Tertiary Mammals and Morphometrics


Prof. Dr. Ottmar Kullmer
Head of Division
Head of Section

Research interests

My research interests are diverse, reaching from general mammalian palaeobiology and palaeoanthropology into almost all topics of bone and dental analyses. In 2001 I have established at SF a virtual imaging lab for 3D-digitalization, editing, reconstruction and morphometrics of vertebrate hard tissue. The focus of my research lays on the structure and function of teeth with a main attention on evolutionary development, dietary and cultural adaptations in the masticatory system. A number of my national and international research projects funded by the DFG (e.g. DFG-FOR771) and the EU (EU-MRTN-CT-2005-019564) have led to the development of the well-known Occlusal Fingerprint Analysis (OFA).

Educational background

2022 Appointment as adjunct Professor
2014 Habilitation, Venia legendi for Zoology (Privatdozent), Faculty of Biosciences, Goethe University Frankfurt a. M., Germany
1997 Dr. rer. nat. graduation in Paleontology and Geology, Faculty of Geosciences, JG University of Mainz, Germany
1991 Diploma in Geology, Paleontology, Zoology and Anthropology, JG University of Mainz, Germany
2014 Habilitation, Venia legendi for Zoology (Privatdozent), Faculty of Biosciences, Goethe University Frankfurt a. M., Germany
Positions held

2016  – Head Department of Paleoanthropology, Senckenberg Research Institute and Natural History Museum Frankfurt, Germany
2001  – Head Tertiary Mammal and Morphometrics Section, Senckenberg Research Institute, Department of Palaeoanthropology, Frankfurt am Main, Germany
2008 – 2014 Provisional Head of Mammalogy Section, Senckenberg Research Institute and Natural History Museum, Department of Messel Research and Mammalogy Frankfurt, Germany
2003, 2004  Guest-professorship, Institute for Anthropology, University of Vienna, Austria.
1994 – 1997  Research Fellow University of Mainz, DFG-Project: Evolution of African Plio-Pleistocene suids based on tooth pattern analysis
1998 Research Assistant Hesse State Museum Darmstadt
1993 – 1994  Research Assistant Hesse State Museum Darmstadt
1999 – 2001 Research Fellow University of Mainz, Hesse State Museum Darmstadt, DFG-Project: Digital Breakage Pattern Analysis
Research profile

  • Palaeoanthropology
  • Virtual Anthropology
  • Hominid Palaeoecology and Palaeobiogeography
  • Life History Reconstruction
  • Palaeodiet
  • Biomechanics
  • Hard Tissue Functional Morphology
  • Palaeobiology of Mammals
  • Bone and tooth morphometry

Field experience

1988 Maramena (Northern Greece): Upper Miocene.
1992 – 1997 Chiwondo Beds (Malawi) ” Hominid Corridor Research Project”, Plio-Pleistocene.
1998 – 1999 Laetoli (Ndolanya Beds) and Kakesio (Kakesio Beds) in Tanzania, Plio-Pleistocene.
2000 – 2009 Gadamaitu (Somali Region) in Ethiopia: Plio-Pleistocene.
2006 Albertine Rift (Nkondo, Kaiso) Uganda: Mio-Peistocene
2008 –  Chiwondo Beds (Malawi) ” Hominid Corridor Research Project”, Plio-Pleistocene.
Third party funded projects (last 10 years)

2019 – 2020 The impact of climate change on growth and development in marsupial and placental mammals. Contribution: Chief Investigator. Cooperation with Dr. L. Fiorenza, Monash University, Australia. Faculty of Science and Faculty of Medicine Interdisciplinary Research Seed Funding Scheme (S01802), Monash University, Australia.                                                        18.000 €
2017 Monash University, Faculty of Medicine Strategic Grant Scheme (SGS16-0344). Understanding the evolution of the human diet. Cooperation with Dr. L. Fiorenza Monash University, Australia.                                                                                               30.000 €
2008 – 2014 DFG (German Science Foundation) reserach unit FOR 771: Occlusal Fingerprint Analyser – OFA software tool for the analysis of occlusal tooth contacts, 370.000 €
2012 – 2014 DFG (German Science Foundation) KU 1260/8 Project A2: Evolution and functional variability of bunodont molars in primates, 220.000 €
2008 – 2014 DFG (German Science Foundation) KU 1260/9 Project D3: Function of the dentition in original Tribosphenida, Co-PI together with Prof. Dr. T. Martin, University Bonn, 210.000 €
2006 – 2009 EU-MRTN-CT-2005-019564 EVAN – European Virtual Anthropology Network,Human Resources and Mobility (HRM) Activity, Marie Curie Actions Research Training Network (RTN), together with 15 European Partners                                                        EVAN total amount, 3.500.000 €
2006 – 2009 EU-MRTN-CT-2005-019564 EVAN own sub-project: Functional pattern analysis of tooth surfaces applying virtual technique, 250.000 €
Academic teaching experiences (selected)          

  • Lectures

2011 – Evolutionary biology of mammals and paleoanthropology Institute of Ecology, Evolution and Diversity, Goethe University Frankfurt, Germany
2006  – Paleoecology of early hominids     (Functional morphology of mammals), Institute of Ecology, Evolution and Diversity, Goethe University Frankfurt, Germany
2003 – 2004 Background to African hominid sites, Department of Anthropology, University Vienna, Austria
2001 – 2002 Palaeobiology of vertebrates (architecture and function), Institute of Geosciences Goethe University Frankfurt, Germany
2000 – 2005 Specific zoology (vertebrates), comparative morphology and function of the Metazoa (amphibians, reptiles, mammals), Institute of Ecology, Evolution and Diversity, Goethe University Frankfurt, Germany

  • Practical courses

2011  – Strukture and function of organisms, Institut für Ökologie, Institute of Ecology, Evolution and Diversity, Goethe University Frankfurt, Germany
2011 –     Practical course and excursion, Evolutionary biology of mammals and paleoanthropology, Institute of Ecology, Evolution and Diversity, Goethe University Frankfurt, Germany
2010 – Evolution and biodiversity of animals (Evolution of architectural and functional morphology of vertebrates Institute of Ecology, Evolution and Diversity, Goethe University Frankfurt, Germany
2006 – Paleoecology of early hominids     (Functional morphology of mammals)         ), Institute of Ecology, Evolution and Diversity, Goethe University Frankfurt, Germany
2001 –       Introduction to the scientific work, Institute of Ecology, Evolution and Diversity, Goethe University Frankfurt, Germany
2003 – 2004    Privatissimum, practical training for fossil sites remote sensing (satellite image analysis), Department of Anthropology, Univeristy Vienna, Austria
2000 – 2005    Specific zoology (vertebrates), comparative morphology and function of the Metazoa (amphibians, reptiles, mammals), Institute of Ecology, Evolution and Diversity, Goethe University Frankfurt, Germany

  •  Seminars

2011 – Seminar of evolutionary biology, Institute of Ecology, Evolution and Diversity, Goethe University Frankfurt, Germany
2007 – Seminar for exam candidates, Institute of Ecology, Evolution and Diversity, Goethe University Frankfurt, Germany
2006 –  Paleoecology of early hominids     (Functional morphology of mammals), Institute of Ecology, Evolution and Diversity, Goethe University Frankfurt, Germany
2001 – Introduction to the scientific work, Examenskandidaten, Institute of Ecology, Evolution and Diversity, Goethe University Frankfurt, Germany
2003 – 2004 Introduction in the Plio-Pleistocene faunas of Africa, Department of Anthropology, Univeristy Vienna, Austria

  • Supervision/Co-supervision of PhD thesis

2018 – Zur Bedeutung der Analyse okklusaler Abrasionsmuster für die klinisch-manuelle Funktionsdiagnostik“, dissertation J. Heil, cooperation with Prof. U. Lotzmann, Phillipps University Marburg, Germany.
2018 – Dietary Changes during the english industrial revolution and transformed masticatory behaviours. Dissertation C. Silvester, cooperation with Prof. S. Hillson University College London, UK.
2011 – 2017 Evolution und Funktionsvariabilität von bunodonten Molaren bei Primaten, disseration U. Menz, Goethe University Frankfurt, Germany
2010 – 2015  Early Homo phylogenetic and functional relationships, reconstructed from mandibular/dental structures and growth patterns, dissertation M. Bäuchle, cooperation with Prof. J.-J. Hublin, Max-Planck Institut for Evolutionary Anthropology Leipzig and Prof. F. Schrenk, Goethe University Frankfurt, Germany
2006 – 2012 Tandem scanning Microscopical Analysis of enamel ultrastructure of Homo erectus, dissertation J. Tausch, cooperation with Prof. F. Schrenk, Goethe University Frankfurt, Germany, and Prof. Dr. T. Bromage New York University (NYU), College of Dentistry
2006 – 2009 Occlusion Wear Pattern Analysis of Functional Morphology in Neanderthal and         Early Homo sapiens, dissertation Luca Fiorenza, cooperation with Prof. F.  Schrenk, Goethe University Frankfurt, Germany
2003 – 2006 Kaumechanik und Funktion der Molaren-Okklusion früher Hominiden, dissertation M.  Huck, cooperation with Prof. F. Schrenk, Goethe University Frankfurt, Germany
2002 – 2006  Molaren – Konstruktion und Funktion der Okklusionsfacetten der Cercopithecoidea, dissertation L. Ulhaas, cooperation with Prof. F. Schrenk, Goethe University Frankfurt, Germany
2001 – 2004 Molaren – Konstruktion, Phylogenese und Paläoökologie der Hominoidea (Hylobates, Pongo, Gorilla, Pan), dissertation K. Engel, cooperation with Prof. F. Schrenk, Goethe University Frankfurt, Germany

