Human Adaptation, Bio-cultural Diversity and Ecology


Peer-reviewed Articles / ISI


Altolaguirre Y, Bruch AA, Gibert L (2020) A long Early Pleistocene pollen record from Baza Basin (SE Spain): Major contributions to the palaeoclimate and palaeovegetation of Southern Europe. Quaternary Science Reviews, 231.

Bataille G, Falcucci A, Tafelmaier Y, Conard NJ (2020) Technological differences between Kostenki 17/II (Spitsynskaya industry, Central Russia) and the Protoaurignacian: Reply to Dinnis et al. (2019). Journal of Human Evolution, 146, 102685.

Baumann C, Bocherens H, Drucker DG, Conard NJ (2020) Fox dietary ecology as a tracer of human impact on Pleistocene ecosystems. PLOS One, 15.

Baumann C, Starkovich B, Drucker DG, Münzel SC, Conard NJ, Bocherens H (2020) Dietary niche partitioning among Magdalenian canids in southwestern Germany and Switzerland. Quaternary Science Reviews, 227.

Baumann C, Wong G, Starkovich B, Münzel SC, Conard NJ (2020) The role of foxes in the Palaeolithic economies of the Swabian Jura (Germany). Archaeological and Anthropological Sciences, 12, 208.

Bosman AM, Reyes-Centeno H, Harvati K (2020) A virtual assessment of the suprainiac depressions on the Eyasi I (Tanzania) and Aduma ADU-VP-1/3 (Ethiopia) Pleistocene hominin crania. Journal of Human Evolution, 145.

Bridault A, Binois-Roman A, Drucker DG, Pion G (2020) Investigating the exploitation pattern of a newly established species, the case of the Late-Glacial roe deer (Capreolus capreolus) at La Fru (Savoie, France). Archaeological and Anthropological Sciences, 12, 229.

Conard NJ, Serangeli J, Bigga G, Rots V (2020) A 300,000-year-old throwing stick from Schöningen, northern Germany, documents the evolution of human hunting. Nature Ecology and Evolution, 4, 690-693.

Denys C, Otero O, Kullmer O, Sandrock O, Bromage TG, Schrenk F, Dauphin Y (2020) Biominerals Fossilisation: Fish Bone Diagenesis in Plio–Pleistocene African Hominid Sites of Malawi. Minerals, 10, 1049.

Drucker DG, Ducrocq TF, Baumann C, Valentin F (2020) Environment and human subsistence in Northern France at the Late-Glacial to early Holocene transition. Archaeological and Anthropological Sciences, 12, 1-16.

Fiorenza L, Menter CG, Fung S, Lee J, Kaidonis J, Moggi-Cecchi J, Townsend G, Kullmer O (2020) The functional role of the Carabelli trait in early and late hominins. Journal of Human Evolution, 145, 102816.

Furtwängler A, Rohrlach AB, Lamnidis TC, Papac L, Neumann GU, Siebke I, Reiter E, Steuri N, Hald JR, Denaire A, Schnitzler B, Wahl J, Ramstein M, Schuenemann VJ, Stockhammer PW, Hafner A, Lösch S, Haak W, Schiffels S, Krause J (2020) Ancient genomes reveal social and genetic structure of Late Neolithic Switzerland. Nature Communications, 11, 1915.

Gokhman D, Nissim-Rafinia M, Agranat-Tamir L, Housman G, García-Pérez R, Lizano E, Cheronet O, Mallick S, Nieves-Colón M, Li H, Alpaslan-Roodenberg S, Novak M, Gu H, Osinski J, Ferrando-Bernal M, Gelabert P, Lipende I, Mjungu D, Kondova I, Bontrop R, Kullmer O, Weber GW, Shahar T, DvirGinzberg M, Faerman M, Quillen E, Meissner A, Lahav Y, Kandel L, Liebergall M, Prada ME, Encinas JMV, Gronostajski R, Stone A, Yakir B, Lalueza-Fox C, Pinhasi R, Reich D, Marques-Bonet T, Meshorer E, Carmel L (2020) Differential DNA methylation of vocal and facial anatomy genes in modern humans. Nature Communications, 11, 1189.

Gunz P, Kozakowski S, Neubauer S, Le Cabec A, Kullmer O, Benazzi S, Hublin JJ, Begun DR (2020) Skull reconstruction of the late Miocene ape Rudapithecus hungaricus from Rudabánya, Hungary. Journal of Human Evolution, 138, 102687.

Habinger S, De Cupere B, Dövener F, Pucher E, Bocherens H (2020) Mobility and origin of camels in the Roman Empire through serial stable carbon and oxygen isotopes variations in tooth enamel. Quaternary International, 557, 80-91.

Haeusler M, Webb NM, Krenn VA, Fornai C (2020) Locomotor and taxonomic diversity of Sterkfontein hominins not supported by current trabecular evidence of the femoral head. Proceedings of the National Academy of Sciences, 117, 28568-28569.

Heidgen S, Marinova E, Krauß R, Nelle O, Ebner M, Märkle T, Miranda T, Bofinger A, Klingler S, Junginger A (2020) Palaeoenvironment and potential resources for early Holocene subsistence in the Ammer River Valley (Germany) based on palaeoecological and bioarchaeological evidence. Quaternary International, 560-561, 259-272.

Hirniak JN, Smith EI, Johnsen R, Ren M, Hodgkins J, Orr C, Negrino F, Riel‐Salvatore J, Fitch S, Miller CE, Zerboni A (2020) Discovery of cryptotephra at Middle–Upper Paleolithic sites Arma Veirana and Riparo Bombrini, Italy: a new link for broader geographic correlations. Journal of Quaternary Science, 35, 199-212.

Iovita R, Varis A, Namen A, Cuthbertson P, Taimagambetov Z, Miller CE (2020) In search of a Paleolithic Silk Road in Kazakhstan. Quaternary International, 559, 119-132.

Jafarzadeh R, Konidaris GE (2020) Choerolophodon (Proboscidea, Mammalia) from a new upper Miocene locality: Kivi-1 (Iran). Palaeobiodiversity and Palaeoenvironments 100, 269–280.

Jarl J, Riehl S, Deckers K, Becker JA (2020) Plant cultivation under climatic fluctuations during the sixth and fifth millennia BC at Tell Tawila (northern Syria). Archaeological and Anthropological Sciences, 12, 249-266.

Kielhofer J, Miller CE, Reuther J, Holmes C, Potter B, Lanoë F, Esdale J, Crass B (2020) The micromorphology of loess‐paleosol sequences in central Alaska: A new perspective on soil formation and landscape evolution since the Late Glacial period (c. 16,000 cal yr BP to present). Geoarchaeology, 35, 701-728.

Kitagawa K, Conard NJ (2020) Split-based Points from the Swabian Jura highlight Aurignacian regional signatures. PLOS One, 15, e0239865.

Konidaris GE, Kostopoulos DS, Koufos GD (2020) Mammuthus meridionalis (Nesti, 1825) from Apollonia-1 (Mygdonia Basin, Northern Greece) and its importance within the Early Pleistocene mammoth evolution in Europe. Geodiversitas, 42, 69-91.

Konidaris GE, Tsoukala E (2020) Proboscideans from the upper Miocene localities of Thermopigi, Neokaisareia and Platania (Northern Greece). Annales de Paleontologie, 106.

Krajcarz M, Krajcarz MT, Baca M, Baumann C, Van Neer W, Popović D, Sudoł-Procyk M, Wach B, Wilczyński J, Wojenka M, Bocherens H (2020) Ancestors of domestic cats in Neolithic Central Europe – isotopic evidence of a synanthropic diet. PNAS, 117, 1771-1779.

Lipson M, Ribot I, Mallick S, Rohland N, Olalde I, Adamski N, Broomandkhoshbacht N, Lawson AM, Lopez S, Oppenheimer J, Stewardson K, Asombang R, Bocherens H, Bradman N, Cornelissen E, Crevecoeur I, Lavachery P, de Maret P, Mindzie CM, et al. EA, Reich D (2020) Ancient West African foragers in the context of deep human genetic history. Nature, 577, 665-670.

Majander K, Pfrengle S, Kocher A, Neukamm J, du Plessis L, Pla-Díaz M, Arora N, Akgül GL, Salo K, Schats R, Inskip S, Oinonen M, Valk H, Malve M, Kriiska A, Onkamo PI, González-Candelas F, Kühnert D, Krause J, Schuenemann VJ (2020) Ancient Bacterial Genomes Reveal a High Diversity of Treponema pallidum Strains in Early Modern Europe. Current Biology, 30, 3788-3803.e10.

