogorodsky, S.V. & Alpermann, T.J. (2016) Maculabatis ambigua sp. nov., a new whipray (Myliobatiformes: Dasyatidae) from the Western Indian Ocean. Zootaxa, 4154 (1): 66–78

John, U., Tillmann, U., Hülskötter, J., Alpermann, T.J., Wohlrab, S. & van der Waal, D.B. (2015) Intraspecific facilitation by allelochemical mediated grazing protection within a toxigenic dinoflagellate population. Proceedings of the Royal Society B, 282 (1798): 20141268

Akselman, R., Krock, B., Alpermann, T.J., Tillmann, U., Borel, C.M., Almandoz, G.O. & Ferrario, M.E. (2015) Protoceratium reticulatum (Dinophyceae) in the austral Southwestern Atlantic and the first report on YTX-production in shelf waters of Argentina. Harmful Algae, 45: 40–52

Alpermann, T.J., Plieske, J., Mal, A.O., Gon, O. & Berumen M.L. (2014) Isolation and characterization of polymorphic microsatellite loci for Siphamia tubifer Weber (Perciformes: Apogonidae) Conservation Genetics Resources, 6: 1031–1034

Bogorodsky, S. V., Alpermann, T.J., Mal, A.O. & Gabr , M. (2014) Survey of demersal fishes from southern Saudi Arabia, with five new records for the Red Sea. Zootaxa, 3852 (4): 401–437

Randall, J.E, Victor, B.C., Alpermann, T.J., Bogorodsky, S.V., Mal, A.O., Satapoomin, U. & Bineesh, K.K. (2014) Rebuttal to Koeda et al. (2014) on the Red Sea fishes of the perciform genus Pempheris. Zootaxa, 3887 (3): 377–392

Bogorodsky, S.V., Last, P.R., Alpermann, T.J. & Mal, A.O. (2014) Records of Himantura granulata (Dasyatidae) and Rhinoptera jayakari (Rhinopteridae) from the Red Sea. Zoology in the Middle East, 60 (2): 144–153.

Alpermann, T.J., Mee, J.K.L. & Berumen, M.L. (2013) First record of the black-fin coral goby, Paragobiodon lacunicolus (Kendall & Goldsborough), from the Red Sea. Journal of Applied Ichthyology, 29 (6): 1361–1363.

Krock, B., Tillmann, T., Alpermann, T.J., Voß, D., Zielinski, O. & Cembella. A.D. (2013) Phycotoxin composition and distribution in plankton fractions from the German Bight and western Danish coast. Journal of Plankton Research, 35 (5): 1093–1108.

Toebe, K., Alpermann, T., Tillmann, U., Krock, B., Cembella, A. & John, U. (2013) Molecular discrimination of toxigenic and non-toxic Alexandrium species in natural phytoplankton assemblages from the Scottish coast of the North Sea. European Journal of Phycology, 48 (1): 12–26.

Randall, J.E., Bogorodsky, S.V., Alpermann, T.J., Satapoomin, U., Mooi, R.D. & Mal, A.O. (2013) Pempheris flavicycla, a new pempherid fish from the Indian Ocean, previously identified as P. vanicolensis Cuvier. Journal of the Ocean Science Foundation, 9: 1–23.

Tahvanainen, P., Alpermann, T.J., Figueroa, R.I., John, U., Hakanen, P., Nagai, S., Blomster, J. & Kremp, A. (2012) Patterns of post-glacial genetic differentiation in marginal populations of a marine microalga. PLoS One, 7 (12): e53602.

Molecular Ecology Resources Primer Development Consortium, C., Abreu, A.G., Albaina, A., Alpermann, T.J., Apkenas, V.E., Bankhead-Dronnet, S., Bergek, S., Berumen, M.L., Cho, C.-H., Clobert, J. et al. (2012). Permanent Genetic Resources added to Molecular Ecology Resources Database 1 October 2011 – 30 November 2011. Molecular Ecology Resources, 12 (2): 374–376.

Anderson, D.M., Alpermann, T.J., Cembella, A.D., Collos, Y., Masseret, E. & Montresor, M. (2012) The globally distributed genus Alexandrium: Multifaceted roles in marine ecosystems and impacts on human health. Harmful Algae, 14: 10–35

Coyer, J.A., Hoarau, G., Pearson, G., Mota, C., Jüterbock, A., Alpermann, T., John, U. & Olson, J.L. (2011) Genomic scans detect signatures of selection along a salinity gradient in populations of the intertidal seaweed Fucus serratus on a 12 km scale. Marine Genomics, 4 (1): 41–49.

Alpermann, T.J., Tillmann, U., Beszteri, B., Cembella, A.D. & John, U. (2010) Phenotypic variation and genotypic diversity in a planktonic population of the toxigenic marine dinoflagellate Alexandrium tamarense. Journal of Phycology, 46: 18–32.

Alpermann, T.J., Beszteri, B., John, U., Tillmann, U. & Cembella, A.D. (2009) Implications of life history transitions on the population genetic structure of the toxigenic marine dinoflagellate Alexandrium tamarense. Molecular Ecology, 18 (10): 2122–2133.

Legrand, C, Alpermann, T., Béchemin, C., Fragoso, B., Huelskötter, J., John, U., Müller, A., Olofsson, M., Rudström, M., Tillmann, U., Weissbach, A., Wohlrab, S. & Cembella, A. (2010) Role of genotypic and phenotypic (allelopathy) diversity of harmful algal blooms (HABs): Case of the dinoflagellate Alexandrium tamarense. In: Proceedings of the HYDRALAB III Joint User Meeting, Hannover, February 2010, J. Grüne & M. Klein Breteler (Eds.), Forschungszentrum Küste, Universität Hannover und Technische Universität Braunschweig (ISBN 78‐3‐00‐030141‐4)

Tillmann, U., Alpermann, T.J., da Purificação, R.C., Krock, B. & Cembella, A.D. (2009) Intra-population clonal variability in allelochemical potency of the toxigenic dinoflagellate Alexandrium tamarense. Harmful Algae, 8 (5): 759–769.

Tillmann, U., Alpermann, T.J., John, U. & Cembella, A.D. (2008) Allelochemical interactions and short- term effects of the dinoflagellate Alexandrium on selected photoautotrophic and heterotrophic protists. Harmful Algae, 7 (1): 52–64.

Krock, B., Alpermann, T.J., Tillmann, U., Pitcher, G.C. & Cembella, A.D. (2008) Yessotoxin profiles of the marine dinoflagellates Protoceratium reticulatum and Gonyaulax spinifera. Proceedings of the 12th International Conference on Harmful Algae, 4-8 September 2006, Copenhagen, Denmark, 303–305.

Alpermann, T.J., Beszteri, B., Tillmann, U., Cembella, A.D. & John, U. (2008) Species discrimination in the genus Alexandrium by amplified fragment length polymorphism. Proceedings of the 12th International Conference on Harmful Algae, 4-8 September 2006, Copenhagen, Denmark, 51–54.

Alpermann, T.J., John, U., Medlin, L.K., Edwards, K.J., Hayes, P.K. & Evans, K.M. (2006) Six new microsatellite markers for the toxic marine dinoflagellate Alexandrium tamarense. Molecular Ecology Notes, 6: 1057–1059.

Koehler, A., Alpermann, T.J., Lauritzen, B. & van Noorden, C.J.F. (2004) Clonal xenobiotic resistance during pollution-induced toxic injury and hepatocellular carcinogenesis in liver of female flounder (Platichthys flesus (L.)) Acta Histochemica, 106 (2): 155–170.

Sergey Bogorodsky

Station of Ichthyologists
Omsk, Rußland