  • Supervision of Diploma, master and bachelor thesis

2001 – Diploma, master and bachelor thesis in the fields of evolution and functional morphology of mammals, including hominins, paleoecology, taphonomy, morphometry, 2D/3D image analysis, virtual paleontology, Paläoökologie, Taphonomie, Morphometrie, 2D/3D Bildverarbeitung; Virtuelle Paläontologie Institute of Ecology, Evolution and Diversity, Goethe University Frankfurt, Germany

  • Long term scientific cooperations (international und national)

2018 – Virtual analyses of functional morphology in modern human teeth. Cooperation: Prof. U. Lotzmann, department of orofacial Prosthetics and function, Phillipps University Marburg, Germany.
2016 – Isotope, element and ultra-structure analyses in dental tissue for life history reconstructions of mammals, and hominins. Cooperation:  Prof. W. Müller, Institute of Geosciences, Goethe University Frankfurt, Germany, Prof. A. Mulch, Senckenberg Biodiversity and Climate Research Center Frankfurt, Germany, Dr. L Bondioli, Museo delle Civiltà Roma, Servizio di Bioarcheologia, Roma, Italy, Prof. R. Joannes-Boyau Southern Cross University Lismore, Australia, Prof. G. Schwartz, School of Human Evolution and Social Changes, Arizona State University Tempe, USA.
2015 – Human Evolution and Biodiversity in South East Asia. Analysis of fossil remains in the G.H.R. von Koenigswald collection. Cooperation: Clément Zanolli, Laboratoire PACEA, University Bordeaux, France.
2010 – Human Paleobiomics Hard Tissue Research Program (www.paleobiomics.org), International research program for structural life history analyses of dental and bone material. Cooperation: Prof. T. Bromage, University of New York (NYU), College of Dentistry, Prof. F. Schrenk JWG – Universität Frankfurt a. M.
2010 –  Phylogenetic and functional relationships in early hominins reconstructed from mandibular/dental structures and growth patterns. Cooperation: Prof. Jean-Jacques Hublin, Prof. K. Kupczik, Dr. P. Gunz, Max-Planck Institute for Evolutionary Anthropology Leipzig, Prof. Schrenk, Institute of Ecology, Evolution and Diversity, Goethe University Frankfurt, Germany
2010 – Combining Finite Element Method (FEM) and Occlusal Fingerprint Analysis (OFA), distribution of loads in dental tissues and mandibular bone, dental arch reconstruction in fossil hominids derived from Occlusal Fingerprint Analysis Prof. S. Benazzi, Department of Cultural Heritage. University Bologna, Italy
2006 – Virtual structure analysis und 3D morphometry in anthropology, EVAN Society (www.evan-society.org). Kooperation: Prof. G. Weber, Department of Anthropology, University Vienna.
2003 – Fundamental research in correlation between the jaw joint motion and native occlusal surface in teeth. Cooperation: Prof. Olaf Winzen, SRH-University Hamm, Dentale Bewegungslehre und Craniomedizin Zentrum Frankfurt, Germany.

  • Organisation of international workshops und conference (selection)

2019  International Senckenberg Workshop “Evolutionary Medicine: The Biocultural Perspective”, Senckenberg Research Institute and Natural History Museum Frankfurt, Germany.
2016  Co-organizer International Senckenberg-Conference 125 Years of Homo erectus; Georgian National Museum, Tbilisi, Georgia
2011 Co-organizer International Senckenberg Konferenz „The World at the Messel Time: Puzzles in Palaeobiology, Palaeoenvironment and the History of the early Primates”. Senckenberg Research Institute and Natural History Museum Frankfurt, Germany.
2007 EVAN „Intensive Training Course“ Evolution und Function of Mammalian Dentition Senckenberg Research Institute and Natural History Museum Frankfurt, Germany.
2005 3D-Advances in Natural History and Cultural Heritage (Symposium with 80 international participants) Senckenberg Research Institute and Natural History Museum Frankfurt, Germany.
Museum activities

  • Exhibits

2010  Scientific supervision „Safari zum Urmensch“, temporary exhibition Senckenberg Research Institute and Natural History Museum Frankfurt, Germany.
2008 – 2010 Idea, concept and organisation „Humans in 3D – In the focus digital techniques, travelling exhibition in the frame of the EU-project EVAN. presentation: Frankfurt, Germany, Vienna Austria, Leipzig, Germany, Mettmann, Germany, Toulouse, France, Meersburg, Germany.
1998 Concept and co-organisation permanent exhibition ”Die Frühzeit des Menschen”, State Museum of Hessia Darmstadt, Germany.                                  
1998 Scientific supervision of eight head-reconstructions of early humans, cooperation with   Wildlife Art (W. Schnaubelt & N. Kieser), State Museum of Hessia Darmstadt, Germany.

  • Curator activities

2001 – Collections of fossil mammal remains dinothere sands of Eppelsheim 1996-2000 and Dorn Dürkheim hosted in the Senckenberg Research Institute and Natural History Museum Frankfurt, Germany.
2001 –  Collection G.H.R. von Koenigswald fossil teeth and bone from Java, Indonesia, Senckenberg Research Institute and Natural History Museum Frankfurt, Germany.
2001 – Cast collection fossil of hominids, Senckenberg Research Institute and Natural History Museum Frankfurt, Germany.
2001 – Collection digital surface and CT data of fossil hominoid teeth, jaws, skulls, Senckenberg Research Institute and Natural History Museum Frankfurt, Germany.
2008-2014 Provisional curator collection of modern mammals, Senckenberg Research Institute and Natural History Museum Frankfurt, Germany.

  • Memberships, commissions and editorial activities

2011 – European Society of Human Evolution (ESHE)
2010 – 2012 Associate Editor Journal of Human Evolution
2009-2017 Scientific BoardWissenschaftsausschuss Senckenberg Gesellschaft für Naturforschung
2009 – EVAN Society e.V. European Virtual Anthropology Network
2006 – Society of Vertebrate Paleontology (SVP)
2006 – IT-user advisory council Senckenberg Gesellschaft für Naturforschung
2005 – Scientific editorial advisory council Senckenberg Journal Nature – Research –Museum
2005 – German Palaeontological Society

Selected publications (peer-reviewed, since 2011)


Fiorenza L, Menter C G, Fung S, Lee J, Kaidonis J,  Moggi-Cecchi J, Townsend G, Kullmer O, 2020. The functional role of the Carabelli trait in early and late hominins. Journal of Human Evolution, 145, 102816   Gokhman D, Nissim-Rafinia M, Agranat-Tamir L, Housman G, García-Pérez R, Lizano E, Cheronet O, Mallick S, Nieves-Colón M, Li H, Alpaslan-Roodenberg S, Novak M, Gu H, Osinski J, Ferrando-Bernal M, Gelabert P, Lipende I, Mjungu D, Kondova I, Bontrop R, Kullmer O, Weber G, Shahar T, Dvir-Ginzberg M, Faerman M, Quillen E, Meissner A, Lahav Y, Kandel L, Liebergall M, Prada M E, Encinas J M V, Gronostajski R, Stone A, Yakir B, Lalueza-Fox C, Pinhasi R, Reich D, Marques-Bonet T, Meshorer E, Carmel L, 2020. Differential DNA methylation of vocal and facial anatomy genes in modern humans. Nature Communications, 11, 1189. https://doi.org/10.1038/s41467-020-15020-6.   Zanolli C, Schillinger B, Kullmer O, Schrenk F, Kelley J, Rössner GE, Macchiarelli R, 2020. When X-Rays Do Not Work. Characterizing the Internal Structure of Fossil Hominid Dentognathic Remains Using High-Resolution Neutron Microtomographic Imaging. Front. Ecol. Evol. 8:42. doi:10.3389/fevo.2020.00042   Gunz P, Kozakowski S, Neubauer S, Le Cabec A, Kullmer O, Benazzi S, Hublin J-J, Begun D R, 2020. Skull reconstruction of the late Miocene ape Rudapithecus hungaricus from Rudabánya, Hungary. Journal of Human Evolution, 138, 102687  
Joannes-Boyau R, Adams J W, Austin C, Arora M, Moffat I, Herries A I, Tonge M P, Benazzi S, Evans A R, Kullmer O, Wroe S, Dosseto A, Fiorenza L 2019. Elemental signatures of Australopithecus africanus teeth reveal seasonal dietary stress. Nature, 572(7767), 112-115.

Zanolli C, Kullmer O, Kelley J, Bacon A-M, Demeter F, Dumoncel J, Fiorenza L, Grine FE, Hublin J-J, Nguyen AT, Nguyen TMH, Lei P, Schillinger B, Schrenk F, Skinner MM, Xueping J, Macchiarelli R, 2019. Evidence for increased hominid diversity in the Early to Middle Pleistocene of Indonesia. Nature Ecology & Evolution, 3: 755–764.
Fiorenza L, Benazzi S, Kullmer O, Zampirolo G, Mazurier A, Zanolli C, Macchiarelli R 2019. Dental macrowear and cortical bone distribution of the Neanderthal mandible from Regourdou (Dordogne, Southwestern France). Journal of Human Evolution, 132, 174-188.
Lüdecke T, Kullmer O, Wacker U, Sandrock O, Fiebig J, Schrenk F, Mulch A 2018. Dietary versatility of Pleistocene hominins. PNAS, 115(52), 13330-13335.
Smith TM, Houssaye A, Kullmer O, Le Cabec A, Olejniczak AJ, Schrenk F, de Vos J, Tafforeau P, 2018. Disentangling isolated dental remains of Asian Pleistocene hominins and pongines. PloS ONE, 13(11), e0204737.
Wroe S, Parr WC, Ledogar JA, Bourke J, Evans SP, Fiorenza L, Benazzi S, Hublin J-J, Stringer C, Kullmer O, Curry M, Rae TC, Yokley TR, 2018. Computer simulations show that Neanderthal facial morphology represents adaptation to cold and high energy demands, but not heavy biting. Proceedings of the Royal Society B, 285(1876), 20180085.
Oxilia G, Bortolini E, Martini S, Papini A, Boggioni M, Buti L, Figus C, Sorrentino R, Townsend G, Kaidonis J, Fiorenza L, Cristiani E, Kullmer O, Moggi-Cecchi J, Benazzi S, 2018. The physiological linkage between molar inclination and dental macrowear pattern. American Journal of Physical Anthropology. 166:941-951.
Zanolli C, Pan L, Dumoncel J, Kullmer O, Kundrát M, Liu W, Macchiarelli R, Mancini L, Schrenk F, Tuniz C, 2018. Inner tooth morphology of Homo erectus from Zhoukoudian. New evidence from an old collection housed at Uppsala, C. 2018. University, Sweden. Journal of Human Evolution, 116, 1-13.
Fiorenza L, Benazzi S, Oxilia G, Kullmer O, 2018. Functional relationship between dental macrowear and diet in Late Pleistocene and recent modern human populations. International Journal of Osteoarchaeology, 28(2), 153-161.
Schillinger B, Beaudet A, Fedrigo A, Grazzi F, Kullmer O, Laaß M, Makowska M, Werneburg I, Zanolli C, 2018. Neutron Imaging in Cultural Heritage Research at the FRM II Reactor of the Heinz Maier-Leibnitz Center. Journal of Imaging, 4, 22; doi:10.3390/jimaging4010022.
Zanolli C, Schillinger B, Beaudet A, Kullmer O, Macchiarelli R, Mancini L, Schrenk F, Tuniz C, Vodopivec V, 2017. Exploring hominin and non-hominin primate dental fossil remains with neutron microtomography. Physics Procedia, 88, 109-115.
Schultz JA, Menz U,  Winkler DE, Schulz-Kornas E, Engels S, Kalthoff DC, von Koenigswald W, Ruf I, Kaiser TM, Kullmer O, Südekum K-H, Martin T 2017. Modular Wear Facet Nomenclature for mammalian post-canine dentitions, Historical Biology: 1-12.
Bocherens H, Schrenk F, Chaimanee Y, Kullmer O, Mörike D, Pushkina D, Jaeger JJ, 2017. Flexibility of diet and habitat in Pleistocene South Asian mammals: Implications for the fate of the giant fossil ape Gigantopithecus. Quaternary International. 434 (A):148-155.
Fiorenza L, Kullmer O 2016. Occlusion in an Adult Male Gorilla with a Supernumerary Maxillary Premolar. International Journal of Primatology. DOI: 10.1007/s10764-016-9937-6.
Luedecke T, Schrenk F, Thiemeyer H, Kullmer O, Bromage TG, Sandrock O, Fiebig J, Mulch A 2016. Persistent C3 vegetation accompanied Plio-Pleistocene hominin evolution in the Malawi Rift (Chiwondo Beds, Malawi). Journal of Human Evolution, 90:163-175.
Benazzi S, Nguyen HN, Kullmer O, Kupczik K 2016. Dynamic Modelling of Tooth Deformation Using Occlusal Kinematics and Finite Element Analysis. PLoS ONE 11(3): e0152663. doi:10.1371/journal.pone.0152663
Luedecke T, Mulch A, Kullmer O, Sandrock O, Thiemeyer H, Fiebig J, Schrenk F 2016. Stable isotope dietary reconstructions of herbivore enamel reveal heterogeneous savanna ecosystems in the Plio-Pleistocene Malawi Rift. Palaeogeography, Palaeoclimatology, Palaeoecology, 479:170-181.
Fiorenza L, Benazzi S, Henry AG, Salazar-García DC, Blasco R, Picin A, Wroe S, Kullmer O 2015. To Meat or Not to Meat? New Perspectives on Neanderthal Ecology. American Journal of Physical Anthropology (Yearbook of Physical Anthropology), 156 (Suppl.59):43–71.
Fiorenza L, Kullmer O 2015. Dental wear patterns in early modern humans from Skhul and Qafzeh: A response to Sarig andTillier. Homo – Journal of Comparative Biology, published online ahead of print, DOI:10.1016/j.jchb.2015.04.002.
Zanolli C, Grine FE, Kullmer O, Schrenk F, Macchiarelli R 2015. Brief Communication: The Early Pleistocene Deciduous Hominid Molar FS-72 From the Sangiran Dome of Java, Indonesia: A Taxonomic Reappraisal Based on its Comparative Endostructural Characterization. American Journal of Physical Anthropology, 157(4):666–674.
Oxilia G, Peresani M, Romandini M, Matteucci C, Spiteri CD, Henry AG, Schulz D, Archer W, Crezzini J, Boschin F, Boscato P, Jaouen K, Dogandzic T, Broglio A, Moggi-Cecchi J, Fiorenza L, Hublin J-J, Kullmer O, Benazzi S 2015. Earliest evidence of dental caries manipulation in the Late Upper Palaeolithic. Scientific Reports, 5, 12150. DOI:10.1038/srep12150
Benazzi S, Nguyen HN, Kullmer O, Hublin J-J 2015. Exploring the biomechanics of taurodontism. Journal of Anatomy, 226 (2):180-188.
Tausch J, Kullmer O, Bromage TG 2015. A New Method for Determining the 3D Spatial Orientation of Molar Microwear, Scanning, 9999:1-12. published online ahead of print DOI: 10.1002/sca.21234.
Shar S, Batsaikhan N, Dolch D, Gardner S, Kullmer O, Lebedev VS, Lkhagvasuren D, Menz U, Samiya R, Stubbe M, Ansorge H 2015. First Report of the Herb Field Mouse, Apodemus uralensis (Pallas, 1811) from Mongolia. Mongolian Journal of Biological Sciences, 13 (1-2). pp. 35-42.
Kullmer O, 2015: Der verlorene Biss: Biologische und kulturelle Evolution der Zähne. In (H. Böhme, B. Kordaß, B. Slominski Hrsg.) Das Dentale – Faszination des oralen Systems in Wissenschaft und Kultur. S.65-86, Quintessenz, Berlin.
Emonet E-G, Kullmer O 2014. Variability in Premolar and Molar Root Number in a Modern Population of Pan troglodytes verus. The Anatomical Record 297(10): 1927-1934. DOI: 10.1002/ar.22959.
Weber GW, Benazzi S, Kullmer O 2014. Mikro-CT-Anwendungen in der biologischen Anthropologie. Journal für Mineralstoffwechsel 21(2):68-68f.
Benazzi S, Grosse,IR, Gruppioni,G, Weber GW, Kullmer O 2014. Comparison of occlusal loading conditions in a lower second premolar using three-dimensional finite element analysis. Clinical Oral Investigations, 18: 369-375.
Benazzi S, Nguyen HN, Kullmer O, Hublin J 2013. Unravelling the Functional Biomechanics of Dental Features and Tooth Wear. PLoS ONE 8(7): e69990. doi:10.1371/journal.pone.0069990
Fiorenza L, Kullmer O 2013. Dental wear and cultural behavior in Middle Paleolithic humans from the Near East. American Journal of Physical Anthropology, 152: 107–117. doi: 10.1002/ajpa.22335
Benazzi S, Nguyen NH, Schulz D, Grosse IR, Gruppioni G, Hublin J-J, Kullmer O 2013. The Evolutionary Paradox of Tooth Wear: Simply Destruction or Inevitable Adaptation? PLoS ONE 8(4): e62263.doi:10.1371/journal.pone.0062263.
Harvati K, Darlas A, Bailey SE, Rein TR, El Zaatari S, Fiorenza F, Kullmer O, Psathi E 2013. New Neanderthal remains from Mani peninsula, Southern Greece: The Kalamakia Middle Paleolithic cave site, Journal of Human Evolution, 64(6):486-499.
Weber GW, Benazzi S, Kullmer O 2013. CT-Anwendungen in der biologischen Anthropologie. Osteologie/Osteology, 22 (1):18-24.
Kullmer O, Benazzi S, Schulz D, Gunz P,  Kordos L, Begun DR 2013. Dental arch restoration using tooth macrowear patterns with application to Rudapithecus hungaricus, from the late Miocene of Rudabánya, Hungary. Journal of Human Evolution, 64(2):151-160.
Benazzi S, Kullmer O, Schulz D, Weber GW 2013. Individual tooth macrowear pattern guides the reconstruction of Sts 52 (A. africanus) dental arches. American Journal of Physical Anthropology, 150: 324-329.
von Koenigswald W, Anders U, Engels S, Schultz JA, Kullmer O 2013. Jaw movement in fossil mammals: analysis, description and visualization. Paläontologische Zeitschrift, 87:141–159, doi:10.1007/s12542-012-0142-4.
Kullmer O, Schulz D, Benazzi S 2012. An Experimental Approach to Evaluate the Correspondence Between Wear Facet Position and Occlusal Movements. The Anatomical Record, 295: 846–852.
Brügmann G, Krause J, Brachert TC, Stoll B, Weis U, Kullmer O, Ssemmanda I, Mertz DF, 2012. Chemical composition of modern and fossil Hippopotamid teeth and implications for paleoenvironmental reconstructions and enamel formation – Part 2: Alkaline earth elements as tracers of watershed hydrochemistry and provenance. Biogeosciences Discuss., 9, 3645-3691, doi:10.5194/bgd-9-3645-2012.
Böhme M, Aiglstorfer M, Uhl D, Kullmer O 2012. The Antiquity of the Rhine River: Stratigraphic Coverage of the Dinotheriensande (Eppelsheim Formation) of the Mainz Basin (Germany). PLoS ONE 7(5): e36817. doi:10.1371/journal.pone.0036817
Brügmann G, Krause J, Brachert TC, Kullmer O, Schrenk F, Ssemmanda I, Mertz DF 2012. Chemical composition of modern and fossil Hippopotamid teeth and implications for paleoenvironmental reconstructions and enamel formation – Part 1: Major and minor element variation, Biogeosciences, 9, 119–139.
Smith TM, Olejniczak AJ, Zermeno JP, Tafforeau P, Skinner MM, Hoffmann A, Radovčić J, Toussaint M, Kruszynski R, Menter C, Moggi-Cecchi J, Glasmacher UA, Kullmer O, Schrenk F, Stringer C, Hublin J-J 2012. Variation in enamel thickness within the genus Homo. Journal of Human Evolution 62:395-411.
Benazzi S, Kullmer O, Grosse IR, Weber GW, 2012. Brief Communication: Comparing Loading Scenarios in Lower First Molar Supporting Bone Structure Using 3D Finite Element Analysis. American Journal of Physical Anthropology 147(1):128-134.
Denys C, Sandrock O, Kullmer O, Ramirez Rozzi F, Bromage TG, Schrenk F 2011. Note on the presence of Hystrix (Mammalia, Rodentia) in the Malawi Chiwondo Beds (Plio-Pleistocene): taphonomical and palaeoecological implications. Geodiversitas 33 (4): 729-738.
Kullmer O, Sandrock O, Kupczik K, Frost SR, Volpato V, Bromage TG, Schrenk F 2011. New Primate Remains from Mwenirondo, Chiwondo Beds in northern Malawi, Journal of Human Evolution, 61:617-623.
Benazzi S, Douka K, Fornai C, Bauer CC, Kullmer O, Svoboda J, Pap I, Mallegni F, Bayle P, Coquerelle M, Condemi S, Ronchitelli A, Harvati K, Weber GW 2011. Early dispersal of modern humans in Europe and implications for Neanderthal behavior. NATURE, 479, 525-528.
Smith TM, Bacon A-M, Demeter F, Kullmer O, Nguyen KT, de Vos J, Wei W, Zermeno JP, Zhao L 2011. Dental tissue proportions in fossil orangutans from mainland Asia and Indonesia. Human Origins Research 2011; volume 1:e1
Benazzi S, Viola B, Kullmer O, Fiorenza L, Harvati K, Paul T, Gruppioni G, Weber GW, Mallegni F 2011. A reassessment of the Neanderthal teeth from Taddeo cave (southern Italy). Journal of Human Evolution, 61(4):377-387.
Benazzi S, Fornai C, Bayle P, Coquerelle M, Kullmer O, Mallegni F, Weber G 2011. Comparison of dental measurement systems for taxonomic assignment of Neanderthal andmodern human lower second deciduous molars. Journal of Human Evolution, 61(3):320-326.
Fiorenza L, Benazzi S, Kullmer O 2011. Para-masticatory wear facets and their functional significance in hunter–gatherer maxillary molars. Journal of Archaeological Sciences, 38 (9): 2182-2189.
Benazzi S, Kullmer O, Grosse IR, Weber GW 2011. Using occlusal wear information and finite element analysis to investigate stress distributions in human molars. Journal of Anatomy, 219:259–272.
Benazzi S, Fiorenza L, Kozakowski S, Kullmer O 2011. Comparing 3D Virtual Methods for Hemimandibular Body Reconstrution. The Anatomical Record, 294: 1116–1125.
Bocherens H, Sandrock O, Kullmer O, Schrenk F 2011. Hominin palaeoecology in Late Pliocene Malawi: First insights from isotopes (13C, 18O) in mammal teeth. South African Journal of Science, 107 (3/4), Art. #331, 6 pages. DOI: 10.4102/sajs.v107i3/4.331
Fiorenza L, Benazzi S, Tausch J, Kullmer O, Bromage TG, Schrenk F 2011. Molar Macrowear Reveals Neanderthal Eco-Geographic Dietary Variation. PLoS ONE 6(3): e14769. doi:10.1371/journal.pone.0014769
Benazzi S, Fiorenza L, Katina S, Bruner E, Kullmer O 2011. Quantitative assessment of interproximal wear facet outlines for the association of isolated molars, American Journal of Physical Anthropology,144(2):309–316.
Fiorenza L, Benazzi S, Kullmer O, Schrenk F 2011. Relationship Between Cusp Size and Occlusal Wear Pattern in Neanderthal and Homo sapiens First Maxillary Molars, The Anatomical Record, 294:453-461.
Benazzi S, Coquerelle M, Fiorenza L, Bookstein F, Katina S, Kullmer O 2011. Comparison of dental measurement systems for taxonomic assignment of first molars. American Journal of Physical Anthropology, 144(3):342–354.

FF PaAn Denkel-Oswalt
Birgit Denkel-Oswalt

Field of work

  • Organisatorisches und administratives Büromanagement
  • Terminkoordination und -controlling
  • Interne/externe Kommunikation
  • Mitwirkung bei der Entwicklung, Planung und Durchführung von internationalen Forschungs- und Kooperationsprojekten
  • Wissenschaftsadministration (u.a. Drittmittel- und Reisekostenabrechnung)
  • Unterstützung bei der Sammlungsbetreuung
  • Vortrags- und Veranstaltungsorganisation
  • Gastforscherbetreuung
  • Ansprechpartnerin für Studierende, Doktoranden und Postdocs
  • Haushaltsüberwachung
FF PaAn Christine Hemm
Christine Hemm
Technical Assistant

Field of work

  • 3 D Labor (3D-Sammlungsbetreuung, 3-D-Vermessungen und Rekonstruktionen,  Ausbilden, Verarbeitung der Daten, Sicherung, Zugriff etc.)
  • Zeichnungen, Graphiken herstellen, Grabungspläne zeichnen
  • Betreuung der Bibliothek (Sonderdrucke, Bibliotheksarbeiten…)
  • Betreuung der Dauerausstellung
  • Ausgrabungen, Survey
  • Betreuung der Praktikanten
  • SNG-Schülerausbildung
  • Veranstaltungsorganisation
  • Gastforscherbetreuung (technisch)
  • Homepage der Sektionen Paläoanthropologie und tertiäreSäugetiere

2007: Einsatzmöglichkeiten von 3D-Scannern in der Paläontologie und deren Anwendung. Der Präparator

Prof. Dr. Katharina Neumann
Honorary post, Project staff

DFG-Projekt: ‘Plant environment and resource exploitation in SW Ethiopia >50 ka to the present’

DFG-Nr NE 408/16-1, 2021-2024. PIs: Katharina Neumann und Karen Hahn.  
Professional background

Associate Professor, Goethe University Frankfurt

Habilitation, Goethe University Frankfurt

Since 1994
Leader of working group „African Archaeobotany“, Institute of Archaeological Sciences, Goethe University Frankfurt

Research assistant, Goethe University Frankfurt

PhD (Dr), Goethe University Frankfurt

Research assistant, Institute for Pre- and Protohistory, University of Cologne

Diploma, botany, Goethe University Frankfurt

* with peer-review, books


* Casas-Gallego, M., Hahn, K., Neumann, K., Demissew, S., Schmidt, M., Bodin, S., Bruch, A.A.: Cooling‐induced expansions of Afromontane forests in the Horn of Africa since the Last Glacial Maximum. Scientific Reports (2023) 13:10323. https://doi.org/10.1038/s41598- 023-37135-8


* Champion, L., Höhn, A., Neumann, K. et al. A question of rite – pearl millet consumption at Nok culture sites, Nigeria (second/first millennium BC). Veget Hist Archaeobot (2022). https://doi.org/10.1007/s00334-022-00902-0

* Dunne, J., Höhn, A., Neumann, K., Franke, G., Breunig, P., Champion, L., Gillard, T., Walter-Doyle, C. & Evershed, R.P.: Making the invisible visible: tracing the origins of plants in West African cuisine through archaeobotanical and organic residue analysis. Archaeological and Anthropological Sciences 14:30. doi.org/10.1007/s12520-021-01476-0

* Hensel, E.A., Kehl, M., Wöstehoff, L., Neumann, K., Vogelsang, R. & Bubenzer, O.: A multi-method approach for deciphering rockshelter microstratigraphies – the role of the Sodicho Rockshelter (SW Ethiopia) as a geoarchaeological archive. Geosciences 2022, 12, 92. https://doi.org/10.3390/geosciences12020092

* Neumann, K., Eichhorn, B. & Wotzka, H.-P.: Iron Age plant subsistence in the Inner Congo Basin (DR Congo). Vegetation History and Archaeobotany (2022) 31:481–509. https://doi.org/10.1007/s00334-021-00865-8


* Dunne, J., Höhn, A., Franke, G., Neumann, K., Breunig, P., Gillard, T., Walter-Doyle, C. & Evershed, R.P.: Honey-collecting in prehistoric West Africa from 3500 years ago. Nature Communications 12: 2227. https://doi.org/10.1038/s41467-021-22425-4


* Franke, G., Höhn, A., Schmidt, A., Ozainne, S., Breunig, P., Neumann, K.: Pits, pots and plants at Pangwari – Deciphering the nature of a Nok Culture site. Azania: Archaeological Research in Africa. https://doi.org/10.1080/0067270X.2020.1757902


* Kay, A.U., Fuller, D.Q., Neumann, K., Eichhorn, B., Höhn, A., Morin‐Rivat, J., Champion, L., Linseele, V., Huysecom, E., Ozainne, S., Lespez, L., Biagetti, S., Madella, M., Salzmann, U. & Kaplan, J.O.: Diversification, Intensification and Specialization: Changing Land Use in Western Africa from 1800 BC to AD 1500. Journal of World Prehistory 32:179–228. doi.org/10.1007/s10963-019-09131-2

* Neumann, K., Albert, R.M., Ball, T., Scott Cummings, L., Strömberg, C.A. & Vrydaghs, L. (International Committee for Phytolith Taxonomy, ICPT): International Code of Phytolith Nomenclature, ICPN 2.0. Annals of Botany 124(2):189-199. https://doi.org/10.1093/aob/mcz064


* Clist, B., Bostoen, K., de Maret, P., Eggert, M., Höhn, A., Mbida Mindzie, C., Neumann, K. & D. Seidensticker: Did human activity really trigger the Late Holocene rainforest crisis in Central Africa? Proceedings of the National Academy of Sciences. www.pnas.org/cgi/doi/10.1073/pnas.1805247115

* Neumann, K.: Development of Plant Food Production in the West African Savannas: Archaeobotanical Perspectives. Oxford Research Encyclopedia of African History. https://doi: 10.1093/acrefore/9780190277734.013.138


* Collura, L.V. & Neumann, K.: Wood and bark phytoliths of West African woody plants. Quaternary International 434: 142-159.

Neumann, K.: Klima, Mensch und Vegetation im Holozän Westafrikas. In: Rupp, N. Beck, C., Franke, G. & Wendt, K.P. (eds.): Winds of Change. Archaeological Contributions in Honour of Peter Breunig. Bonn (Rudolf Habelt): 343-354.

* Neumann, K., Fahmy, A., Müller-Scheeßel, N. & Schmidt, M.: Taxonomic, ecological and palaeoecological significance of leaf phytoliths in West African grasses. Quaternary International 434: 13-32.


* Höhn, A. & Neumann, K.: Palaeovegetation of Janruwa (Nigeria) and its implications for the decline of the Nok Culture. Journal of African Archaeology 14(3): 331-353.

* Neumann, K., Chevalier, A. & Vrydaghs, L. Phytoliths in archaeology: recent advances. Vegetation History and Archaeobotany. doi 10.1007/s00334-016-0598-3


Neumann, K.: In memoriam Ahmad Gamal el-Din Fahmy, March 5th 1962 to December 18th 2013. Vegetation History and Archaeobotany 24: 3-4.


* Eichhorn, B. & Neumann, K.: Holocene vegetation change and land use at Ounjougou, Mali. – In: Stevens, C.J., Nixon, S., Murray, M.A. & Fuller, D. (eds.): Archaeology of African Plant Use. Walnut Creek (Left Coast Press): 83-96.

Höhn, A. & Neumann, K.: Millet and more: Farming and food in the time of the Nok Culture. – In: In: Breunig, P. (ed.): Nok – African Sculpture in Archaeological Context. Frankfurt am Main (Africa Magna Verlag): 178-185.

* Kahlheber, S., A. Höhn & Neumann, K.: Plant use in southern Cameroon between 400 BC and 400 AD. – In: Stevens, C.J., Nixon, S., Murray, M.A. & Fuller, D. (eds.): Archaeology of African Plant Use. Walnut Creek (Left Coast Press): 113-128.

Neumann, K. & Höhn, A.: The Nok region – a savanna landscape and its history. – In: Breunig, P. (ed.): Nok – African Sculpture in Archaeological Context. Frankfurt am Main (Africa Magna Verlag): 40-49.

* Ozainne, S., Lespez, L., Garnier, A., Ballouche, A., Neumann, K., Pays, O. & Huysecom, E.: A question of timing: spatio-temporal structure and mechanisms of early agriculture expansion in West Africa. Journal of Archaeological Science 50: 359-368.


* Garnier, A., Neumann, K., Eichhorn, B. & Lespez, L. Phytolith taphonomy in the middle- to late-Holocene fluvial sediments of Ounjougou (Mali, West Africa). The Holocene 23(3): 416-431.

Wittig, R. & Neumann, K. Die westafrikanische Savanne – ein Lebensraum unter starkem anthropogenem Einfluss. Berichte der Reinhold-Tüxen-Gesellschaft 25. 150-165.


Barton, H., T. Denham, K. Neumann, M. Arroyo-Kalin: Long-term perspectives on human occupation of tropical rainforests: An introductory overview. Quaternary International 249: 1- 3.

* Höhn, A. & Neumann, K.: Shifting cultivation and cultural landscape development during the Iron Age (0-1500 AD) in the Sahel of Burkina Faso, West Africa – Insights from archaeological charcoal. Quaternary International 249: 72-83.

Neumann, K. & Ballouche, A. Das Pollenprofil von Oursi und die Diskussion um die “menschengemachte” westafrikanische Savanne. In: Stobbe, A. & Tegtmeier, U. (ed.) Verzweigungen. Eine Würdigung für A.J. Kalis und J. Meurers-Balke. Rudolf Habelt, Köln, pp. 199-206.

* Neumann, K., K. Bostoen, A. Höhn, S. Kahlheber, A. Ngomanda & Tchiengué, B.: First farmers in the Central African rainforest: A view from southern Cameroon. Quaternary International 249: 53-62. doi:10.1016/j.quaint.2011.03.024

* Neumann, K., M.K.H. Eggert, R. Oslisly, B. Clist, T. Denham, P. de Maret, S. Ozainne, E. Hildebrand, K. Bostoen, U. Salzmann, D. Schwartz, B. Eichhorn, B. Tchiengué & Höhn, A.: Comment on “Intensifying Weathering and Land Use in Iron Age Central Africa”. Science 337: 1040. doi: 0.1126/science.1221747


* Lespez, L., Y. Le Drezen, A. Garnier, M. Rasse, B. Eichhorn, S. Ozainne, A. Ballouche, K. Neumann & Huysecom, E.: High-resolution fluvial records of Holocene environmental changes in the Sahel: the Yamé River at Ounjougou (Mali, West Africa). Quaternary Science Reviews 30 (5-6): 737-756.

* Radomski, K.U. & Neumann, K.: Grasses and grinding stones: Inflorescence phytoliths from modern West African Poaceae and archaeological stone artefacts. – In: Fahmy, A.G., S. Kahlheber & A.C. Andrea (eds): Windows on the African Past. Current Approaches to African Archaeobotany. Frankfurt am Main (Africa Magna Verlag): 153-166.

* Sangen, M., K. Neumann & Eisenberg, J.: Climate-induced dynamics in tropical Africa around the last glacial maximum? Quaternary Research 76: 417-429.


* Eichhorn, B., Neumann, K., Garnier, A.: Seed phytoliths in West African Commelinaceae and their potential for palaeoecological studies. Palaeogeography, Palaeoclimatology, Palaeoecology 298: 300-310. doi:10.1016/j.palaeo.2010.10.004


* Huysecom, E., M. Rasse, L. Lespez, K. Neumann, A. Fahmy, A. Ballouche, S. Ozainne, M. Maggetti, Ch. Tribolo, S. Soriano: The emergence of pottery in Africa during the 10th millennium cal BC: new evidence from Ounjougou (Mali). Antiquity 83 (322): 905-917.

* Kahlheber, S., K. Bostoen & K. Neumann: Early plant cultivation in the Central African rain forest: first millennium BC pearl millet from South Cameroon. Journal of African Archaeology 7(2): 253-272.

* Neumann, K., A. Fahmy, L. Lespez, A. Ballouche, E. Huysecom: The Early Holocene palaeoenvironment of Ounjougou (Mali): Phytoliths in a multiproxy context. Palaeogeograpphy, Palaeoclimatology, Palaeoecology 276: 187-206. doi:10.1016/j.palaeo.2009.03.001

* Neumann, K. & E. Hildebrand: Early bananas in Africa: The state of the art. Ethnobotany Research and Applications 7: 353-362.

* Ngomanda, A., K. Neumann, A. Schweizer & J. Maley: Seasonality change and the third millennium BP rainforest crisis in Central Africa: a high resolution pollen profile from Nyabessan, southern Cameroon. Quaternary Research 71: 307-318. doi:10.1016/j.yqres.2008.12.002

Ozainne, S., L. Lespez, Y. Le Drezen, B. Eichhorn, K. Neumann & E. Huysecom: Developing a chronology integrating archaeological and environmental data from different contexts: the Late Holocene sequence of Ounjougou (Mali). Radiocarbon 51(2): 457–470.


* Kahlheber, S. & K. Neumann: The development of plant cultivation in semi-arid west Africa. In: Denham, T. P., J. Iriarte & L. Vrydaghs (eds.): Rethinking Agriculture: Archaeological and Ethnoarchaeological Perspectives. One World Archaeology 51. Walnut Creek (Left Coast Press): 320-346.

* Höhn, A., S. Kahlheber, K. Neumann & A. Schweizer: Settling the rain forest – the environment of farming communities in Southern Cameroon during the first millennium BC. In: Runge, J. (ed.): Palaeoecology of Africa 28. London (Taylor & Francis): 29-41.


Breunig, P., B. Eichhorn, S. Kahlheber, V. Linseele, C. Magnavita, K. Neumann, M. Posselt & N. Rupp: G(l)anz ohne Eisen: Große Siedlungen aus der Mitte des ersten Jahrtausends BC im Tschadbecken von Nordost-Nigeria. In: Wotzka, H.-P. (Hrsg.), Grundlegungen. Beiträge zur afrikanischen Archäologie für Manfred K.H. Eggert. Tübingen (Francke): 255-270.

Neumann, K.: Ölpalme, Perlhirse und Banane. Wie kam die Landwirtschaft in den Regenwald Zentralafrikas? Forschung Frankfurt 2-3/2006: 38-41.

* Eggert, M.K.H., A. Höhn, S. Kahlheber, C. Meister, K. Neumann & A. Schweizer: Pits, graves and grains. Archaeological and archaeobotanical research in southern Cameroun. Journal of African Archaeology 4(2): 273-298.

* Rasse, M., A. Ballouche, E. Huysecom, C. Tribolo, S. Ozainne, Y. Le Drezen, S. Stokes & K. Neumann: Évolution géomorphologique, enregistrements sédimentaires et dynamiques paléoenvironnementales Holocènes à Ounjougou (Plateau Dogon, Mali, Afrique de l’Ouest). Quaternaire 17 (1): 61-74.


* Neumann, K.: The romance of farming – plant cultivation and domestication in Africa. In: A.B. Stahl (ed.): African Archaeology. A Critical Introduction: 249-275. Malden (Blackwell).


Albert, K.-D., D. Löhr & K. Neumann (eds.): Mensch und Natur in Westafrika. Abschlussbuch des Sonderforschungsbereichs 268. Weinheim (Wiley-VCH).

Breunig, P. & K. Neumann: Zwischen Wüste und Regenwald. Besiedlungsgeschichte der westafrikanischen Savannen im Holozän. In: Albert, K.-D., Löhr, D. & K. Neumann (eds.): Mensch und Natur in Westafrika. Abschlussbuch des Sonderforschungsbereichs 268: 93-138. Weinheim (Wiley-VCH).

Neumann, K., K. Hahn-Hadjali & U. Salzmann: Die Savannen der Sudanzone in Westafrika – natürlich oder menschengemacht? In: Albert, K.-D., D. Löhr & K. Neumann (eds.): Mensch und Natur in Westafrika. Abschlussbuch des Sonderforschungsbereichs 268: 39-68. Weinheim (Wiley – VCH).


*  Neumann, K.: New Guinea: A Cradle of Agriculture. Science 301: 180-181.

*  Neumann, K.: The late emergence of agriculture in Sub-Saharan Africa: Archaeological evidence and ecological considerations. In: Neumann, K., A. Butler & S. Kahlheber (eds.): Food, Fuel and Fields. Progress in African Archaeobotany: 71-92. Africa Praehistorica 15. Köln (Heinrich-Barth-Institut).

* Neumann, K., A. Butler & S. Kahlheber (eds.): Food, Fuel and Fields. Progress in African Archaeobotany. Africa Praehistorica 15. Köln (Heinrich-Barth-Institut).


* Breunig, P. & K. Neumann: From hunters and gatherers to food producers: new archaeological and archaeobotanical evidence from the West African Sahel. In: F. Hassan (ed.): Ecological Change and Food Security in Africa’s Later Prehistory: 123-155. New York (Kluwer Academic / Plenum Publishers).

* Breunig, P. & K. Neumann: Continuity or discontinuity? The 1st millennium BC-crisis in West African prehistory. – In: Lenssen-Erz, T. et al. (ed.): Tides of the Desert. Contributions to the Archaeology and Environmental History of Africa in Honour of Rudolph Kuper: 491- 505. Africa Praehistorica 14, Köln (Heinrich-Barth-Institut).

Neumann, K.: Die westafrikanische Savanne – eine Kulturlandschaft. In: Reikat, A. (ed.): Leben in Westafrika: 70-83. Frankfurt a. Main (Plexus-Verlag, Johann Wolfgang Goethe- Universität, SFB 268 “Kulturentwicklung und Sprachgeschichte im Naturraum Westafrikanische Savanne).


Ballouche, A., A. Akoègninou, K. Neumann, U. Salzmann & M.A. Sowunmi: Le projet “Dahomey Gap”: une contribution à l’histoire de la végétation au Sud-Bénin et Sud-Ouest du Nigéria. Berichte des Sonderforschungsbereichs 268, 14: 237-251. Frankfurt am Main.

Frank, Th., P. Breunig, P. Müller-Haude, W. Van Neer, K. Neumann, R. Vogelsang & H.-P. Wotzka: The Chaîne de Gobnangou in SE Burkina Faso: archaeological, archaeobotanical, archaeozoological and geomorphological studies. Ber. Allg. Vergl. Arch. 127-190.

Kahlheber, S. & K. Neumann (eds.): Man and Environment in the West African Sahel – an Interdisciplinary Approach. Berichte des Sonderforschungsbereichs 268, 17, Frankfurt am Main.

Neumann, K.: Masakwa farming in the Chad Basin of northeast Nigeria – Introduction. In: Kahlheber, S. & K. Neumann (eds.): Man and Environment in the West African Sahel – an Interdisciplinary Approach: 9-13. Berichte des Sonderforschungsbereichs 268, 17, Frankfurt am Main.

* Neumann, K., W. Schoch, F.H. Schweingruber & P. Détienne: Hölzer der Sahara und des Sahel – ein anatomischer Atlas. Paul Haupt, Bern & Stuttgart.


* Klee, M., B. Zach & K. Neumann: Four thousand years of plant exploitation in the Chad Basin of northeast Nigeria: The archaeobotany of Kursakata. Vegetation History and Archaeobotany 9: 223-237.


Breunig, P. & K. Neumann: Archäologische und archäobotanische Forschung in Westafrika. Archäologisches Nachrichtenblatt 4: 336-357.

* Neumann, K.: Charcoal from West African savanna sites – questions of identification and interpretation. In: Van der Veen M (ed.) The exploitation of plant resources in ancient Africa. Plenum Publishers, New York: 205-219.

* Neumann, K.: Early plant food production in the West African Sahel – new evidence from the Frankfurt project. In: Van der Veen M (ed.) The Exploitation of Plant Resources in Ancient Africa. Plenum Publishers, New York: 73-80.

* Neumann, K. & P. Müller-Haude: Forêts sèches au sud-ouest du Burkina Faso: végétations – sols – action de l’homme. Phytocoenologia 29(1): 53-85.


* Neumann, K., S. Kahlheber & D. Uebel: Remains of woody plants from Saouga, a medieval West African village. Vegetation History and Archaeobotany 7(2): 57-77.


Neumann, K.: Report on the second workshop on the archaeobotany of North Africa. Nyame Akuma 48: 79.


* Breunig, P., Neumann, K. & W. Van Neer: New research on the Holocene settlement and environment of the Chad Basin in Nigeria. African Archaeological Review 13(2): 111-145.

* Neumann, K., A. Ballouche & M. Klee: The emergence of plant food production in the West African Sahel: new evidence from northeast Nigeria and northern Burkina Faso. In: Pwiti, G. & R. Soper (eds.): Aspects of African Archaeology. Papers from the 10th Congress of the PanAfrican Association for Prehistory and Related Studies: 441-448; Harare (University of Harare Publications).

Zach, B., H. Kirscht, D. Löhr, K. Neumann & E. Platte: Masakwa dry season cropping in the Chad Basin. Berichte des Sonderforschungsbereichs 268 8: 349-356, Frankfurt am Main.


* Ballouche, A. & K. Neumann: A new contribution to the Holocene vegetation history of the West African Sahel: pollen from Oursi, Burkina Faso and charcoal from three sites in northeast Nigeria. Vegetation History and Archaeobotany 4: 31-39.

Ballouche, A. & K. Neumann: La végétation du Sahel Burkinabé à l’Holocène: La Mare d’Oursi. In: 2nd Symposium on African Palynology, Tervuren (Belgium): 19-25; Occasional Publication CIFEG 31, Orléans.

Neumann, K. & A. Ballouche: Anthropogenic change in the vegetation history of the Sahel: only a question of viewpoint? In: 2nd Symposium on African Palynology, Tervuren (Belgium): 27-33; Occasional Publication CIFEG 31, Orléans.


Neumann, K.: Wirtschaftsweisen im Neolithikum der Ostsahara und ihr Einfluß auf die Vegetation. In: Bollig, M. & F. Klees (ed.): Überlebensstrategien in Afrika: 47-65; Köln (Heinrich-Barth-Institut).


Ballouche, A., K. Küppers, K. Neumann & H.P. Wotzka: Aspects de l’occupation humaine et de l’histoire de la végétation au cours de l’Holocène dans la région de la Chaine de Gobnangou, S.E. Burkina Faso. Berichte des Sonderforschungsbereichs 268, 1: 13-31; Frankfurt am Main.

Breunig, P., A. Ballouche, K. Neumann, F.W. Rösing, H. Thiemeyer, P. Wendt & W. van Neer: Gajiganna. New data on early settlement and environment in the Chad basin. Berichte des Sonderforschungsbereichs 268, 2: 51-74; Frankfurt am Main.

Neumann, K.: Holocene vegetation of the Eastern Sahara: charcoal from prehistoric sites. In: Krzyzaniak, L., M. Kobusiewicz & J. Alexander (eds.): Environmental change and human culture in the Nile basin and Northern Africa until the second millenium B.C.: 153-169. Studies in African Archaeology 4, Poznan (Poznan Archaeological Museum).


* Neumann, K.: The contribution of anthracology to the study of the late Quaternary vegetation history of the Mediterranean region and Africa. Bulletin de la Societé botanique de France139, Actualités botaniques 2/3/4: 421-440

* Neumann, K.: Une flore soudanienne au Sahara central vers 7000 B.P.: les charbons de bois de Fachi/Niger. Bulletin de la Societé botanique de France 139, Actualités botaniques 2/3/4: 565-570.

Neumann, K. & A. Ballouche: Die Chaine de Gobnangou in SE Burkina Faso – ein Beitrag zur Vegetationsgeschichte der Sudanzone W-Afrikas. Geobot. Kolloq. 8: 53-68.

Vermeersch, P.M., E. Paulissen, D. Huyge, K. Neumann, W. Van Neer & P. Van Peer: Predynastic hearths in Upper Egypt. In: Friedman, R. & B. Adams (eds.): The Followers of Horus. Studies dedicated to M. A. Hoffman: 163-172; Egyptian Studies Assoc. Publ. 2. Oxbow Monograph 20.


* Neumann, K.: In search for the green Sahara: Palynology and botanical macroremains. Palaeoecology of Africa 22: 203-212.


* Neumann, K.: Holocene vegetation of the Eastern Sahara: charcoal from prehistoric sites. Afr. Arch. Rev. 7: 97-116.

Neumann, K.: Vegetationsgeschichte der Ostsahara im Holozän. Holzkohlen aus prähistorischen Fundstellen. In: Kuper, R. (ed.): Forschungen zur Umweltgeschichte der Ostsahara: 13-181; Köln (Heinrich-Barth-Institut).


Neumann, K.: Die Bedeutung von Holzkohleuntersuchungen für die Vegetationsgeschichte der Sahara: Das Beispiel Fachi/Niger. Würzburger Geogr. Arb. 69: 71-85.


* Neumann, K.: Middle Holocene vegetation of the Gilf Kebir/SW-Egypt – a reconstruction. Palaeoecol. Africa 18: 179-188.

* Neumann, K.: The Jebel Tagern: A contribution to the flora of the Southern Libyan Desert. J. Arid Environments 12: 27-39.

Neumann, K. & E. Schulz: Végétation holocène dans le Sahara Central. Géodynamique 2(2): 150-153; Bondy (O.R.S.T.O.M.).

* Neumann, K. & E. Schulz: Middle Holocene savanna vegetation in the Central Sahara – preliminary report. Palaeoecology of Africa 18: 163-166.


Neumann, K.: Untersuchung der Holzkohlen. In: E. Cziesla: Bericht über die Grabungen 1980 und 1983 in der Weidentalhöhle bei Wilgartswiesen, Pfälzer Wald. Mitt. Hist. Vereins Pfalz 84: 48-51.

Stephanie Bodin
Phd. Stephanie Bodin

‘Plant environment and resource exploitation in SW Ethiopia >50 ka to the present’

I am currently working on wood charcoal in the framework of a multidisciplinary research project focussing on plant environment and resource exploitation in SW Ethiopia, >50 ka to the present. The aim is to reconstruct the paleoenvironment and characterize human impact on the landscape. I am also working on a database that list the woody species of the Dry Afromontane of Ethiopia, their wood anatomical features and includes an electronic identification key. PIs: Katharina Neumann and Karen Hahn.


2020 | Anthracologist at CNRS, Montpellier, France

I identified wood charcoal from Holocene to Pleistocene assemblages from Kourou and Apatou, French Guiana, in the frame of the SOURCE multidisciplinary research project. The aim was to help reconstructing the paleoenvironment. PI: Pierre-Olivier Antoine.

2020 | Anthracologist for the Arkaeos Association, Montpellier, France

I identified wood charcoal from a 19th century lime kiln of the Sucrerie plantation, Martinique. The aim was to better understand wood management and wood economy by the lime kiln workers from this period. Archaeological excavations led by Christophe Vaschalde and David Ollivier.

2019 | PhD at University of Montpellier

My thesis subject was the ‘Characterization of the impact of pre-Columbian anthropogenic fires on the forest vegetation of the Nouragues Natural Reserve in French Guiana using an anthracological approach’. I identified wood charcoal from anthropogenic sites and curated the wood charcoal collection from French Guiana which is at the Institut des Sciences de l’Evolution, Montpellier. I was supervised by Laurent Bremond and Jean-François Molino. My PhD was funded by the Center for the study of biodiversity in Amazonia (LabEx CEBA).

Academic background

2019 | University of Montpellier
PhD in Ecology, Evolution, Genetics, Palaeobiology.
Topic: ‘Characterization of the impact of pre-Columbian anthropogenic fires on the forest vegetation of the Nouragues Natural Reserve in French Guiana using an anthracological approach’. Supervisors: Laurent Bremond and Jean-François Molino.
Funded by the Center for the study of biodiversity in Amazonia (LabEx CEBA).

2015 | University of Montpellier & Montpellier SupAgro
M. Sc. in Ecosystem Ecology.

2013 | University of Montpellier
M. Sc. (1st year) in Evolutionary Biology and Ecology.

2012 | University of Poitiers
M. Sc. (1st year) in Paleontology, Palaeobiology, Phylogeny.

2011 | University of Montpellier
B. Sc. in Earth and Life Sciences.

Articles in peer-reviewed journals

Under review | Antoine P.-O., Wieringa L., Adnet S., Aguilera O., Bodin S. C., et al. Hyperdiverse Pleistocene coastal ecosystems of the Guianas at the Ariane 6 ELA4 launchpad, Kourou, French Guiana. PNAS.

Under review | Bodin S. C., Neumann K., Hensel E. A., Vogelsang R., Demissew S., Casas-Gallego M., Hahn K. Afromontane forests and human impact after the African Humid Period: wood charcoal from the Sodicho rock shelter, SW Ethiopian highlands. Vegetation History and Archaeobotany.  

2023 | Casas-Gallego M., Hahn K., Neumann K., Demissew S., Schmidt M., Bodin S. C., Bruch A. A. Cooling-induced expansions of Afromontane forests in the Horn of Africa since the Last Glacial Maximum. Scientific Reports, 13, 10323. https://doi.org/10.1038/s41598-023-37135-8  

2021 | Bodin S. C., Feiss T., Molino J.-F. & Bremond L. Assessment of the efficiency of three sampling methods for the recovery of soil charcoals in tropical anthropogenic sites. Quaternary International, 595, 145-154. https://doi.org/10.1016/j.quaint.2021.04.023  

2021 | Bodin S. C., Vaschalde C. & Ollivier D. First insights into the wood management for the production of lime given by the anthracological study of a 19th century lime kiln from Martinique, Lesser Antilles. Journal of Archaeological Science: Reports, 37, 102926. https://doi.org/10.1016/j.jasrep.2021.102926

 2020 | Bodin S. C., Molino J. F., Odonne G. & Bremond L. Unraveling pre-Columbian occupation patterns in the tropical forest of French Guiana through an anthracological approach. Vegetation history and Archaeobotany, 1-14. https://doi.org/10.1007/s00334-019-00767-w  

2019 | Bodin S. C., Scheel-Ybert, R., Beauchêne, J., Molino, J. F. & Bremond, L. CharKey: An electronic identification key for wood charcoals of French Guiana. IAWA Journal, 40(1), 75-S20. https://doi.org/10.1163/22941932-40190227  

2017 | Bremond L., Bodin S. C., Bentaleb I., Favier C. & Canal S. Past tree cover of the Congo Basin recovered by phytoliths and δ13C along soil profiles. Quaternary international, 434, 91-101. https://doi.org/10.1016/j.quaint.2015.11.150  

Book chapter

2020 | Bodin S. C., Morin-Rivat J., Bremond L., Scheel-Ybert R., Tardy C. & Vaschalde C. Anthracology in the tropics: how wood charcoals help us to better understand today ecosystems. In: G. Odonne and J.-F. Molino (eds.). Methods in Historical Ecology – Insights from the Neotropics. Routledge. https://www.routledge.com/Methods-in-Historical-Ecology-Insights-from-Amazonia/Odonne-Molino/p/book/9780367182212  

Professional article

2020 | Antoine P.-O., Heuret A., Bodin S., Boivin M., Pujos F. & Marivaux L. Un inventaire fossile de la Guyane : historique et nouvelles perspectives. Géologues, 206, 27-29.  

Articles for the general public

2021 | Bodin S., Bremond L., Mestre M., Lévêque F. Roy M., Molino J.-F., Odonne G. et al. LongTIme. Ces mondes disparus que cache la forêt. Une Saison en Guyane, Hors-Série n°7 Archéologie.  

2021 | Heuret, A., Bodin, S., Marivaux, L. & Antoine, P.-O. Des étoiles aux fossiles. Quand Ariane 6 dévoile la Guyane pléistocène. Une Saison en Guyane, Hors-Série n°7 Archéologie.

Jennifer Markwirth
Technische Assistentin
Project staff

Post-doctoral fellows

Alessandro Urciuoli
Dr. Alessandro Urciuoli
Post-doctoral researcher

Ph.D. in Biological Anthropology and Biodiversity, Alessandro Urciuoli is a Margarita Salas Postdoctoral Fellow at the Autonomous University of Barcelona (Spain). He is also visiting researcher at the Paleoanthropology Division of the Senckenberg Research Institute and Natural History Museum Frankfurt, as well as associate researcher at the Catalan Paleontological Institute Miquel Crusafont (Spain). His main research line deals with the evolution of the inner ear anatomy in catarrhines primates, with particular emphasis on that of hominins and of the ape-and-human clade. Author of several papers in JCR journals of high impact (e.g., Proceedings of the National Academy of Sciences USA, eLife, Journal of Human Evolution, and American Journal of Biological Anthropology), he aims to clarify the evolutionary history of early Homo species (Homo ergaster and Homo erectus) in Africa and Asia through the inspection semicircular canal shape with different geometric morphometrics methods. Most recently, he has been dedicating efforts to perform morphofunctional analyses of the middle ear and the cochlea to reconstruct the hearing capabilities of extinct hominins, and provide insights into the evolution of language within the human clade.

 31/12/2021/ – 31/12/2024

Margarita Salas Fellowship
Universitat Autònoma de Barcelona

  01/12/2015 – 16/04/2021

Ph.D in Biological Anthropology
Institut Català de Paleontologia – Campus UAB, Barcelona, Spain
The evolution of semicircular canals in anthropoid primates: Phylogenetic implications for Miocene catarrhines
Final note: Excellent – Cum Laude – International Doctor Mention

  01/04/2015 – 01/07/2015

Erasmus + Traineeship LLPInstitut Català de Paleontologia – Campus UAB, Barcelona, Spain
Analysis of artiodactyl remains, learning of 3D geometric morphometrics techniques and use of micro CT scanning.

  26/09/2012 – 30/09/2014

Master Degree: Scienze della Natura e dell’Uomo. Nivel QEQ: 7LM60
Unversity of Florence – Facoltà Scienze MM.FF.NN.
Graduation mark: 110/110 – Cum Laude 

Urciuoli, A., Alba, D.M., 2023. Systematics of Miocene apes: State of the art of a neverending controversy. Hum. Evol. 175,103309.

Arias-Martorell, J., Urciuoli, A., Almécija, S., Alba, D.M., Nakatsukasa, M., 2023. The proximal radial morphology the Middle Miocene ape Nacholapithecus kerioi: Morphometric affinities, locomotor inferences, and implications for the evolution of the hominoid humeroradial joint. Hum. Evol. 178, 103345.

Frosali, S., Bartolini-Lucenti, S., Madurell-Malapeira, J., Urciuoli, A., Costeur, L., Rook, L., in press. First digital study of the frontal sinus of stem-Canini (Canidae, Carnivora): evolutionary and ecological insights. Ecol. Evol. (in press).

Urciuoli, A., Kubat, J., Schisanowski, L., Schrenk, F., Zipfel, B., Tawane, M., Bam, L., Alba, D.M., Kullmer, O., 2022. Cochlear morphology of Indonesian Homo erectus from Sangiran. Hum. Evol. 163, 103163.

Urciuoli, A., Zanolli, C., Bouchet, F., Almécija, S., Moyà-Solà, S., Alba, D.M., 2022. Semicircular canal morphology of the Miocene small-bodied catarrhine Pliobates cataloniae: Phylogenetic implications. J. Biol. Anthropol. 177(S73), 187.

Urciuoli, A., Florian, B., Quirós-Sánchez, A., Conde-Valverde, M., 2022. The human cochlea: a test case for endocranial morphology comparisons on a small scale. PaleoAnthropology 2,

Urciuoli, A., Beaudet, A., Heaton, J., Alba, D.M., 2022. Papionin affinities in the endocranial anatomy of Cercopithecoides williamsi.In: Belvedere M., Mecozzi B., Amore O., Sardella R (Eds.). Abstract book of the XIX Annual Conference of the European Association of Vertebrate Palaeontologists, Benevento/Pietraroja, Italy, 27th June-2nd July 2022. PalaeoVertebrata, Special Volume 12022, 207.

López-Onaindia, D., Crevecoeur, I., Thibeault, A., Urciuoli, A., Lucas-Aragay, M., Pruvost, M., Zanolli, C., Sala, N., Balzeau, A., Couture-Veschambre, C., Gómez-Olivencia, A., 2022. Variation of bony labyrinth morphology in late Middle Pleistocene Neanderthals from Abri Suard, France. J. Biol. Anthropol. 177(S73), 111.

Zanolli, C., Urciuoli, A., Delgado, M., Davies, T.W., Bouchet, F., Fortuny, J., Beaudet, A., Alba, D.M., Kullmer, O., Skinner, M.M., 2022. The phylogenetic signal of the enamel-dentine junction of primate molars. J. Biol. Anthropol. 177(S73), 202–203.

Fornai, C., Webb, N.M., Urciuoli, A., Krenn, V.A., Corron, L.K., Haeusler, M., 2021. New insights on hip bone sexual dimorphism in adolescents and adults using deformation-based geometric morphometrics. Anthropol. Sci. 99, 117-134.

Bouchet, F., Urciuoli, A., Beaudet, A., Pina, M., Moyà-Solà, S., Alba, D.M., 2021. Comparative anatomy of the carotid canal in the Miocene small-bodied catarrhine Pliobates cataloniae.   Hum. Evol. 161, 103073.

Arias-Martorell, J., Almécija, S., Urciuoli, A., Nakatsukasa, M., Moyà-Solà, S., Alba, D.M. 2021. A proximal radius of Barberapithecus huerzeleri from Castell de Barberà: Implications for locomotor diversity among pliopithecoids. Hum. Evol. 157, 103032.

Fornai, C., Urciuoli, A., Kren, A.V., Hausler, M., Webb, N.M., 2021. Deformation-based geometric morpho-metrics captures the morphology of the modern human hipbone. J. Phys. Anthropol. 174 (S71), 33.

Urciuoli, A., Zanolli, C., Almécija, S., Beaudet, A., Dumoncel, J., Morimoto, N., Nakatsukasa, M., Moyà-Solà, S., Begun, D. R., Alba, D. M., 2021. Reassessment of the phylogenetic relationships of the late Miocene apes Hispanopithecus and Rudapithecusbased on vestibular morphology. Natl. Acad. Sci. USA 118, e2015215118.

Urciuoli, A., Zanolli, C., Beaudet, A., Pina, M., Almécija, S., Moyà-Solà, S., Alba, D. M., 2021. A comparative analysis of the vestibular apparatus in Epipliopithecus vindobonensis: Phylogenetic implications. Hum. Evol. 151, 102930.

Bouchet, F., Urciuoli, A., Beaudet, A., Moyà-Solà, S., Alba, D.M., 2020. Orientation and proportions of the carotid canal in Epipliopithecus and Pliobates. ESHE 9, 18.

Urciuoli A., Zanolli C., Beaudet A., Dumoncel J., Santos F., Moyà-Solà S., Alba D.M. (2020). The evolution of the vestibular apparatus in apes and humans. eLife 9, e51261.

Urciuoli A., Zanolli C., Begun D., Almécija S., Dumoncel J., Moyà-Solà S., Alba, D.M. (2019). A deformation-based geometric morphometric analysis of the vestibular apparatus in the Miocene apes Hispanopithecus laietanus and Rudapithecus hungaricus. J. Phys. Anthropol. 168 (S68), 253.

Urciuoli A., Zanolli C., Fortuny J., Almécija S., Schillinger B., Moyà-Solà S., Alba D.M. (2018). Neutron-Based Computed Microtomography: Pliobates cataloniae and Barberapithecus huerzeleri as a Test-Case Study. J. Phys. Anthropol. 166, 987–993.

Urciuoli A., Pina M., Moya-Sola S., Alba D.M. (2018) The inner ear of Epipliopithecus vindobonensis: preliminary results. J. Phys. Anthropol. 165 (S66), 281

Urciuoli A., Zanolli C., Almécija S., Moyà-Solà S., Alba D.M. (2018). Analysis of the primate vestibular apparatus: a comparison of 3D geometric morphometric and diffeomorphism approaches. ESHE 7, 193.

Delfino M., Candilio F., Carnevale G., Coppa A., Medin T., Pavia M., Rook L., Urciuoli A., Villa A. (2018) The early Pleistocene vertebrate fauna of Mulhuli-Amo (Buia area, Danakil Depression, Eritrea). Soc. Paleontol. Ital. 57, 27–44.

Urciuoli A., DeMiguel D., Moyà-Solà S., Rook L. (2016). New Hoplitomeryx Leinders, 1984 remains from the Late Miocene of Gargano (Apulia, Italy). Hystrix 27, 1–8.

Riczar Fuentes
Dr. Riczar Belcena Fuentes
Post-doctoral researcher

Dr. Riczar Fuentes awarded with the Southeast-Asia Biocultural Evolution Research Fund, promoted by the Senckenberg Gesellschaft für Naturforschung, the Daimler and Benz Foundation and the Werner Reimers Foundation

Research interests

Prehistoric archaeology, human evolution, prehistoric migration and island adaptation, stone tool technology, lithic use-wear analysis, Southeast Asian prehistory.

Academic positions:


8/2021 – present

Assistant Professor, Department of Sociology and Anthropology, Ateneo de Manila University, Quezon City, Philippines

01/2023 – 12/2024

Post-doctoral researcher, Department of Palaeoanthropology, Senckenberg Research Institute and Natural History Museum Frankfurt, Frankfurt am Main, Germany

08/2020 – 07/2021

Lecturer, Department of Sociology and Anthropology, Ateneo de Manila University, Quezon City, Philippines

10/2016 – 09/2019

Ph.D. scholar, Lisa Maskell Fellowships – Gerda Henkel Foundation, Düsseldorf, Germany, Project titled Island Adaptation and Maritime Interaction in Changing Environments from the Terminal Pleistocene to the Early Holocene: A Comparative Study of Prehistoric Technology in Northeastern Indonesia

08/2015 – 05/2016

Senior lecturer, Arkiyoloji 1 (Ang Pilipinas: Arkiyoloji at Kasaysayan, The Philippines: Archaeology and History), Archaeological Studies Program (ASP), University of the Philippines Diliman

10/2012 – 09/2016

Research associate/ laboratory technician, Emerging Interdisciplinary Research (EIDR) Grant – “Palaeoenvironmental and Biodiversity Study of Mindoro Island: An Archaeological Science Initiative”, ASP, UP Diliman

07/2012 – 06/2013

Research associate, under Dr. Alfred F. Pawlik, Office of the Vice-Chancellor for Research and Development (OVCRD) Outright Research Grant, Project No. 121206 PNSE titled “Microwear and Residue Analysis of Lithic Artefacts from Vito Cave, Peñablanca, Cagayan and the Reconstruction of Prehistoric Technology, Cave Function and Site Activities”, ASP, UP Diliman



Researcher, Mindoro Archaeological Project
A joint project of UP Diliman and The University of Western Australia ASP, UP Diliman



10/2016 – 03/2020 Dr. rer. nat. (Ph.D.) in Archaeological Sciences (magna cum laude) Eberhard Karls Universität Tübingen, 72070 Tübingen, Germany 06/2009 – 06/2015 Master of Arts in Archaeology University of the Philippines Diliman, Quezon City, 1101 Philippines 06/2003 – 04/2008 Bachelor of Arts (History) University of the Philippines Visayas, Miagao, Iloilo, 5023 Philippines





Ono, R., Sofian, H.O., Fuentes, R., Aziz, N. and Pawlik, A., 2023. The Goa Topogaro complex: Human migration and mortuary practice in Sulawesi during the Late Pleistocene and Holocene. L’Anthropologie, p.103155.

Pawlik, A. and Fuentes, R. 2023. Prehistoric Hunter-Gatherers in the Philippines—Subsistence strategies, adaptation, and behaviour in maritime environments. Frontiers in Earth Science, 11, p.574.

Ono, R., Sofian, H.O., Fuentes, R., Aziz, N., Ririmasse, M., Geria, I.M., Katagiri, C. and Pawlik, A., 2023. Early modern human migration into Sulawesi and Island adaptation in Wallacea. World Archaeology, 54(2), pp.229-243.

Pawlik, A. and Fuentes, R., 2023. Traceology. Reference Module in Social Sciences, