Senckenberg am Meer, Wilhelmshaven & Hamburg

Library Senckenberg Wilhelmshaven

Marine Research Division

The library is a scientific reference library with a focus on marine geology and marine biology. The inventory comprises approx. 3 450 book titles, 500 maps and 250 journals and series, and approx. 26 000 separate prints. Archive material from estates is incorporated into the inventory. The main task is to provide the staff of SaM with literature. We do not take part in interlibrary loan.

Our library is connected to WIFI (Wilhelmshaven Fachinformationseinrichtungen und wissenschaftliche Bibliotheken) and to AmB (German Association of Marine Science Libraries and Information Centers) and also to the Senckenberg working group Libraries and Exchange of Publications.

The stock of Marine Research Division includes approx. 3,100 book titles, 420 maps, 330 journals and series as well as approx. 21,500 reprints.

The DZMB’s stock includes around 400 books / monographs and around 2,800 reprints. With approx. 23,000 titles, the Monoculus Library is one of the largest collections of literature in the world on the subject of copepods.

Archival material from bequests is incorporated into the library holdings.

The library has permanent access to Senckenberg licences and the DFG national licenses. Interlibrary loans are made through the GBV. The electronic recording of the library stock is done with JabRef: an open source bibliography reference manager.

The use for external users is possible after registering by telephone. Guided tours through the library are only possible by prior arrangement.

Scientific direction

Prof. Dr.  André Freiwald
Direktor Senckenberg am Meer; Abteilungsleiter Meeresforschung; Fachgebietsleiter Meeresgeologie

Functions within Senckenberg:

  • Director of SaM
  • Head of Division Marine Geology (SaM)
  • Speaker of RA 4.1: Evolving Earth and Environment

Research profile

  • Non-tropical biosedimentary systems
  • Cold-water coral ecosystems
  • Bioerosion
  • Paleoecology

Research interests

My research focuses on geobiological studies of recent and fossil cold-water corals, with a focus on occurrences along the increasingly oxygen-poor upwelling areas of West Africa (Mauritania and Angola-Namibia). Another topic is research on the quantitative degradation of carbonate by biological processes (bioerosion). Finally, we are developing a global database on habitat-forming cold-water coral and associated communities across all phyla. In this database, the current state of knowledge of the composition of the reef inhabitants from the unicellular organisms to the fish, their geographical changes and their biological functions will be depicted for the first time.

Professional positions
Since 2018 Director Senckenberg am Meer, Wilhelmshaven
Since 2010 Head of Division Meeresforschung at Senckenberg am Meer, Wilhelmshaven
Since 2010 Professor, University of Bremen
2002-2010 Professor, Friedrich-Alexander-Universität Erlangen-Nürnberg
2000-2002 Professor, University of Tübingen

2000 Heisenberg Fellow
1999 Habilitation University of Bremen
1993-1999 Researcher at University of Bremen
1993 PhD, Kiel University

Relevant scientific projects  2017-2019
2018-2019 Atlantic Thermocline Ocean and Ecosystems Dynamic during Natural Climate
Change (ATHENA) (DFG) (externally administered through University of
2019-2022 Cold-water Coral sensitivity (MARUM Excellence Cluster)
2019-2022 West-African Biodiversity under Pressure (WASP) (GIZ)

Professional service, membership associations, cooperation in national and intl. committees
Since 2017 Advisory Board of the Northwest German University Society
Since 2015 Chairman of the DFG-BMBF Advisory Councils of the research vessels METEOR
Since 2015 of the BMBF advisory board research vessel SONNE
2011-2015 Supervisory Board at the Center for Tropical Marine Ecology
Since 2010 Member of the MARUM board of the University of Bremen
​2008-2011 Speaker of DFG Geoforum
2004-2008 Member of the DFG Review Board Geology- Paleontology
2001-2006 Member of the DFG Senate Commission for Oceanography
2000-2003 Coordinator of the EU-FP5 Atlantic Coral Ecosystem Study (ACES) project
2019 Foundation member of the Deutsche Allianz Meeresforschung

Editorial activities
2005-2010 Publisher of the Springer book series “Erlangen Earth Conference Series”
2004-2010 Publisher of the ISI journal FACIES together with Springer

Awards and awarded memberships
2016 Gustav-Steinmann-Medal of the Geological Association
2015 Northern German Science Award for innovative research on cold-water corals


Tief- und Kaltwasserkorallen: ihre Lebensgemeinschaften, Bildungs- und Umweltbedingungen sowie Einwirkungen des Menschen

Addamo AM, Reimer JD, Taviani M, Freiwald A, Machordom A (2012) Desmophyllum dianthus (Esper, 1794) in the scleractinian phylogeny and its intraspecific diversity. Plos One 7: e50215

Barnich R, Beuck L, Freiwald A (2013) Scale worms (Polychaeta: Aphroditiformia) associated with cold-water corals in the eastern Gulf of Mexico. Journal of the Marine Biological Association of the United Kingdom 93: 2129-2143

Beuck L, Freiwald A (2005) Bioerosion patterns in a deep-water Lophelia pertusa (Scleractinia) thicket (Propeller Mound, northern Porcupine Seabight). In: Freiwald A, Roberts JM (eds) Cold-water corals and ecosystems. Springer, pp 915-936

Beuck L, Freiwald A, Taviani M (2010) Spatiotemporal bioerosion patterns in deep-water scleractinians from off Santa Maria di Leuca (Apulia, Ionian Sea). Deep-Sea Research II 57: 458-470

Beuck L, López Correa M, Freiwald A (2008) Biogeographical distribution of Hyrrokkin (Rosalinidae, Foraminifera) and its host-specific morphological and textural trace variability. In: Wisshak M, Tapanila L (eds) Current Developments in Bioerosion. Springer, Heidelberg, pp 329-360

Copard K, Colin C, Douville E, Freiwald A, Gudmundsson G, De Mol B, Frank N (2010) Nd isotopes in deep-sea corals in the North-eastern Atlantic. Quaternary Science Reviews 29: 2499-2508

Cordes EE, Arnaud-Haond S, Bergstad OA, Da Costa Falcao AP, Freiwald A, Roberts JM, Bernal P (2016) Cold-water corals. In: Inniss L, Simcock A, Ajawin AY, et al. (eds) World Ocean Assessment I. United Nations, pp 1-28

Costello MJ, McCrea M, Freiwald A, Lundälv T, Jonsson L, Bett BJ, van Weering TCE, de Haas H, Roberts JM, Allen D (2005) Role of cold-water Lophelia pertusa coral reefs as fish habitat in the NE Atlantic. In: Freiwald A, Roberts JM (eds) Cold-water corals and ecosystems. Springer, Heidelberg, pp 771-805

Cuif JP, Dauphin Y, Freiwald A, Gautret P, Zibrowius H (1999) Biochemical markers of zooxanthellae symbiosis in soluble matrices of skeleton of 24 Scleractinia species. Comparative Biochemistry and Physiology, Part A: Physiology 123: 269-278

Dorschel B, Hebbeln D, Rüggeberg A, Dullo WC, Freiwald A (2005) Growth and erosion of a cold-water coral covered carbonate mound in the Northeast Atlantic during the Late Pleistocene and Holocene. Earth and Planetary Science Letters 233: 33-44

Eisele M, Frank N, Wienberg C, Hebbeln D, López Correa M, Douville E, Freiwald A (2011) Productivity controlled cold-water coral growth periods during the last glacial off Mauritania. Marine Geology 280: 143-149

Fabri MC, Pedel L, Beuck L, Galgani F, Hebbeln D, Freiwald A (2014) Megafauna of vulnerable marine ecosystems in French Mediterranean submarine canyons: Spatial distribution and anthropogenic impacts. Deep-Sea Research II 104: 184-207

Fink HG, Wienberg C, Hebbeln D, McGregor HV, Schmiedl G, Taviani M, Freiwald A (2012) Oxygen control on Holocene cold-water coral development in the eastern Mediterranean Sea. Deep-Sea Research I 62: 89-96

Försterra G, Beuck L, Häussermann V, Freiwald A (2005) Shallow-water Desmophyllum dianthus (Scleractinia) from Chile: characteristics of the biocoenoses, the bioeroding community, heterotrophic interactions and (Paleo)-bathymetric implications. In: Freiwald A, Roberts JM (eds) Cold-water corals and ecosystems. Springer, Heidelberg, pp 937-977

Frank N, Freiwald A, López Correa M, Wienberg C, Eisele M, Hebbeln D, van Rooij D, Henriet JP, Colin C, van Weering T, De Haas H, Buhl-Mortensen P, Roberts JM, De Mol B, Douville E, Blamart D, Hatte C (2011) Northeastern Atlantic cold water coral reefs and climate. Geology 39: 743-746

Freiwald A (2002) Reef-forming cold-water corals. In: Wefer G, Billett D, Hebbeln D, Jørgensen BB, Schlüter M, van Weering TCE (eds) Ocean Margin Systems. Springer, pp 365-385

Freiwald A (2011) Cold-water coral reefs. In: Hopley D (ed) Encyclopedia of modern coral reefs. Springer, Heidelberg, pp 225-229

Freiwald A (2019) Messinian Salinity Crisis: What happened to cold-water corals? In: Orejas C, Jiménez C (eds) Mediterranean Cold-Water Corals: Past, Present and Future. Springer, pp 47-50

Freiwald A, Beuck L, Rüggeberg A, Taviani M, Hebbeln D (2009) The white coral community in the central Mediterranean Sea revealed by ROV surveys. Oceanography 22: 58-74

Freiwald A, Fosså JH, Grehan A, Koslow T, Roberts JM (2004) Cold-water coral reefs. UNEP-WCMC Biodiversity Series 22: 1-84

Freiwald A, Henrich R, Pätzold J (1997) Anatomy of a deep-water coral reef mound from Stjernsund, West-Finnmark, northern Norway. In: James NP, Clarke JAD (eds) Cool-water carbonates. SEPM, Special Publication, pp 141-161

Freiwald A, Hühnerbach V, Lindberg B, Wilson JB, Campbell J (2002) The Sula Reef complex, Norwegian Shelf. Facies 47: 179-200

Freiwald A, Mortensen PB (2000) The first record of the deep-water coral Stenocyathus vermiformis (Pourtalès, 1868) (Scleractinia, Guyniidae) from Norwegian waters. Sarsia 85: 275-276

Freiwald A, Reitner J, Krutschinna J (1997) Microbial alteration of the deep-water coral Lophelia pertusa: early postmortem processes. In: Neuweiler F, Reitner J, Monty C (eds) Biosedimentology of microbial buildups IGCP Project No. 380 – Proceedings of 2nd Meeting. Facies, pp 223-226

Freiwald A, Rogers A, Hall-Spencer J, Guinotte JM, Davies AJ, Yesson C, Martin CS, Weatherdon LV (2017) Global distribution of cold-water corals (version 5.0). Fifth update to the dataset in Freiwald et al. (2004) by UNEP-WCMC.

Freiwald A, Schönfeld J (1996) Substrate pitting and boring pattern of Hyrrokkin sarcophaga Cedhagen, 1994 (Foraminifera) in a modern deep-water coral reef mound. Marine Micropaleontology 28: 199-207

Freiwald A, Wilson JB (1998) Taphonomy of modern deep, cold-temperate water coral reefs. Historical Biology 13: 37-52

Freiwald A, Wilson JB, Henrich R (1999) Grounding Pleistocene icebergs shape recent deep-water coral reefs. Sedimentary Geology 125: 1-8

Gautret P, Cuif JP, Freiwald A (1997) Composition of soluble mineralizing matrices in zooxanthellate and non-zooxanthellate scleractinian corals: biochemical assessment of photosynthetic metabolism through the study of a skeletal feature. Facies 36: 189-194

Göcke C, Hestetun JT, Uhlir C, Freiwald A, Beuck L, Janussen D (2016) Cladorhiza corallophila sp. nov., a new carnivorous sponge (Cladorhizidae, Demospongiae) living in close association with Lophelia pertusa and Madrepora oculata (Scleractinia). Zootaxa 4168: 512-524

Grehan AJ, Freiwald A, Consortium A (2001) The Atlantic Coral Ecosystem Study (ACES): Forging a new partnership between scientists and principal stakeholders. In: Willison JHM, Hall J, Gass S, Kenchington ELR, Butler M, Doherty P (eds) Proceedings of the 1st International Symposium on Deep-Sea Corals. Ecology Action Centre and Nova Scotia Museum, pp 95-105

Guinotte JM, Orr J, Cairns S, Freiwald A, Morgan L, George R (2006) Will human-induced changes in seawater chemistry alter the distribution of deep-sea scleractinian corals? Frontiers in the Ecology and the Environment 4: 141-146

Hanz U, Wienberg C, Hebbeln D, Duineveld G, Lavaleye M, Juva K, Dullo WC, Freiwald A, Tamborrino L, Reichart GJ, Flögel S, Mienis F (2019) Environmental factors influencing cold-water coral ecosystems in the oxygen minimum zones on the Angolan and Namibian margins. Biogeosciences 16: 4337-4356

Hebbeln D, Wienberg C, Dullo WC, Freiwald A, Mienis F, Orejas C, Titschack J (Online) Cold-water coral reefs thriving under hypoxia. Coral Reefs

Hebbeln D, Wienberg C, Wintersteller P, Freiwald A, Becker M, Beuck L, Dullo C, Eberli GP, Glogowski S, Matos L, Forster N, Reyes-Bonilla H, Taviani M (2014) Environmental forcing of the Campeche cold-water coral province, southern Gulf of Mexico. Biogeosciences 11: 1799-1815

Heindel K, Titschack J, Dorschel B, Huvenne VAI, Freiwald A (2010) The sediment composition and predictive mapping of facies on the Propeller Mound – A cold-water coral mound (Porcupine Seabight, NE Atlantic). Continental Shelf Research 30: 1814-1829

Hoffman L, Beuck L, Van Heugten B, Lavaleye M, Freiwald A (2019) Last snails standing since the Pleistocene, a tale of Calliostomatidae (Gastropoda) living in deep-water coral habitats in the Northeast Atlantic. Zootaxa 4613: 93-110

Hoffman L, Fraussen K, Freiwald A (2018) The genus Chauvetia (Gastropoda, Buccinoidea) in deep-water coral habitats on the continental slope of Mauritania. Miscellanea Malacologica 7: 127-138

Hoffman L, Freiwald A (2017) A unique and diverse amalgamated mollusk shell assemblage from the Coral Patch Seamount, eastern Atlantic. Miscellanea Malacologica 7: 55-73

Hoffman L, Freiwald A (2018) Last snails standing: a tale of Fissurellidae (Gastropoda) from deep-water coral habitats off Mauritania since the Pleistocene. Miscellanea Malacologica 7: 115-126

Hoffman L, Freiwald A (2018) A new genus and two new species in Calliotropidae (Mollusca: Gastropoda) from NE Atlantic cold-water coral habitats. Miscellanea Malacologica 7: 89-96

Hoffman L, Freiwald A (2019) Neocardia kandeli n. sp. (Bivalvia: Philobryidae) from Namibia. Miscellanea Malacologica 8: 55-58

Hoffman L, Freiwald A (2019) Vexillum monscorallum n. sp. (Gastropoda: Costellariidae) from deep-water coral habitats off Mauritania. Miscellanea Malacologica 8: 37-40

Hoffman L, Freiwald A, van Heugten B, Lavaleye M (2018) New information on Ganesa, Lopheliella and Trochaclis (Gastropoda) in cold-water coral habitats in the temperate and tropical North Atlantic  Miscellanea Malacologica 8: 27-34

Hoffman L, Gofas S, Freiwald A (2020) New and little-known Seguenziidae (Vetigastropoda, Gastropoda) from the South Azorean Seamount Chain. Iberus 38: 1-18

Hoffman L, Gofas S, Freiwald A (2020) Ten new species in Papuliscala de Boury, 1911 (Gastropoda, Epitoniidae) from the South Azorean Seamount Chain. Iberus 38: 29-53

Hoffman L, Lavaleye M, Van Heugten B, Freiwald A (2018) Lepetella (Gastropoda: Lepetellidae) in deep-water coral habitats in the North Atlantic. Miscellanea Malacologica 8: 19-26

Hoffman L, Monsecour K, Freiwald A (2019) Columbellidae (Gastropoda) from deep-water coral habitats off Mauritania. Miscellanea Malacologica 8: 41-49

Hoffman L, Von Cosel R, Freiwald A (2019) New and little-known Pliocardiinae (Bivalvia, Vesicomyidae) from the continental slope off Mauritania. Basteria 83: 151-157

Joseph N, López Correa M, Schönfeld J, Rüggeberg A, Freiwald A (2013) Subarctic Holocene climatic and oceanographic variability in Stjernsund, northern Norway: evidence from benthic foraminifera and stable isotopes. Boreas 42: 511-531

Kaszemeik K, Freiwald A (2003) Lophelia pertusa (Scleractinia) — from skeletal structures to growth patterns and morphotypes. Institute for Paleontology, Erlangen, pp 1-26

Kiriakoulakis K, Fisher E, Wolff GA, Freiwald A, Grehan A, Roberts JM (2005) Lipids and nitrogen isotopes of two deep-water corals from the North-East Atlantic: initial results and implications for their nutrition. In: Freiwald A, Roberts JM (eds) Cold-water corals and ecosystems. Springer, Heidelberg, pp 715-729

Kiriakoulakis K, Freiwald A, Fisher E, Wolff G (2007) Organic matter quality and supply to deep-water coral/mound systems of the NW European continental margin. International Journal of Earth Sciences 96: 159-170

Könnecker G, Freiwald A (2005) Plectroninia celtica n.sp. (Calcarea, Minchinellidae), a new species of “pharetronid” sponge from bathyal depths in the northern Porcupine Seabight, NE Atlantic. Facies 51: 53-59

López Correa M, Freiwald A, Hall-Spencer J, Taviani M (2005) Distribution and habitats of Acesta excavata (Bivalvia: Limidae) with new data on its shell ultrastructure. In: Freiwald A, Roberts JM (eds) Cold-water corals and ecosystems. Springer, Heidelberg, pp 173-205

López Correa M, Montagna P, Joseph N, Rüggeberg A, Fietzke J, Flögel S, Dorschel B, Goldstein SL, Wheeler A, Freiwald A (2012) Preboreal onset of cold-water coral growth beyond the Arctic Circle revealed by coupled radiocarbon and U-series dating and neodymium isotopes. Quaternary Science Reviews 34: 24-43

Macpherson E, Beuck L, Freiwald A (2016) Some species of Munidopsis from the Gulf of Mexico, Florida Straits and Caribbean Sea (Decapoda: Munidopsidae), with the description of two new species. Zootaxa 4137: 405-416

Marali S, Wisshak M, López Correa M, Freiwald A (2013) Skeletal microstructure and stable isotope signature of three bathyal solitary cold-water corals from the Azores. Palaeogeography, Palaeoclimatology, Palaeoecology 373: 25-38

Matsuyama K, Janssen A, Martínez Arbizu P, Martha SO, Freiwald A (2014) Bryozoans from RV Sonne deep-sea cruises SO 167 ‘Louisville’ and SO 205 ‘Mangan’. Zootaxa 3856: 100-116

Matsuyama K, Titschack J, Baum D, Freiwald A (2015) Two new species of erect Bryozoa (Gymnolaemata: Cheilostomata) and the application of non-destructive imaging methods for quantitative taxonomy. Zootaxa 4020: 81-100

McCulloch M, Trotter J, Montagna P, Falter J, Dunbar R, Freiwald A, Försterra G, López Correa M, Maier C, Rüggeberg A, Taviani M (2012) Resilience of cold-water scleractinian corals to ocean acidification: Boron isotopic systematics of pH and saturation state up-regulation. Geochimica et Cosmochimica Acta 87: 21-34

Mienert J, Weaver PPE, Berné S, Dullo WC, Evans D, Freiwald A, Henriet JP, Joergensen BB, Lericolas G, Lykousis V, Parkes J, Trincardi F, Westbrook G (2004) Dynamics and evolution of European margins. Oceanography 17: 12-29

Montagna P, López Correa M, Rüggeberg A, McCulloch M, Rodolfo-Metalpa R, Dullo WC, Ferrier-Pagès C, Freiwald A, Henderson GM, Mazzoli C, Russo S, Silenzi S, Taviani M (2008) Coral Li/Ca in micro-structural domains as a temperature proxy. Geochimica et Cosmochimica Acta 72: A645

Montagna P, McCulloch M, Douville E, López Correa M, Trotter J, Rodolfo-Metalpa R, Dissard D, Ferrier-Pagès C, Frank N, Freiwald A, Goldstein S, Mazzoli C, Reynaud S, Rüggeberg A, Russo S, Taviani M (2014) Li/Mg systematics in scleractinian corals: Calibration of the thermometer. Geochimica et Cosmochimica Acta 132: 288-310

Morato T, González-Irusta JM, Dominguez-Carrió C, Wei CL, Davies A, Sweetman AK, Taranto GH, Beazley L, García-Alegre A, Grehan A, Laffargue P, Murillo FJ, Sacau M, Vaz S, Kenchington E, Arnaud-Haond S, Callery O, Chimienti G, Cordes E, Egilsdottir H, Freiwald A, Gasbarro R, Gutiérrez-Zárate C, Gianni M, Gilkinson K, Wareham Hayes VE, Hebbeln D, Hedges K, Henry LA, Johnson D, Koen-Alonso M, Lirette C, Mastrototaro F, Menot L, Molodtsova T, Muñoz PD, Orejas C, Pennino MG, Puerta P, Ragnarsson SA, Ramiro-Sánchez B, Rice J, Rivera J, Roberts JM, Ross SW, Rueda JL, Sampaio Í, Snelgrove P, Stirling D, Treble MA, Urra J, Vad J, Van Oevelen D, Watling L, Walkusz W, Wienberg C, Woillez M, Levin LA, Carreiro-Silva M (2020) Climate-induced changes in the suitable habitat of cold-water corals and commercially important deep-sea fishes in the North Atlantic. Global Change Biology 26: 2181-2202

Noé S, Lembke-Jene L, Reveillaud J, Freiwald A (2007) Microstructure, growth banding and age determination of a primnoid gorgonian skeleton (Octocorallia) from the late Younger Dryas to earliest Holocene of the Bay of Biscay. Facies 53: 177-188

Noé S, Titschack J, Freiwald A, Dullo WC (2006) From sediment to rock: diagenetic processes of hardground formation in deep-water carbonate mounds of the NE Atlantic. Facies 52: 183-208

Raddatz J, Liebetrau V, Rüggeberg A, Hathorne E, Krabbenhöft A, Eisenhauer A, Böhm F, Vollstaedt H, Fietzke J, López Correa M, Freiwald A, Dullo WC (2013) Stable Sr-isotope, Sr/Ca, Mg/Ca, Li/Ca and Mg/Li ratios in the scleractinian cold-water coral Lophelia pertusa. Chemical Geology 352: 143-152

Raddatz J, Titschack J, Frank N, Freiwald A, Conforti A, Osborne A, Skornitzke S, Stiller W, Rüggeberg A, Voigt S, Albuquerque ALS, Vertino A, Schröder-Ritzrau A, Bahr A (2020) Solenosmilia variabilis-bearing cold-water coral mounds off Brazil. Coral Reefs 39: 69-83

Reveillaud J, Freiwald A, Van Rooij D, Le Guilloux E, Altuna A, Foubert A, Vanreusel A, Olu-Le Roy K, Henriet JP (2008) The distribution of scleractinian corals in the Bay of Biscay, NE Atlantic. Facies 54: 317-331

Roberts JM, Freiwald A (2005) Integrated European research into cold-water coral reefs. Current 21: 41-45

Roberts JM, Wheeler AJ, Freiwald A (2006) Reefs of the deep: the biology and geology of cold-water coral ecosystems. Science 312: 543-547

Roberts JM, Wheeler AJ, Freiwald A, Cairns SD (2009) Cold-water corals – The biology and geology of deep-sea coral habitats. Cambridge University Press

Rosso A, Beuck L, Vertino A, Sanfilippo A, Freiwald A (2018) Cribrilinids (Bryozoa, Cheilostomata) associated with deep-water coral habitats at the Great Bahama Bank slope (NW Atlantic), with description of new taxa. Zootaxa 4524: 401-439

Rüggeberg A, Fietzke J, Liebetrau V, Eisenhauer A, Dullo WC, Freiwald A (2008) Stable strontium isotopes (d88/86Sr) in cold-water corals— a new proxy for reconstruction of intermediate ocean water temperatures. Earth and Planetary Science Letters 269: 569-574

Rüggeberg A, Flögel S, Dullo WC, Hissmann K, Freiwald A (2011) Water mass characteristics and sill dynamics in a subpolar cold-water coral reef setting at Stjernsund, northern Norway. Marine Geology 282: 5-12

Sakai K, Türkay M, Beuck L, Freiwald A (2015) A collection of the Infraorder Callianassidea (Decapoda, Pleocyemata) with one new genus and five new species from the Eastern Atlantic off Mauritania (R/V Maria S. Merian cruise MSM 16/3 “PHAETON”). Marine Biodiversity 45: 113-133

Sampaio I, Carreiro-Silva M, Freiwald A, Menezes G, Grasshoff M (2019) Natural history collections as a basis for sound biodiversity assessments: Plexauridae (Octocorallia, Holaxonia) of the Naturalis CANCAP and Tyro Mauritania II expeditions. ZooKeys 870: 1-32

Sampaio I, Freiwald A, Porteiro FM, Menezes G, Carreiro-Silva M (2019) Census of Octocorallia (Cnidaria: Anthozoa) of the Azores (NE Atlantic) with a nomenclature update. Zootaxa 4550: 451-498

Sanfilippo R, Vertino A, Rosso A, Beuck L, Freiwald A, Taviani M (2013) Serpula aggregates and their role in deep-sea coral communities in the southern Adriatic Sea. Facies 59: 663-677

Savini A, Vertino A, Marchese F, Beuck L, Freiwald A (2014) Mapping cold-water coral habitats at different scales within the northern Ionian Sea (Central Mediterranean): An assessment of coral coverage and associated vulnerability. Plos One 9: e87108

Schleinkofer N, Raddatz J, Freiwald A, Evans D, Beuck L, Rüggeberg A, Liebetrau V (2019) Environmental and biological controls on Na/Ca ratios in scleractinian cold-water corals. Biogeosciences 16: 3565-3582

Schönfeld J, Dullo WC, Pfannkuche O, Freiwald A, Rüggeberg A, Schmidt S, Weston J (2011) Recent benthic foraminiferal assemblages from cold-water coral mounds in the Porcupine Seabight. Facies 57: 187-213

Schröder-Ritzrau A, Freiwald A, Mangini A (2005) U/Th-dating of deep-water corals from the eastern North Atlantic and the western Mediterranean Sea. In: Freiwald A, Roberts JM (eds) Cold-water corals and ecosystems. Springer, Heidelberg, pp 157-172

Sorauf JE, Freiwald A (2002) Skeletal structure in the rugosan genus Calophyllum from Permian strata of Timor: Comparison with the living deep water coral Lophelia pertusa. Coral Research Bulletin 7: 191-207

Tamborrino L, Wienberg C, Titschack J, Wintersteller P, Mienis F, Schröder-Ritzrau A, Freiwald A, Orejas C, Dullo WC, Haberkern J, Hebbeln D (2019) Mid-Holocene extinction of cold-water corals on the Namibian shelf steered by the Benguela oxygen minimum zone. Geology 47: 1185-1188

Taviani M, Angeletti L, Beuck L, Campiani E, Canese S, Foglini F, Freiwald A, Montagna P, Trincardi F (2015) On and off the beaten track: Megafaunal sessile life and Adriatic cascading processes. Marine Geology 369: 273-287

Taviani M, Angeletti L, Dimech M, Mifsud C, Freiwald A, Harasewych MG, Oliverio M (2009) Coralliophilinae (Gastropoda: Muricidae) associated with deep-water coral banks in the Mediterranean. The Nautilus 123: 106-112

Taviani M, Freiwald A, Beuck L, Angeletti L, Remia A, Vertino A, Dimech M, Schembri PJ (2010) The deepest known occurrence of the precious red coral Corallium rubrum (L., 1758) in the Mediterranean Sea. In: Bussoletti E, Cottingham D, Bruckner A, Roberts G, Sandulli R (eds) Proceedings of the International Workshop on Red Coral Science, Management, and Trade: Lessons from the Mediterranean. NOAA Technical Memorandum CRCP-13, pp 87-93

Taviani M, Freiwald A, Zibrowius H (2005) Deep coral growth in the Mediterranean Sea: an overview. In: Freiwald A, Roberts JM (eds) Cold-water corals and ecosystems. Springer, Heidelberg, pp 137-156

Taviani M, Remia A, Corselli C, Freiwald A, Malinverno E, Mastrototaro F, Savini A, Tursi A (2005) First geo-marine survey of living cold-water Lophelia reefs in the Ionian Sea (Mediterranean basin). Facies 50: 409-417

Titschack J, Baum D, De Pol-Holz R, López Correa M, Forster N, Flögel S, Hebbeln D, Freiwald A (2015) Aggradation and carbonate accumulation of Holocene Norwegian cold-water coral reefs. Sedimentology 62: 1873-1898

Titschack J, Bromley RG, Freiwald A (2005) Plio-Pleistocene cliff-bound, wedge-shaped, warm-temperate carbonate deposits from Rhodes (Greece): Sedimentology and facies. Sedimentary Geology 180: 29-56

Titschack J, Fink HG, Baum D, Wienberg C, Hebbeln D, Freiwald A (2016) Mediterranean cold-water corals – an important regional carbonate factory? Depositional Record 2: 74-96

Titschack J, Freiwald A (2005) Growth, deposition, and facies of Pleistocene bathyal coral communities from Rhodes, Greece. In: Freiwald A, Roberts JM (eds) Cold-water corals and ecosystems. Springer, Heidelberg, pp 41-59

Titschack J, Thierens M, Dorschel B, Schulbert C, Freiwald A, Kano A, Takashima C, Kawagoe N, Li X (2009) Carbonate budget of a cold-water coral mound (Challenger Mound, IODP Exp. 307). Marine Geology 259: 36-46

Weaver PPE, Billett DSM, Boetius A, Danovaro R, Freiwald A, Sibuet M (2004) Hotspot ecosystem research on Europe’s deep-ocean margins. Oceanography 17: 132-143

Weaver PPE, Boetius A, Danovaro R, Freiwald A, Gunn V, Heussner S, Morato T, Schewe I, van den Howe S (2009) The future of integrated deep-sea research in Europe—The HERMIONE Project. Oceanography 22: 177-191

Welte C, Wacker L, Hattendorf B, Christl M, Fohlmeister J, Breitenbach SFM, Robinson LF, Andrews AH, Freiwald A, Farmer JR, Yeman C, Synal HA, Günther D (2016) Laser ablation − accelerator mass spectrometry: An approach for rapid radiocarbon analyses of carbonate archives at high spatial resolution. Analytical Chemistry 88: 8570-8576

Wheeler AJ, Beyer A, Freiwald A, de Haas H, Huvenne VAI, Kozachenko M, Olu-Le Roy K, Opderbecke J (2007) Morphology and environment of cold-water coral carbonate mounds on the NE European margin. International Journal of Earth Sciences 96: 37-57

Wienberg C, Beuck L, Heidkamp S, Hebbeln D, Freiwald A, Pfannkuche O, Monteys X (2008) Franken Mound: facies and biocoenoses on a newly-discovered “carbonate mound” on the western Rockall Bank, NE Atlantic. Facies 54: 1-24

Wienberg C, Hebbeln D, Fink HG, Mienis F, Dorschel B, Vertino A, López Correa M, Freiwald A (2009) Scleractinian cold-water corals in the Gulf of Cádiz—First clues about their spatial and temporal distribution. Deep-Sea Research I 56: 1873-1893

Wienberg C, Titschack J, Freiwald A, Frank N, Lundälv T, Taviani M, Beuck L, Schröder-Ritzrau A, Krengel T, Hebbeln D (2018) The giant Mauritanian cold-water coral mound province: Oxygen control on coral mound formation. Quaternary Science Reviews 185: 135-152

Wisshak M, Freiwald A, Lundälv T, Gektidis M (2005) The physical niche of the bathyal Lophelia pertusa in a non-bathyal setting: environmental controls and palaeoecological implications. In: Freiwald A, Roberts JM (eds) Cold-water corals and ecosystems. Springer, Heidelberg, pp 979-1001

Wisshak M, López Correa M, Gofas S, Salas C, Taviani M, Jakobsen J, Freiwald A (2009) Shell architecture, element composition, and stable isotope signature of the giant deep-sea oyster Neopycnodonte zibrowii sp. n. from the NE Atlantic. Deep-Sea Research I 66: 374-407

Wisshak M, López Correa M, Zibrowius H, Jakobsen J, Freiwald A (2009) Skeletal reorganisation affects geochemical signals, exemplified in the stylasterid hydrocoral Errina dabneyi (Azores Archipelago). Marine Ecology Progress Series 397: 197-208

Wisshak M, Neumann C, Jakobsen J, Freiwald A (2009) The ‘living-fossil community’ of the cyrtocrinoid Cyathidium foresti and the deep-sea oyster Neopycnodonte zibrowii (Azores Archipelago). Palaeogeography, Palaeoclimatology, Palaeoecology 271: 77-83

Zettler ML, Freiwald A, Guerra-García JM (2018) Cold-water corals off Angola as refuge for a new Aeginella species (Crustacea: Amphipoda: Caprellidae). Zootaxa 4462: 535-546


Populärwissenschaftliche Publikationen zum Thema Kaltwasserkorallen 

Freiwald A (2003) Korallengärten in kalten Tiefen. Spektrum der Wissenschaft 2/2003: 666-673

Freiwald A (2004) Kaltwasser-Karbonate und Tiefwasser-Riffe – eine geobiologische Herausforderung. Gmit 18: 8-14

Freiwald A (2014) Kaltwasserkorallen – aus der Grundlagenforschung auf die politische Agenda. Natur, Forschung, Museum 144: 6-17

Freiwald A (2015) Kaltwasserriffe als Geobiodiversitäts-Hotspots. Natur, Forschung und Museum 145: 6-13

Freiwald A (2015) Tiefseeforschungsschiff „Sonne“ nimmt Fahrt auf. Natur, Forschung und Museum 145: 34-37

Freiwald A, Beuck L (2011) Sind Kaltwasserkorallen durch den Klimawandel gefährdet? In: Lozán JL, Graßl H, Karbe L, Reise K (eds) Warnsignal Klima: Die Meere – Änderungen und Risiken, Hamburg, pp 220-224

Freiwald A, Beuck L, Wisshak M (2013) Korallenriffe im kalten Wasser des Nordatlantiks. In: Beck E (ed) Die Vielfalt des Lebens: Wie hoch, wie komplex, warum? Wiley-VCH, pp 88-98

George KH, Wehrmann A, Freiwald A (2019) Gipfelstürmer der Tiefe. Natur, Forschung und Museum 149: 72-77

Hebbeln D, Freiwald A (2010) Manche mögen’s kalt – Der europäische Korallengürtel von Norwegen bis ins Mittelmeer. In: Wefer G, Schmieder F (eds) Expedition Erde. Marum Bibliothek verständliche Geowissenschaften, Bremen, pp 394-401

Schmidt H, Freiwald A (1993) Rezente gesteinsbohrende Kleinorganismen des norwegischen Schelfs. Natur und Museum 123: 149-155


Heutige nicht-tropische biosedimentäre Systeme (Kaltwasserkarbonate im Flachwasser)

Andruleit H, Freiwald A, Schäfer P (1996) Bioclastic carbonate sediments on the southwestern Svalbard Shelf. Marine Geology 134: 163-182

Färber C, Titschack J, Schönberg CHL, Ehrig K, Boos K, Baum D, Illerhaus B, Asgaard U, Bromley RG, Freiwald A, Wisshak M (2016) Long-term macrobioerosion in the Mediterranean Sea assessed by micro-computed tomography. Biogeosciences 13: 3461-3474

Färber C, Wisshak M, Pyko I, Bellou N, Freiwald A (2015) Effects of water depth, seasonal exposure, and substrate orientation on microbial bioerosion in the Ionian Sea (Eastern Mediterranean). Plos One 10: e0126495

Freiwald A (1993) Coralline algal maerl frameworks-Islands within the phaeophytic kelp belt. Facies 29: 133-148

Freiwald A (1995) Bacteria-induced carbonate degradation: a taphonomic case study of Cibicides lobatulus from a high-boreal carbonate setting. Palaios 10: 337-346

Freiwald A (1995) Sedimentological and biological aspects in the formation of branched rhodoliths in northern Norway. Beiträge zur Paläontologie 20: 7-19

Freiwald A (1998) Microbial maceration and carbonate dissolution on cold-temperate shelves. Historical Biology 13: 27-35

Freiwald A (1998) Modern nearshore cold-temperate calcareous sediments in the Troms District, northern Norway. Journal of Sedimentary Research 68: 763-776

Freiwald A, Henrich R (1994) Reefal coralline algal build-ups within the Arctic Circle: morphology and sedimentary dynamics under extreme environmental seasonality. Sedimentology 41: 963-984

Freiwald A, Henrich R, Schäfer P, Willkomm H (1991) The significance of high-boreal to subarctic maerl deposits in northern Norway to reconstruct Holocene climatic changes and sea level oscillations. Facies 25: 315-340

Freiwald A, Mostafawi N (1998) Ostracods in a cold-temperate coastal environment, western Troms, northern Norway: sedimentary aspects and assemblages. Facies 38: 255-274

Henrich R, Freiwald A, Betzler CB, Bader B, Schäfer P, Samtleben C, Brachert TC, Wehrmann A, Zankl H, Kühlmann DHH (1995) Controls on modern carbonate sedimentation on warm-temperate to Arctic coasts, shelves and seamounts in the northern Hemisphere: implications for fossil counterparts. Facies 32: 71-108

Henrich R, Freiwald A, Bickert T, Schäfer P (1997) Evolution of an Arctic open-shelf carbonate platform, Spitsbergen Bank (Barents Sea). In: James NP, Clarke JAD (eds) Cool-water carbonates. SEPM, Special Volume, pp 163-181

Henrich R, Freiwald A, Wehrmann A, Schäfer P, Samtleben C, Zankl H (1996) Nordic cold-water carbonates: occurrences and controls. Göttinger Arbeiten zur Geologie und Paläontologie Sb 2: 35-52

Henrich R, Hartmann M, Reitner J, Schäfer P, Freiwald A, Steinmetz S, Dietrich P, Thiede J (1992) Facies belts and communities of the Arctic Vesterisbanken Seamount (Central Greenland Sea). Facies 27: 71-104

Teichert S, Freiwald A (2014) Polar coralline algal CaCO3-production rates correspond to intensity and duration of the solar radiation. Biogeosciences 11: 833-842

Teichert S, Woelkerling W, Rüggeberg A, Wisshak M, Piepenburg D, Meyerhöfer M, Form A, Büdenbender J, Freiwald A (2012) Rhodolith beds (Corallinales, Rhodophyta) and their physical and biological environment at 80°31’N in Nordkappbukta (Nordaustlandet, Svalbard Archipelago, Norway). Phycologia 51: 371-390

Teichert S, Woelkerling W, Rüggeberg A, Wisshak M, Piepenburg D, Meyerhöfer M, Form A, Freiwald A (2014) Arctic rhodolith beds and their environmental controls (Spitsbergen, Norway). Facies 60: 15-37

Wehrmann A, Freiwald A, Zankl H (1995) Formation of cold-temperate water multispecies rhodoliths in intertidal gravel pools from northern Brittany, France. Senckenbergiana maritima 26: 51-71

Westphal H, Halfar J, Freiwald A (2010) Heterozoan carbonates in subtropical to tropical settings in the present and in the past. International Journal of Earth Sciences 99 Supplement 1: S153-S169

Wisshak M, Berning B, Jakobsen J, Freiwald A (2015) Temperate carbonate production: biodiversity of calcareous epiliths from intertidal to bathyal depths (Azores). Marine Biodiversity 45: 87-112

Wisshak M, Form A, Jakobsen J, Freiwald A (2010) Temperate carbonate cycling and water mass properties from intertidal to bathyal depths (Azores, N-Atlantic). Biogeosciences 7: 2379-2396

Wisshak M, Gektidis M, Freiwald A, Lundälv T (2005) Bioerosion along a bathymetric gradient in a cold-temperate setting (Kosterfjord, SW Sweden): an experimental study. Facies 51: 93-117

Wisshak M, Schönberg CHL, Form A, Freiwald A (2012) Ocean acidification accelerates reef bioerosion. Plos One 7: e45124

Wisshak M, Schönberg CHL, Form A, Freiwald A (2013) Effects of ocean acidification and global warming on reef bioerosion—lessons from a clionaid sponge. Aquatic Biology 19: 111-127

Wisshak M, Schönberg CHL, Form A, Freiwald A (2014) Sponge bioerosion accelerated by ocean acidification across species and latitudes? Helgoland Marine Research 68: 253-262

Wisshak M, Tribollet A, Golubic S, Jakobsen J, Freiwald A (2011) Temperate bioerosion: ichnodiversity and biodiversity from intertidal to bathyal depths (Azores). Geobiology 9: 492-520


Geologische und paläontologische Studien

Beuck L, Wisshak M, Munnecke A, Freiwald A (2008) A giant boring in a Silurian stromatoporoid analysed by computer tomography. Acta Palaeontologica Polonica 53: 149-160

Birgel D, Himmler T, Freiwald A, Peckmann J (2008) A new constraint on the antiquity of anaerobic oxidation of methane: Late Pennsylvanian seep limestones from southern Namibia. Geology 36: 543-546

Blomeier D, Wisshak M, Dallmann W, Volohonsky E, Freiwald A (2003) Facies analysis of the Old Red sandstone of Spitsbergen (Wood Bay Formation): Reconstruction of the depositional environments and implications of basin development. Facies 49: 151-174

Blomeier D, Wisshak M, Joachimski M, Freiwald A, Volohonsky E (2003) Calcareous, alluvial and lacustrine deposits in the Old Red Sandstone of central north Spitsbergen (Wood Bay Formation, Early Devonian). Norwegian Journal of Geology 83: 281-298

Freiwald A, Krzeminski W (1991) Cylindrotomidae (Diptera, Tipulomorpha) from the Paleogene of Bolshaya Svetlovodnaya (Eastern Asiatic USSR). Paläontologische Zeitschrift 65: 339-344

Himmler T, Freiwald A, Stollhofen H, Peckmann J (2008) Late Carboniferous hydrocarbon-seep carbonates from the glaciomarine Dwyka Group, southern Namibia. Palaeogeography, Palaeoclimatology, Palaeoecology 257: 185-197

Krzeminski W, Freiwald A (1991) Toxorhina (Ceratocheilus) caucasiensis, a new species from the Middle Miocene of Stavropol (northern Caucasus, USSR) (Diptera, Limoniidae). Paläontologische Zeitschrift 65: 153-155

Kuechler RR, Birgel D, Kiel S, Freiwald A, Goedert J, Thiel V, Peckmann J (2012) Miocene methane-derived carbonates from southwestern Washington, USA and a model for silification at seeps. Lethaia 45: 259-273

Milker Y, Weinkauf MFG, Titschack J, Freiwald A, Krüger S, Jorissen FJ, Schmiedl G (2017) Testing the applicability of a benthic foraminiferal-based transfer function for the reconstruction of paleowater depth changes in Rhodes (Greece) during the early Pleistocene. PlosOne 12: e0188447

Nelson CS, Freiwald A, Titschack J, List S (2001) Lithostratigraphy and sequence architecture of temperate mixed siliciclastic-carbonate facies in a new Plio-Pleistocene section at Plimiri, Rhodes Island (Greece). Occasional Report Department of Earth Sciences, University of Waikato 25: 1-50

Titschack J, Joseph N, Fietzke J, Freiwald A, Bromley RG (2013) Record of a tectonically-controlled regression captured by changes in carbonate skeletal associations on a structured island shelf (mid-Pleistocene, Rhodes, Greece). Sedimentary Geology 283: 15-33

Titschack J, Nelson CS, Beck T, Freiwald A (2008) Sedimentary evolution of a Late Pleistocene temperate red algal reef (Coralligène) on Rhodes, Greece: correlation with global sea-level fluctuations. Sedimentology 55: 1747-1776

Titschack J, Radtke U, Freiwald A (2009) Dating and characterization of polymorphic transformation of aragonite to calcite in Pleistocene bivalves from Rhodes (Greece) by combined shell microstructure, stable isotope, and electron spin resonance study. Journal of Sedimentary Research 79: 332-346

Wisshak M, Volohonsky E, Seilacher A, Freiwald A (2004) A trace fossil assemblage from fluvial Old Red deposits (Wood Bay Formation; Lower to Middle Devonian) of NW-Spitsbergen, Svalbard. Lethaia 37: 149-163






Prof. Dr. Pedro Martínez Arbizu
Head of DZMB

Library administration