Meadows J, Lozovskaya O, Bondetti M, Drucker DG, Moiseyev V (2020) Human palaeodiet at Zamostje 2, Central Russia: results of radiocarbon and stable isotope analyses. Quaternary International.

Moncel MHLN, Biddittu I, Manzi G, Saracino B, Pereira A, Nomade SB, Hertler C, Voinchet P, Bahain JJ (2020) Emergence of regional cultural traditions during the Lower Palaeolithic: the case of Frosinone-Ceprano basin (Central Italy) at the MIS 11–10 transition. Archaeological and Anthropological Sciences, 12.

Naito YI, Meleg IN, Robu M, Drucker DG, Wißing C, Hofreiter M, Barlow A, Bocherens H (2020) Heavy reliance on plants for Romanian cave bears evidenced by amino acid nitrogen isotope analysis. Scientific Reports, 10.

Neukamm J, Pfrengle S, Molak M, Seitz A, Francken M, Eppenberger P, Avanzi C, Reiter E, Urban C, Welte B, Stockhammer PW, Teßmann B, Herbig A, Harvati K, Nieselt K, Krause J, Schuenemann VJ (2020) 2,000-year-old pathogen genomes reconstructed from metagenomic analysis of Egyptian mummified individuals. BMC BIOLOGY, 108.

Oinonen M, Alenius T, Arppe L, Bocherens H, Etu-Sihvola H, Helama S, Huhtamaa H, Lahtinen M, Mannermaa K, Onkamo P, Vanhanen S, Wessman A (2020) Buried in water, burdened by nature-Resilience carried the Iron Age people through Fimbulvinter. PLOS One, 15.

Richard M, Falguères C, Pons-Branchu E, Richter D, Beutelspacher T, Conard NJ, Kind CJ (2020) The Middle to Upper Palaeolithic transition in Hohlenstein Stadel cave (Swabian Jura, Germany): A comparison between ESR, U-series and radiocarbon dating. Quaternary International, 556, 49-57.

Rivals F, Drucker DG, Weber MJ, Audouze F, Enloe JG (2020) Dietary traits and habitats of the reindeer (Rangifer tarandus) during the Late Glacial. Archaeological and Anthropological Sciences, 12, 1-13.

Rufi I, Drucker DG, Bocherens H, Lloveras L, Madurell-Malapeira J, Maroto J, Soler J, Soler N (2020) Revision of the occurrence of muskox (Ovibos moschatus Zimmermann 1780) from the Gravettian of Arbreda Cave (Serinyà, northeast of Iberian Peninsula): new insights for the study of Iberian cold-adapted faunas. Boreas, 49, 858-872.

Siebke I, Furtwängler A, Steuri N, Hafner A, Ramstein M, Krause J, Lösch S (2020) Crops vs. animals: regional differences in subsistence strategies of Swiss Neolithic farmers revealed by stable isotopes. Archaeological and Anthropological Sciences, 12, 235.

Silva K, Bräuer J, de Sousa L, Lima M, O’Hara R, Belger J, Epperlein T, Tennie C (2020) An attempt to test whether dogs (Canis familiaris) show increased preference towards humans who match their behaviour. Journal of Ethology, 38, 223-232.

Simons EA, Frost SR, Harvati K, McNulty K, Singleton M (2020) Comparing rates of lineage diversification with rates of size and shape evolution in catarrhine crania. Evolutionary Biology, 47, 152-163.

Uhl A, Karakostis FA, Wahl J, Harvati K (2020) A cross-population study of sexual dimorphism in the bony labyrinth. Archaeol Anthropol Science, 12, 132.

Will M, Conard NJ (2020) Regional patterns of diachronic technological change in the Howiesons Poort of southern Africa. PLOS One, 15.

Wong G, Drucker DG, Starkovich B, Conard NJ (2020) Latest Pleistocene paleoenvironmental reconstructions from the Swabian Jura, southwestern Germany: Evidence from stable isotope analysis and micromammal remains. Palaeogeography, Palaeoclimatology, Palaeoecology, 540.

Wong G, Starkovich B, Drucker DG, Conard NJ (2020) New perspectives on human subsistence during the Magdalenian in the Swabian Jura, Germany. Archaeologcial and Anthropological Sciences, 12, 217.

Zanolli C, Schllinger B, Kullmer O, Schrenk F, Kelley J, Rössner GE, Macchiarelli R (2020) When X-Rays Do Not Work. Characterizing the Internal Structure of Fossil Hominid Dentognathic Remains Using High-Resolution Neutron Microtomographic Imaging. Frontiers in Ecology and Evolution, 8, 42.


Abell JT, Quade J, Duru G, Mentzer SM, Stiner MC, Uzdurum M, Özbaşaran M (2019) Urine salts elucidate Early Neolithic animal management at Asikli Hoyuk, Turkey. Science Advances, 5, eaaw0038.

Altolaguirre Y, Postigo-Mijarra JM, Barrón E, Carrion JS, Leroy S, Bruch AA (2019) An environmental scenario for the earliest hominins in the Iberian Peninsula: Early Pleistocene palaeovegetation and palaeoclimate. Review of Palaeobotany and Palynology, 260, 51-64.

Anastopoulou I, Karakostis FA, Moraitis K (2019) A reliable regression-based approach for reassociating human skeletal elements of the lower limbs from commingled assemblages. Journal of Forensic Sciences, 64, 502-506.

Arppe L, Karhu JA, Vartanyan S, Drucker DG, Etu-Sihvola H, Bocherens H (2019) Thriving or surviving? The isotopic record of the Wrangel Island woolly mammoth population. Quaternary Science Reviews, 222.

Bons PD, Bauer CC, Bocherens H, de Riese T, Drucker DG, Francken M, Menéndez LP, Uhl A, van Milligen BP, Wißing C (2019) Out of Africa by spontaneous migration waves. PLOS One, 14.

Bosman AM, Harvati K (2019) A virtual assessment of the propose suprainiac fossa on the Early Modern European Calvaria from Cioclovina, Romania. American Journal of Physical Anthropology, 169, 567-574.

Bromage TG, Juwayeyi YM, Katris JA, Gomez S, Ovsiy O, Goldstein J, Janal MN, Hu B, Schrenk F (2019) Corrigendum to “The scaling of human osteocyte lacuna density with body size and metabolism” [C. R. Palevol. 15 (2016) 33–40]. Comptes Rendus Palevol, 18, 492.

Bulut O, Freudenstein N, Hekimoglu B, Gurcan S (2019) Dilemma of Gonial Angle in Sex Determination Sexually Dimorphic or Not? The American Journal of Forensic Medicine and Pathology, 40, 361-365.

Bulut O, Liu C, Gurcan S, Hekimoglu B (2019) Prediction of nasal morphology in facial reconstruction: Validation and recalibration of the Rynn method. Legal Medicine, 40, 26-31.

Díaz-Zorita Bonilla M, Aranda Jimènez G, Bocherens H, Escudero Carillo J, Sánchez Romero M, Lozano Medina A, Alarcón García E, Milesi Garcia L (2019) Multi-isotopic diet determination of Southeastern Iberian Megalithic populations: The cemeteries of El Barranquete and Panoría. Archaeological and Anthropological Sciences, 11, 3681-3698.

Fiorenza L, Benazzi S, Kullmer O, Zampirolo G, Mazurier A, Zanolli C, Macchiarelli R (2019) Dental macrowear and cortical bone distribution of the Neanderthal mandible from Regourdou (Dordogne, Southwestern France). Journal of Human Evolution.

Fuentes R, Ono R, Nakajima N, Nishizawa H, Siswanto J, azis N, Sofian HO, Miranda T, Pawlik AF (2019) Technological and behavioural complexity in expedient industries: The importance of use-wear analysis for understanding flake assemblages. Journal of Archeological Science, 112.

Giusti D, Konidaris GE, Tourloukis V, Marini M, Maron M, Zerboni A, Thompson N, Koufos GD, Kostopoulos DS, Harvati K (2019) Recursive anisotropy: a spatial taphonomic study of the Early Pleistocene vertebrate assemblage of Tsiotra Vryssi, Mygdonia Basin, Greece. Boreas, 48, 713-730.

Gretzinger J, Molak M, Reiter E, Pfrengle S, et al. EA, Conard NJ, et al. EA, Drucker DG, et al. EA, Bocherens H, Schuenemann VJ (2019) Large-scale mitogenomic analysis of the phylogeography of the Late Pleistocene cave bear. Scientific Reports, 9, 10700.

Haaland M, Czechowski M, Carpentier F, Lejay M, Vandermeulen B (2019) Documenting archaeological thin sections in high-resolution: A comparison of methods and discussion of applications. Geoarchaeology, 34, 100-114.

Habermann JRM, Alberti M, Aldeias V, Alemseged Z, Archer W, Bamford M, Biro D, Braun DR, Capelli C, Cunha EN, da Silva MF, Lüdecke T, Madiquida HR, Martinez FI, Mathe J, Negash E, Paulo LM, Pinto M, Stalmans M, Regala FTT, Wynn JG, Bobe R, Carvalho S (2019) Gorongosa by the sea: First Miocene fossil sites from the Urema Rift, central Mozambique, and their coastal paleoenvironmental and paleoecological contexts. Palaeogeography, Palaeoclimatology, Palaeoecology, 514, 723-738.

Harvati K, Röding C, Bosman AM, Karakostis FA, Grün R, Stringer C, Karkanas P, Thompson N, Koutoulidis V, Moulopoulos LA, Gorgoulis VG, Kouloukoussa M (2019) Apidima Cave fossils provide earliest evidence of Homo sapiens in Eurasia. Nature, 571, 500-504.

Hofman-Kaminska E, Bocherens HDrucker DG, Fyfe RM, Gumiński W, Makowiecki D, Pacher M, Piličiauskienė G, Samojlik T, Woodbridge J, Kowalczyk R (2019) Adapt or die – response of large herbivores to environmental changes in Europe during the Holocene. Global Change Biology, 25, 2915-2930.

Ioannidou M, Koufos GD, de Bonis L, Harvati K (2019) A new three-dimensional geometric morphometrics analysis of the Ouranopithecus macedoniensis cranium (Late Miocene, Central Macedonia, Greece). American Journal of Physical Anthropology, 170, 295-307.

Joannes-Boyau R, Adams JW, Austin C, Arora M, Moffart I, Herries AI, Tonge MP, Benazzi S, Evans AR, Kullmer O, Wroe S, Dosseto A, Fiorenza L (2019) Elemental signatures of Australopithecus africanus teeth reveal seasonal dietary stress. Nature, 572, 112–115.

Karakostis FA, Jeffery N, Harvati K (2019) Experimental proof that multivariate patterns among muscle attachments (entheses) can reflect repetitive muscle use. Scientific Reports, 9.

Karakostis FA, Vlachodimitropoulos D, Piagkou M, Scherf HHarvati K, Moraitis K (2019) Is bone elevation in hand muscle attachments associated with biomechanical stress? A histological approach to an anthropological question. Anatomical Record – Advances in Integrative Anatomy and Evolutionary Biology, 302, 1093-1103.

Karakostis FA, Wallace I, Konow N, Harvati K (2019) Experimental evidence that physical activity affects the multivariate associations among muscle attachments. Journal of Experimental Biology.

Kochey JK, Daniels SR, Plagge C, Mehrabi S, Hartmann L, Schrenk F, Plath M, Klaus S (2019) Genetic differentiation of the Malawi blue crab reflects Pleistocene desiccation of Lake Malawi (Brachyura, Potamonautidae: Potamonautes lirrangensis (Rathbun, 1904)). Hydrobiologia, 843, 1-11.

Konidaris GE, Koufos G (2019) Late Miocene proboscideans from Samos Island (Greece) revisited: new specimens from old collections. Paläontologische Zeitschrift, 93, 115-134.

Krajcarz MT, Krajcarz M, Drucker DGBocherens H (2019) Prey-to-fox isotopic enrichment of 34S in bone collagen: Implications for palaeoecological studies. Rapid Communications in Mass Spectrometry, 33, 1311-1317.

Kranioti E, Grigorescu D, Harvati K (2019) State of the art forensic techniques reveal evidence for interpersonal violence ca. 30.000 years ago. PLOS One, 14.

Krijgsman W, Tesakov A, Yanina T, Lazarev S, Danukalova G, van Baak CGC, Agusti J, Alcicek MC, Aliyeva E, Bista D, Bruch AA, Büyükmeric Y, Bukhsianidze M, Flecker R, Frolov P, Hoyle T, Jorissen EL, Kirscher U, Koriche SA, Kroonenberg SB, Lordkipanidze D, Oms O, Rausch L, Singarayer J, Stoica M, van de Velde S, Titov VV, Wesselingh FP (2019) Quaternary time scales for the Pontocaspian domain: Interbasinal connectivity and faunal evolution. Earth-Science Reviews, 188, 1-40.

Lacruz R, Stringer CB, Kimbel WH, Wood B, Harvati K, O’Higgins P, Bromage TG, Arsuaga JL (2019) The evolutionary history of the human face. Nature Ecology & Evolution, 3, 726-736.

Larbey C, Mentzer SM, Ligouis B, Wurz S, Jones M (2019) Cooked starchy food in a 120,000-year-old hearth from Klasies River Cave, South Africa. Journal of Human Evolution, 131, 210-227.

Martinez FI, Capelli C, Ferreira da Silva MJ, Aldeias V, Alemseged Z, Archer W, Bamford M, Biro D, Bobe R, Braun DR, Habermann JRM, Lüdecke T, Madiquida HR, Mathe J, Negash E, Paulo LM, Pinto M, Stalmans M, Tátá F, Carvalho S (2019) A missing piece of the Papio puzzle: Gorongosa baboon phenostructure and intrageneric relationships. Journal of Human Evolution, 130, 1-20.

McGrath K, Reid D, Guatelli-Steinberg D, Arbenz-Smith K, El Zaatari S, et al. EA, McFarlin S (2019) Faster growth corresponds with shallower linear hypoplastic defects in great ape canines. Journal of Human Evolution, 137, 102691.

Mori T, Harvati K (2019) Basicranial ontogeny comparison in Pan and Homo and its use for developmental stage definition of KNM-ER42700. American Journal of Physical Anthropology, 170, 579-594.

Popova L, Maul LC, Zagorodniuk IV, Veklych YM, Shydlovskiy PS, Pogodina NV, Bondar K, Strukova TV, Parfitt SA (2019) ‘Good fences make good neighbours’: Concepts and records of range dynamics in ground squirrels and geographical barriers in the Pleistocene of the Circum-Black Sea area. Quaternary International, 509, 103-120.

Rathmann H, Kyle B, Nikita E, Harvati K, Semerari GS (2019) Population history of southern Italy during Greek colonization inferred from dental remains. American Journal of Physical Anthropology.

Rhodes SE, Starkovich BConard NJ (2019) Did climate determine Late Pleistocene settlement dynamics in the Ach Valley, SW Germany? PLOS One, 14, e0215172.

Richard M, Falguères C, Valladas H, Ghaleb B, Pons-Branchu E, Mercier N, Richter D, Conard NJ (2019) New electron spin resonance (ESR) ages from Geißenklösterle Cave: A chronological study of the Middle and early Upper Paleolithic layers. Journal of Human Evolution, 133, 133-145.

Riga A, Mori T, Pickering TR, Moggi-Cecchi J, Menter CG (2019) Ages-at-death distribution of the early Pleistocene hominin fossil assemblage from Drimolen (South Africa). American Journal of Physical Anthropology, 168, 632-636.

Suraprasit K, Jongautchariyakul S, Yamee C, Pothichaiya C, Bocherens H (2019) New fossil and isotope evidence for the Pleistocene zoogeographic transition and hypothesized savanna corridor in Peninsular Thailand. Quaternary Science Reviews, 221.

Terlato G, Bocherens H, Romandini M, Nannini N, Hobson KA, Peresani M (2019) Chronological and isotopic data support a revision for the timing of cave bear extinction in Mediterranean Europe. Historical Biology, 31, 474-484.

Will M, El-Zaatari S, Havarti K, Conard NJ (2019) Human teeth from securely stratified Middle Stone Age contexts at Sibudu, South Africa. Archaeological and Anthropological Sciences, 11, 3491-3501.

Wirkner MHertler C (2019) Feeding ecology of Late Pleistocene Muntiacus muntjak in the Padang Highlands (Sumatra) Écologie alimentaire de Muntiacus muntjak au Pléistocène Terminal dans les Highlands de Padang (Sumatra). Comptes Rendus Palevol, 18, 541-554.

Wißing C, Rougier H, Baumann C, Comeyne A, Crevecoeur I, Drucker DG, Gaudzinski-Windheuser S, Germonpre M, Gómez-Olivencia A, Krause J, Matthies T, Naito YI, Posth C, Semal P, Street M, Bocherens H (2019) Stable isotopes reveal patterns of diet and mobility in last Neandertals and first modern humans in Europe. Scientific Reports, 9, 4433.

Zanolli C, Kullmer O, Kelly J, Bacon AM, Demeter F, Dumoncel J, Fiorenza L, Grine FE, Hublin JJ, Nguyen AT, Nguyen TMH, Lei P, Schillinger B, Schrenk F, Skinner MM, Xueping J, Macchiarelli R (2019) Evidence for increased hominid diversity in the Early to Middle Pleistocene of Indonesia. Nature Ecology & Evolution.


Aluwé K, Starkovich B, Van Vaerenbergh J (2018) A noble diet at the Hof van Leugenhaeghe (Steendorp, Belgium): pig skulls as a fourteenth-fifteenth century delicacy? Archaeological and Anthropological Sciences, 10, 247-257.

Anastopoulou I, Karakostis FA, Borrini M, Moraitis K (2018) A statistical method for reassociating human tali and calcanei from a commingled context. Journal of Forensic Sciences, 63, 381-385.

Athanassiou A, Michailidis D, Vlachos E, Tourloukis V, Thompson N, Harvati K (2018) Pleistocene vertebrates from the Kyparissia lignite mine, Megalopolis Basin, S. Greece: Testudines, Aves, Suiformes. Quaternary International, 497, 178-197.

Barbieri A, Leven-Pfister C, Hodgins G, Kind KJ, Conard NJMiller CE (2018) Bridging Prehistoric caves with buried landscapes in the Swabian Jura (southwestern Germany). Quaternary International, 485, 23-43.

Bauer CC, Benazzi S, Darlas A, Harvati K (2018) Geometric morphometric analysis and internal structure measurements of the Neanderthal lower fourth premolars from Kalamakia, Greece. Quaternary International, 497, 14-21.

Beier J, Anthes N, Wahl JHarvati K (2018) Similar cranial trauma prevalence among Neanderthals and Upper Paleolithic humans. Nature, 563, 686-690.

Bocherens H (2018) The rise of the anthroposphere since 50,000 years: an ecological replacement of megaherbivores by humans in terrestrial ecoystems? Frontiers in Ecology and Evolution, 6, UNSP 3.

Bocherens H, Cotte M, Bonini M, Straccia P, Scian D, Soibelzon L, Prevosti FJ (2018) Reply to “Comment on “Isotopic insight on paleodiet of” by Bocherens et al. (Gondwana Research, 48(1), 7-14)”. Gondwana Research, 58, 243-245.

Bretzke K, Yousif E, Jasim S (2018) Filling in the gap – The Acheulean site Suhailah 1 from the central region of the Emirate of Sharjah, UAE. Quaternary International, 466, 23-32.

Doukas C, van Kolfschoten T, Papayianni K, Panagopoulou E, Harvati K (2018) The small mammal fauna from the Paleolithic site Marathousa 1 (Greece). Quaternary International, 497, 95-107.

Dutkiewicz E, Wolf SConard NJ (2018) Early symbolism in the Ach and the Lone valleys of southwestern Germany. Quaternary International, 491, 30-45.

Dutkiewicz E, Wolf S, Floss H, Conard NJ (2018) Les objets en ivoire du Jura Souabe. The ivory objects of the Swabian Jura. L’Anthropologie, 122, 447-468.

Ebinger-Rist N, Wolf S, Wehrberger K, Kind CJ (2018) L`homme-lion d’ Hohlenstein-Stadel. The lion-man of Hohlenstein-Stadel. L’Anthropologie, 122, 415-436.

El Zaatari S (2018) The central Levantine corridor: The Paleolithic of Lebanon. Quaternary International, 466, 33-47.

El Zaatari SHarvati K, Bae C (2018) Filling the Gaps in the Human Evolutionary Story. Quaternary International, 466 A, 1-106.

Exposito-Alonso M, Becker C, Schuenemann VJ, Reiter E, Setzer C, Slovak R, Brachi B, Hagmann J, Grimm DG, Chen J, Busch W, Bergelson J, et al. EA, Weigel D (2018) The rate and potential relevance of new mutations in a colonizing plant lineage . PLOS Genetics, 14, e10071.

Ferreira G, Iori F, Hermanson G, Langer MC (2018) New turtle remains from the Late Cretaceous of Monte Alto-SP, Brazil, including cranial osteology, neuroanatomy and phylogenetic position of a new taxon. Paläontologische Zeitschrift, 92, 481-498.

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Giusti DTourloukis VKonidaris GE, Thompson N, Karkanas P, Panagopoulou E, Harvati K (2018) Beyond maps: patterns of formation processes at the Middle Pleistocene open-air site of Marathousa 1, Megalopolis Basin, Greece. Quaternary International, 497, 137-153.

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Harvati KKonidaris GTourloukis V (2018) Paleoanthropology at the Gates of Europe: Recent research in Greece in the frame of the PaGE ERC Starting Grant project. Quaternary International, 497, 1-3.

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Kandel AW, Bretzke KConard NJ (2018) Eipaleolithic shell beads from Damascus Province, Syria. Quaternary International, 464, 126-140.

Karakostis FA, Hotz G, Scherf HWahl JHarvati K (2018) A repeatable geometric morphometric approach to the analysis of hand entheseal three-dimensional form. American Journal of Physical Anthropology, 166, 246-260.

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Oxilia G, Bortolini E, Martini S, Papini A, Boggioni M, Buti L, Figus C, Sorrentino R, Townsend G, Kaidonis J, Fiorenza L, Cristiniani E, Kullmer O, Moggi-Cecchi J, Benazzi S (2018) The physiological linkage between molar inclination and dental macrowear pattern. American Journal of Physical Anthropology, 166, 941-951.

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Porraz G, Val A, Tribolo C, Mercier N, de la Pena P, Haaland MM, Igreja M, Miller CE, Schmid VC (2018) The MIS5 Pietersburg at “28” Bushman Rock Shelter, Limpopo Province, South Africa. PLOS One, 13, e02028.

Posth C, Nakatsuka N, Lazaridis I, Skoglund P, Mallick S, Lamnidis TC, Rohland N, et al. EA, Harvati K, et al. EA, Krause J, Reich D (2018) Reconstructing the deep population history of Central and South America. Cell, 175, 1185+.

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Rocholl A, Schaltegger U, Gilg HA, Wijbrans JR, Böhme M (2018) The age of volcanic tuffs from the Upper Freshwater Molasse (North Alpine Foreland Basin) and their possible use for tephrostratigraphic correlations across Europe for the Middle Miocene. International Journal of Earth Sciences, 107, 387-407.

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Schuenemann VJ, Lankapalli AK, Barquera R, Nelson EA, Hernandez DI, Alonzo CA, Bos KI, Morfin LM, Krause J (2018) Historic Treponema pallidum genomes from Colonial Mexico retrieved from archaeological remains. PLOS Neglected Tropical Diseases, 12, e00064.

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Thompson N, Tourloukis V, Panagopoulou E, Harvati K (2018) In search of Pleistocene remains at the Gates of Europe: Directed surface survey of the Megalopolis Basin (Greece). Quaternary International, 497, 22-32.

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Tourloukis V, Thompson N, Panagopoulou E, Giusti DKonidaris GE, Karkanas P, Harvati K (2018) Lithic artifacts and bone tools from the Lower Paleolithic site Marathousa 1, Megalopolis, Greece: Preliminary results. Quaternary International, 497, 47-64.

Trauth MH, Foerster V, Junginger A, Asrat A, Lamb H, Schaebitz F (2018) Abrupt or Gradual? Change Point Analysis of the Late Pleistocene-Holocene Climate Record from Chew Bahir, Southern Ethiopia. Quaternary Research, 90, 321-330.

Vercoutère C, Wolf S (2018) Gravettian tear-drop-shaped beads Perles gravettiennes en forme de larme. L’Anthropologie, 122, 385-401.

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Zanolli C, Pan L, Dumoncel J, Kullmer O, Kundrát M, Liu W, Macchiarelli R, Mancini L, Schrenk F, Tuniz C (2018) Inner tooth morphology of Homo erectus from Zhoukoudian. New evidence from an old collection housed at Uppsala University, Sweden. Journal of Human Evolution, 116, 1-13.


Bocherens H, Schrenk F, Chaimane Y, Kullmer O, Mörike D, Pushkina D, Jaeger JJ (2017) Flexibility of diet and habitat in Pleistocene South Asian mammals: Implications for the fate of the giant fossil ape. Quaternary International, 434, 148-155.

Böhme M, Spassov N, Ebner M, Geraads D, Hristova L, Kirscher U, Kötter S, Linnemann UPrieto J, Roussiakis SL, Theodorou GE, Uhlig G, Winklhofer M (2017) Messinian age and savannah environment of the possible hominin Graecopithecus from Europe. PLOS One, 12, e0177347.

Bretzke K, Kandel AW, Conard NJ (2017) The Middle Paleolithic sequence of Wadi Mushkuna Rockshelter and its implications for hominin settlement dynamics in western Syria. Quaternary International, 435, 106-114.

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Gosh R, Bruch AAPortmann FT, Bera S, Paruya DK, Morthekai P, Ali SN (2017) A modern pollen–climate dataset from the Darjeeling area, eastern Himalaya: Assessing its potential for past climate reconstruction. Quaternary Science Reviews, 174, 63-79.

Grabowski M, Jungers W (2017) Evidence of a chimpanzee-sized ancestor of humans but a gibbon-sized ancestor of apes. Nature Communications, 8, 880.

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Hublin JJ, Ben-Ncer A, Bailey S, Freideline S, Neubauer S, Skinner MM, Bergmann I, le Cabec A, Benazzi S, Harvati K, Gunz P (2017) New fossils from Jebel Irhoud (Morocco) and the Pan-African origin of Homo sapiens. Nature, 546, 289-292.

Kandel AW, Bretzke KConard NJ (2017) Epipaleolithic shell beads from Damascus Province, Syria. Quaternary International, 464, 126-140.

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Karakostis FA, Hotz G, Scherf HWahl JHarvati K (2017) Occupational manual activity is reflected on the patterns among hand entheses. American Journal of Physical Anthropology, 164, 30-40.

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Kirscher U, Oms O, Bruch AA, Shatilova II, Chochishvili G, Bachtadse V (2017) The Calabrian in the Western Transcaucasian basin (Georgia): Paleomagnetic constraints from the Gurian regional stage. Quaternary International, 160, 96-107.

Konidaris GE, Roussiakis SL, Athanassiou A, Theodorou GE (2017) Reprint of: The huge-sized deinothere Deinotherium proavum (Proboscidea, Mammalia) from the Late Miocene localities Pikermi and Halmyropotamos (Greece). Quaternary International, 445, 5-22.

Konidaris GE, Roussiakis SL, Athanassiou A, Theodorou GE (2017) The huge-sized deinothere Deinotherium proavum (Proboscidea, Mammalia) from the Late Miocene localities Pikermi and Halmyropotamos (Greece). Quaternary International, 430, 5-21.

Münzel SC, Wolf S, Drucker DG, Conard NJ (2017) The exploitation of mammoth in the Swabian Jura (SW-Germany) during the Aurignacian and Gravettian period. Quaternary International, 445, 184-199.

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Sahle Y, El Zaatari S, White TD (2017) Hominid butchers and biting crocodiles in the African Plio–Pleistocene. Proceedings of the National Academy of Sciences of the United States of America, 114, 13164-13169.

Schünemann V, Peltzer A, Welte B, van Pelt PW, Molak M, Wang CC, Furtwängler A, Urban C, Reiter E, Nieselt K, Teßmann B, Francken MHarvati K, Haak W, Schiffels S, Krause J (2017) Ancient Egyptian mummy genomes suggest an increase of Sub-Saharan African ancestry in post-Roman periods. Nature Communications, 8, 15694.

Singh N, Albert FW, Plyusnina I, Trut L, Pääbo S, Harvati K (2017) Facial shape differences between rats selected for tame and aggressive behaviors. PLOS One, 12, e0175043.

Starkovich B (2017) Paleolithic Subsistence Strategies and Changes in Site Use at Klissoura Cave 1 (Peloponnese, Greece). Journal of Human Evolution, 111, 63-84.

Starkovich BM, Ntinou M (2017) Climate change, human population growth, or both? Upper Paleolithic subsistence shifts in southern Greece. Quaternary International, 428, 17-32.

Swarts K, Gutaker RM, Benz B, Blake M, Bukowski R, Holland J, Kruse-Peeples M, Lepak N, Prim L, Romay M, Rosslbarra J, et al. EA, Schünemann V, Krause J, et al. EA, Burbano HA (2017) Genomic estimation of complex traits reveals ancient maize adaptation to temperate North America. Science, 357, 512-515.

Teodoridis V, Bruch AA, Kvacek Z, Vassio E, Martinetto E, Stuchlik L (2017) Plio-Pleistocene floras of the Vildštejn Formation in the Cheb Basin, Czech Republic — A floristic and palaeoenvironmental review. Palaeogeography, Palaeoclimatology, Palaeoecology, 467, 166-190.

Toffolo MB, Regev L, Mintz E, Poduska K, Shahack-Gross R, Berthold C, Miller CE, Boaretto E (2017) Accurate Radiocarbon dating of archaeological ash using pyrogenic aragonite. Radiocarbon, 59, 231-249.

Valdivia LB, Douka K, Comeskey D, Bazgir B, Conard NJ, Marean CV, Oll A, Otte M, Tumung L, Zeidi Kulehparcheh M, Higham TFG (2017) Chronometric investigations of the Middle to Upper Paleolithic transition in the Zagros Mountains using AMS radiocarbon dating and Bayesian age modelling. Journal of Human Evolution, 109, 57-69.

Vasilyan D, Zazhigin V, Böhme M (2017) Neogene amphibians and reptiles (Caudata, Anura, Gekkota, Lacertilia, and Testudines) from the south of Western Siberia, Russia, and Northeastern Kazakhstan. PeerJ, 5, e3025.

Villagran XS, Strauss A, Miller CE, Ligouis B, Oliveira R (2017) Buried in ashes: Site formation processes at Lapa do Santo rockshelter, east-central Brazil. Journal of Archaeological Science, 77, 10-34.

Volmer R, Hölzchen E, Wurster A, Ferreras MR, Hertler C (2017) Did Panthera pardus (Linnaeus, 1758) become extinct in Sumatra because of competition for prey? Modeling interspecific competition within the Late Pleistocene carnivore guild of the Padang Highlands, Sumatra. Palaeogeography, Palaeoclimatology, Palaeoecology, 487, 175-186.

Weide A, Riehl SZeidi Kulehparcheh MConard NJ (2017) Reconstructing subsistence practices: taphonomic constraints and the interpretation of wild plant remains at aceramic Neolithic Chogha Golan, Iran. Vegetation History and Archaeobotany, 26, 487-504.

Weyrich LS, Duchene S, Soubrier J, Arriola L, Llamas B, Breen J, Morris AG, Alt KW, Caramelli D, Dresely V, Farrell M, Farrer AG, Francken M, Gully N, Haak W, Hardy K, Harvati K, Held P, Holmes EC, Kaidonis J, Lalueza-Fox C, de la Rasilla M, Rosas A, Semal P, Soltysiak A, Townsend G, Usai D, Wahl J, Huson DH, Dobney K, Cooper A (2017) Neanderthal behaviour, diet, and disease inferred from ancient DNA in dental calculus. Nature, 544, 357-361.

Wright DK, Thompson JC, Schilt F, Cohen AS, Jeong-Heon C, et. al. EA, Miller CEMentzer SM, et. al. EA, Gomani-Chindebvu E (2017) Approaches to Middle Stone Age landscape archaeology in tropical Africa. Journal of Archaeological Science, 77, 64-77.

Peer-reviewed Articles / Non-ISI


Bader GD, Linstädter J, Schoeman A (2020) Uncovering the late Pleistocene LSA of Mpumalanga province, South Africa. Early results from Iron Pig Rock Shelter. Journal of African Archaeology, 18, 19-37.

Conard NJ, Janas A, Zeidi Kulehparcheh M (2020) Ausgrabungen an der Langmahdhalde im Lonetal: Raumnutzung und Siedlungsdynamik im Magdalénien. Archäologische Ausgrabungen in Baden-Württemberg 2019, 51-55.

Conard NJ, Malina M (2020) Fortsetzung der Ausgrabungen am Hohle Fels und neue aurignacienzeitliche Werkzeuge aus Mammutelfenbein. Archäologische Ausgrabungen in Baden-Württemberg 2019, 47-51.

Fuentes R, Ono R, Carlos J, Kerfant C, Sriwigati A, Miranda T, Aziz N, Sofian HO, Pawlik AF (2020) Stuck within notsches: Direct evidence of plant processing during the last glacial maximum to Holocene in North Sulawesi. Jouranal of Archaeological Science: Reports, 39.

Haidle MN, Schlaudt O (2020) Where Does Cumulative Culture Begin? A Plea for a Sociologically Informed Perspective. Biological Theory.

Hutterer R, Schröder O, Linstädter J (2020) Food and Ornament: Use of Shellfish at Ifri Oudadane, a Holocene Settlement in NE Morocco. African Archaeological Review 38, 73–94.

Lehmann J, Verheijen I, Serangeli J, Mertens W, Conard NJ (2020) Schöningen FStNr. 13II, Gde. Stadt Schöningen, Ldkr. Helmstedt. Fundchronik Niedersachsen 2018, 22, 151-152.

Masini F, Maul LC, Abbazzi L, Petruso D, Savorelli A (2020) Independent water vole (Mimomys savini, Arvicola: Rodentia, Mammalia) lineages in Italy and Central Europe. Fossil Imprint, 76, 59-83.

Pomeroy E, Hunt CO, Reynolds T, Abdulmutalb D, Asouti E, Bennett P, Bosch M, Burke A, Farr L, Foley R, French C, Frumkin A, Goldberg P, Hill E, Kabukcu C, Lahr M, Lane R, Marean CV, Maureille B, Mutri G, Miller CE, Mustafa KA, et al. EA, Barker GM (2020) Issues of theory and method in the analysis of Paleolithic mortuary behavior: A view from Shanidar Cave. Evolutionary Anthropology, 29, 263-279.

Porraz G, Parkington JE, Schmidt P, Bereiziat G, Brugal JP, Dayet L, Igreja M, Miller CE, Schmid VC, Tribolo C, Val A, Verna C, Texier PJ (2020) Experimentation preceding innovation in a MIS5 Pre-Still Bay layer from Diepkloof Rock Shelter (South Africa): emerging technologies and symbols. EcoEvoRxiv.

Schlaudt O (2020) Type and Token in the Prehistoric Origins of Numbers. Cambridge Archaeological Journal, 30, 629-646.

Schmidt P, Stynder D, Conard NJ, Parkington JE (2020) When was silcrete heat treatment invented in South Africa? Palgrave Communications, 6, 1-10.

Serangeli J, Conard NJ (2020) Forschung in Schöningen, Steinzeit trifft Moderne. Denkmalpflege, Berichte zur Denkmalpflege in Niedersachsen, 1, 8-11.

Serangeli J, Verheijen I, Rodríguez Álvarez B, Altamura F, Lehmann J, Conard NJ (2020) Elefanten in Schöningen. Archäologie in Deutschland, 3, 8-13.

Stahlschmidt MC, Mallol CM, Miller CE (2020) Fire as an Artifact—Advances in Paleolithic Combustion Structure Studies: Introduction to the Special Issue. Journal of Paleolithic Archaeology.

Willmes C, Niedziolka K, Serbe B, Grimm SB, Groß D, Miebach A, Märker M, Henselowsky F, Gamisch A, Rostami M, Mateos A, Rodriguez J, Limberg H, Schmidt I, Müller M, Hölzchen E, Holthausen M, Klein K, Wegener C, Weninger B, Kellberg Nielsen T, Otto T, Weniger GC, Bubenzer O, Bareth G (2020) State of the Art in Paleoenvironment Mapping for Modeling Applications in Archeology—Summary, Conclusions, and Future Directions from the PaleoMaps Workshop . Quaternary, 3, 13.


Conard NJZeidi Kulehparcheh M (2019) New Research on the Palaeolithic Occupation of Ghar-e Boof, Fars Province. Archaeology, 2.

Etu-Sihvola H, Bocherens HDrucker DG, Junno AP, Mannermaa K, Oinonen M, Uusitaol J, Arppe L (2019) The dIANA database – isotopic baseline for paleodietary research in the Baltic Sea area. Journal of Archaeological Science: Reports, 24, 1003-1013.

Pothier Bouchard G, Mentzer SM, Buckley M, Wogelius M, Negrino F, Hodgkins J, Riel-Salvatore J, Miller CE (2019) Portable FTIR for on-site screening of archaeological bone intended for ZooMS collagen fingerprint analysis. Journal of Archaeological Science: Reports, 24.

Sahle Y, Giusti D, Gossa T, Ashkenazy H (2019) Exploring karst landscapes: new prehistoric sites in south-central Ethiopia. Antiquity Project Gallery, 93, 1-6.

Sahle Y, Giusti D, Tourloukis V (2019) Newly discovered Middle Pleistocene hominid-bearing deposits from the Lower Awash basin, Ethiopia. Anthropological Science.

Taller A, Conard NJ (2019) Transition or Replacement? Radiocarbon dates from Hohle Fels Cave (Alb-Donau-Kreis/ DI and the Passage from Aurignacian to Gravettian. Archäologisches Korrespondenzblatt, 49, 165-181.

Wolf S (2019) Bemalte Steine aus dem Magdalénien der Hohle Fels Höhle bei Schelklingen. Laichinger Höhlenfreund, 53, 51-62.


Bader GD, Tribolo C, Conard NJ (2018) A return to Umbeli Belli: New insights of recent excavations and implications for the final MSA of eastern South Africa. Journal of Archaeological Science: Reports, 21, 733-757.

Beck J, Díaz-Zorita Bonilla M, Bocherens H, Diaz-del-Rio P (2018) Feeding a third millennium BC mega-site: Bioarchaeological analyses of palaeodiet and dental disease at Marroquíes (Jaén, Spain). Journal of Anthropological Archaeology, 52, 23-43.

Blackwell BAB, Sakhrani N, Singh I, Gopalkrishna K, Tourloukis V, Panagopoulou E, Karkanas P, Blickstein JIB, Skinner AR, Florentin JA, Harvati K (2018) ESR Dating Ungulate Teeth and Molluscs from the Paleolithic Site Marathousa 1, Megalopolis Basin, Greece. Quaternary, 22, 1-25.

Chala-Aldana D, Bocherens HMiller CE, Moore K, Hodgins G, Rademaker K (2018) Investigating mobility and highland occupation at the Cuncaicha rock shelter through Strontium and Oxygen isotopes. Journal of Archaeological Science: Reports, 19, 811-827.

Conard NJ (2018) Science and Paleolithic Archaeology, Harold L. Dibble (July 26, 1951 – June 10, 2018). Mitteilungen der Gesellschaft für Urgeschichte, 27, 189-193.

Conard NJ (2018) Preliminary Reflections on the Life and Times of Hansjürgen Müller-Beck (* 13. August 1927 in Apolda – † 02. August 2018 in Bern). Mitteilungen der Gesellschaft für Urgeschichte, 27, 209-246.

Conard NJ, Janus A, Zeidi Kulehparcheh M (2018) Ausgrabungen in den magdalénienzeitlichen Schichten der Langmahdhalde im Lonetal. Archäologische Ausgrabungen Baden-Württemberg, 55-59.

Conard NJ, Rudolf S, Toniato G (2018) Fortsetzungen der paläolithischen Ausgrabungen am Abri Schafstall II im Lauchertal bei Veringenstadt. Archäologische Ausgrabungen Baden-Württemberg, 60-63.

Díaz-Zorita Bonilla M, Beck J, Bocherens H, Diaz-del-Rio P (2018) Isotopic evidence for human mobility at the Copper Age mega-site of Marroquíes Bajos (Jaén, Spain). Antiquity, 92 (364), 991-1007.

Fiorenza L, Benazzi S, Oxilia G, Kullmer O (2018) Functional relationship between dental macrowear and diet in Late Pleistocene and modern human populations. International Journal of Osteoarchaeology, 28, 153-161.

Giemsch L, Hertler C, Märker M, Quénéhervé G, Saanane C, Schrenk F (2018) Acheulean Sites at Makuyuni (Lake Manyara, Tanzania): Results of Archaeological Fieldwork and Classification of the Lithic Assemblages. African Archaeological Review, 35, 87-106.

Lombard M, Högberg A, Haidle MN (2018) Cognition: From Capuchin Rock Pounding to Lomekwian Flake Production. Cambridge Archaeological Journal, 1-31.

Schillinger B, Beaudet A, Fedrigo A, Grazzi F, Kullmer O, Laaß M, Makowska M, Werneburg I, Zanolli C (2018) Neutron Imaging in Cultural Heritage research at the FRM II reactor of Heinz Maier-Leibnitz Center. Journal of Imaging, 4, 22.

Wolf S, Dapschauskas R, Velliky E, Floss H, Kandel AW, Conard NJ (2018) The use of ochre and painting during the Upper Paleolithic of the Swabian Jura in the context of the development of ochre use in Africa and Europe. Open Archaeology, 4, 185-205.


Bretzke KConard NJ (2017) Not Just a Crossroad: Population Dynamics and Changing Material Culture in Southwestern Asia during the Late Pleistocene. Current Anthropology, 58, 449-462.

Camarós E, Cueto M, Rosell J, Díez JC, Blasco R, Duhig C, Darlas A, Harvati K, EA, Rivals F (2017) Hunted or Scavenged Neanderthals? Taphonomic Approach to Hominin Fossils with Carnivore Damage. International Journal of Osteoarchaeology, 27, 606-620.

Conard NJ (2017) The path to UNESCO World Cultural Heritage status for the Caves and Ice Age Art in the Swabian Jura. Mitteilungen der Gesellschaft für Urgeschichte, 26, 153-168.

Conard NJ, Janas A, Rudolf S (2017) Weitere Grabungsergebnisse zu den Aurignacienzeitlichen Schichten vom Hohle Fels bei Schelklingen. Archäologische Ausgrabungen Baden-Württemberg 2016, 54-58.

Conard NJ, Janas A, Zeidi Kulehparcheh M (2017) Neue Ausgrabungen unter den Felsdächern in der Langmahdhalde bei Lonetal. Archäologische Ausgrabungen Baden-Württemberg 2016, 58-63.

Conard NJ, Rudolf S, Toniato G (2017) Neu Ausgrabungen in den Felsdächern Schafstall I und II in Veringenstadt. Archäologische Ausgrabungen Baden-Württemberg 2016, 58-63.

Goldberg P, Miller CEMentzer SM (2017) Recognizing fire in the Paleolithic archaeological record. Current Anthropology, 58, 175-190.

Haidle MN (2017) Development of teaching performance: Comment on P. Gärdenfors and A. Högberg, The archaeology of teaching and the evolution of Homo docens. Current Anthropology, 58, 202-204.

Karakostis FA, Velliky E, Kandel AW (2017) Sixth annual meeting of the European Society for the Study of Human Evolution. Evolutionary Anthropology, 26, 7-8.

Stahlschmidt MC, Miller CE, Kandel AW, Conard NJ (2017) Site formation processes and Late Natufian domestic spaces at Baaz Rockshelter, Syria: A micromorphological perspective. Journal of Archaeological Science: Reports, 12, 499-514.

Starkovich BConard NJ (2017) What were they up against? Lower Paleolithic hominin meat acquisition and competition with Plio-Pleistocene carinvores. RGZM.

Zanolli C, Schillinger B, Beaudet A, Kullmer O, Macchiarelli R, Mancini L, Schrenk F, Tuniz C, Vodopivec V (2017) Exploring hominin and non-hominin primate dental fossil remains with neutron microtomography. Physics Procedia, 88, 109-115.

Articles in other Journals


Hayrapetyan A, Kvavadze EV, Shatilova II, Gabrielyan I, Bruch AA (2020) Subfossil palynological spectra from the surroundings of the village Tsovinar (lake Sevan, Armenia). Biological Journal of Armenia, 72, 52-58.

Lockey AL, Limmer LS, McCartin M, Mentzer SM (2020) Ninth Annual meeting of the European Society for the Study of Human Evolution. Evolutionary Anthropology, 29, 46-48.

Marianyan MA, Harutyunyan RG, Bruch AA, Gabrielyan I (2020) To the knowledge of the Paleofauna of the Subfamily Lixinae (Coleoptera, Curculionidae) of Sisian Suite (Southern Armenia, Early Pleistocene). Biological Journal of Armenia, 72, 110-114.


Bader GDConard NJ (2019) Umbeli Belli Rock Shelter in KwaZulu-Natal – Current results on the final MSA of southern Africa. Roceeh Newsletter 15-2019.

Conard NJ, Malina M (2019) Weiterführende Ausgrabungen im Hohle Fels bei Schelklingen und neue Einblicke in die Nutzung von Ocker im Jungpaläolithikum. Archäologische Ausgrabungen in Baden-Württemberg 2018, 56-59.

Ioannidou M, Falcucci A, Röding C, Kandel AW (2019) Eighth Annual Meeting of the European Society for the Study of Human Evolution. Evolutionary Anthropology Issues News and Reviews, 28.

Lehmann J, Serangeli J, Conard NJ (2019) Schöningen FStNr. 13II, Gde. Stadt Schöningen, Ldkr. Helmstedt. Fundchronik Niedersachsen 2017, 22, 135-139.


Bocherens HDrucker DG, Rivals F (2018) L’alimentation et l’environnement (micro-usure et isotopes) des mammouths et elephants du Pléistocène. Catalogue de l’exposition “Mémoires de Mammouths”, Musée national de Préhistoire Les Eyzies-de-Tayac, 35-39.

Conard NJ (2018) Als die moderne Kultur Begann. Die Anfänge der Kunst und der Musik sowie die Bedeutung der Funde aus den Höhlen der Schwäbischen Alb. Bewegte Zeiten. Archäologie in Deutschland, 338-349.

Conard NJ, Janas A (2018) Fortsetzung der Ausgrabungen am Hohle Fels und die Entdeckung einer markierten Mammutrippe aus dem Gravettien. Archäologische Ausgrabungen Baden-Württemberg, 52-55.

Terlato G, Bocherens H, Romandini M, Nannini N, Hobson KA, Peresani M (2018) Extinction and paleoecology of the Late Pleistocene cave bear from northeastern Italy: Radiocarbon and stable isotope evidence. Alpine and Mediterranean Quaternary, 31, 98-103.

Zeidi Kulehparcheh M (2018) Report on joint Iran-Japan Paleolithic excavations in Bolaghi valley. Parsa Pasargadae World Heritage Site Newsletter, 7, 4-7.


Conard NJ (2017) Cultural Developments during the Terminal Pleistocene and Early Holocene in the Central Levant and the Western Foothills of the Zagros Mountains. Neolithic World Cultural Heritage Sites.

Conard NJ (2017) The contributions of Hans-Peter Uerpmann to the Archaeozoology of the Near East. Archaeozoology of the Near East.

Ghasidian E, Bretzke KConard NJ (2017) Excavations at Ghar-e Boof in the Fars Province of Iran and its bearing on models for the evolution of the Upper Palaeolithic in the Zagros Mountains. Journal of Anthropological Archaeology, 47, 33-49.

Karakostis FA, Lorenzo C, Moraitis K (2017) Morphometric variation and ray allocation of human proximal hand phalanges. Anthropologischer Anzeiger.

Marwik B, d’Alpoim Guedes J, Barton DM, Bates LA, Baxter M, et. al. EA, Giusti D (2017) Open Science in Archaeology. SAA Archaeological Record, 17, 8-14.

Maul LC, Löscher M, Keller T, Henkel T, Schreiber HD (2017) Mauer – Mosbach – Voigtstedt. Drei überregional bedeutende Fundstellen mit frühmittelpleistozänen Kleinsäugerfaunen aus Zentraleuropa. Palaeos, 6, 18-26.

Napierala H, Kandel AW, Conard NJ (2017) Small game and schifting subsistence patterns from the Upper Paleolithic to the Natufian at Baaz Rockshelter, Syria. Archaeozoology of the Near East.



Conard NJ, Wolf S (2020) Der Hohle Fels in Schelklingen: Anfänge der Kunst und Musik. Tübingen: Kerns Verlag, 104pp.




Harvati K, Jäger G, Reyes-Centeno H (2018) New Perspectives on the Peopling of the Americas. Kerns Verlag, Tübingen, 224pp.


Conard NJ, Kind CJ (2017) Als der Mensch die Kunst erfand: Eiszeithöhlen der Schäbischen Alb. Wissenschaftliche Buchgesellschaft, Darmstadt, 192pp.

Shatilova II, Kvavadze EV, Kokolashvili IM, Bruch AA (2017) Atlas of Pollen of the Georgian upper Cenozoic – Gymnosperms and Angiosperms. Georgian National Museum, Tblisi, Georgia, 379pp.

Book section


Fiorenza L, Benazzi S, Estalrrich A, Kullmer O (2020) Diet and cultural diversity in Neanderthals and modern humans from dental macrowear analyses. In: Schmidt CW, Watson JT (Eds) Dental Wear in Evolutionary and Biocultural Contexts. Elsevier, 39-72.

Germonpre M, Lázniçková-Galetová M, Sablin MV, Bocherens H (2020) Could incipient dogs have enhanced differential access to resources among Upper Palaeolithic hunter-gatherers in Europe? In: Moreau L (Eds) Social Inequality before farming? Multidisciplinary approaches to the investigation of egalitarian and non-egalitarian social relationships in prehistoric and extant hunter-gatherer societies. McDonald institute of Archaeology, Cambrigde.

Höflmayer F, Dee MW, Riehl S (2020) The Radiocarbon Dating of the Middle Bronze Age Monumental Building. In: Kamlah J, Sader H (Eds) Tell el-Burak I. The Middle Bronze Age, Harrassowitz, Wiesbaden, 210-226.

Kamlah J, Riehl S (2020) Agriculture in the Bronze Age Levant (ch. 10). In: Hollander D, Howe T (Eds) A Companion to Ancient Agriculture. John Wiley & Sons, Inc., 193-209.

Kullmer O, Menz U, Fiorenza L (2020) Occlusal Fingerprint Analysis (OFA) Reveals Dental Occlusal Behavior in Primate Teeth. Mammalian Teeth – Form and Function. Dr. Friedrich Pfeil München.

Martinetto E, Bertini A, Bhandari S, Bruch AA, Cherin M, Field JH, Gabrielyan I, Gianotti F, Kern AK, Kienast F, Lindsey EL, Momohara A, Ravazzi C, Thomas ER (2020) The last three millions of unequal spring thaws. In: Martinetto E, Tschopp E, Gastaldo RA (Eds) Nature through Time, Virtual field trips through the Nature of the past. Springer International Publishing, Switzerland.

Riehl S (2020) Stable isotopes in ancient agriculture. In: Hollander D, Howe T (Eds) A Companion to Ancient Agriculture. John Wiley & Sons, Inc., 55-81.

Riehl S, Orendi A (2020) Archaeobotanical Samples from Middle and Late Bronze Age Contexts at Tell el-Burak. In: Kamlah J, Sader H (Eds) Tell el-Burak I. The Middle Bronze Age, Harrassowitz, Wiesbaden, 360-368.

Schmidt C, El Zaatari S, Van Sessen A (2020) Dental microwear texture analysis in bioarchaeology. In: Schmidt C, Watson JV (Eds) Dental Wear in Evolutionary and Biocultural Contexts. Academic Press, 143-168.

Starkovich B, Conard NJ (2020) What were they up against? Lower Paleolithic hominin meat acquisition and competition with Plio-Pleistocene carnivores. In: Garcia-Moreno A (Eds) Human behavioural adaptations to interglacial lakeshore environments. Heidelberg: Propylaeum, 105-130.

Taipale N, Rots V, Conard NJ (2020) Cold-climate toolkits: fire-making, lithic recycling and assemblage formation in the Magdalenian of Hohle Fels Cave. In: Gibaja J, Marreiros J, Mazzucco N, Clemente I (Eds) Hunter-gatherers’ tool-kit: a functioanl perspective. Cambrdige Scholar Publishing: Newcastle, 154-176.


Blanco-Lapaz A (2019) Forschungen und Berichte zur Archäologie in Baden-Württemberg. In: Kind CJ (Eds) Löwenmensch und mehr. Die Ausgrabungen 2008-2013 in den altsteinzeitlichen Schichten der Stadel-Höhle im Hohlenstein (Lonetal). Museum Ulm, 175-178.

Bolus M, Conard NJ (2019) Paläolithforschung in den Höhlen der Schwäbischen Alb. Forschungsgeschichte – Kenntnisstand – Ausblick. In: Baales M, Pasda C (Eds) „All der holden Hügel ist keiner mir frend…“ Festschrift zum 65. Geburtstag von Claus-Joachim Kind. Verlag Dr. Rudolf Habelt GmbH Bonn, 43-66.

Bosman AM, Buck L, Reyes-Centeno H, Mirazón Lahr M, Stringer C, Harvati K (2019) The Kabua 1 cranium: Virtual anatomical reconstructions. In: Sahle Y, Reyes-Centeno H, Bentz C (Eds) Modern Human Origins and Dispersal. Kerns Verlag, Tübingen, 137-170.

Conard NJ (2019) Excavations at Geißenklösterle Cave. In: Conard NJ, Bolus M, Münzel SC (Eds) Geißenklösterle: Chronostratigraphie, Paläoumwelt und Subsistenz imMittel- und Jungpaläolithikum der Schwäbischen Alb / Chronostratigraphy, Paleoenvironment and Subsistence during the Middle amd Upper Paleolithic of the Swabian Jura. Kerns Verlag, Tübingen, 9-21.

El Zaatari S, Harvati K (2019) Forschungen und Berichte zur Archaeologie in Baden-Württemberg. In: Kind CJ (Eds) Löwenmensch und mehr. Die Ausgrabungen 2008-2013 in den altsteinzeitlichen Schichten der Stadel-Höhle im Hohlenstein (Lonetal). Reichert Verlag Stuttgart, 171-174.

Goldberg P, Miller CEConard NJ (2019) Geißenklösterle Stratigraphy and Micromorphology. In: Conard NJ, Bolus M, Münzel SC (Eds) Geißenklösterle: Chronostratigraphie, Paläoumwelt und Subsistenz imMittel- und Jungpaläolithikum der Schwäbischen Alb / Chronostratigraphy, Paleoenvironment and Subsistence during the Middle amd Upper Paleolithic of the Swabian Jura. Kerns Verlag, Tübingen, 25-61.

Hahn K, Schmidt M, Thiombiano A (2019) Use patterns of wild food plants in West Africa. .) Trees, grasses and crops. People and plants in Sub-saharan Africa and beyond. Dr. Rudolf Habelt GmbH, Bonn., Bonn, 157-172.

Haidle MN (2019) The origin of cumulative culture – not a single-trait event, but multifactorial processes. In: Overman K, Coolidge FL (Eds) Squeezing minds from stones. Oxford University Press, Oxford, 128-148.

Höflmayer F, Dee MW, Riehl S (2019) Radiocarbon dates. In: Yasur-Landau A, Cline EH (Eds) Tel Kabri II. The 2005-2011 Excavation Seasons.

Höhn A, Hahn K, Schmidt M (2019) The search for Isoberlinia – investigations into today’s woody vegetation of the Nok project’s key research area in Central Nigeria. Trees, grasses and crops. People and plants in Sub-saharan Africa and beyond. Dr. Rudolf Habelt GmbH, Bonn., Bonn, 221-232.

Riehl S (2019) Resource management at Tell el-Abd. Economic specialization despite or because of environmental limitations? In: Finkbeiner U (Eds) Final Reports of the Syrian-German Excavations at Tell el-ʻAbd. Zaphon, 169-185.

Sala N, Conard NJ (2019) Mortuary practices and the culture of death in the Swabian Jura during the Magdalenian. In: Floss H (Eds) Das Magdalénien im Südwesten Deutschlands, im Elsass und in der Schweiz. Kerns Verlag, Tübingen, 225-247.

Schmidt C, El Zaatari S, Van Sessen A (2019) Dental microwear texture analysis in bioarchaeology. In: Schmidt C, Watson JV (Eds) Dental Wear in Evolutionary and Biocultural Contexts. Academic Press, 143-168.

Taller A, Conard NJ (2019) Das Magdalénien im Hohle Fels. In: Baales M, Pasda C (Eds) „All der holden Hügel ist keiner mir frend…“ Festschrift zum 65. Geburtstag von Claus-Joachim Kind. Verlag Dr. Rudolf Habelt GmbH Bonn, 285-299.

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Work and Discussion Paper




Conard NJ, Janas A, Zeidi Kulehparcheh M (2019). Ausgrabungen an der Langmahdhalde liefern weitere Einblicke in magdalénienzeitliche Lebensweisen auf der Schwäbischen Alb., Arch. Ausgr. Baden-Württemberg 2018, 4pp.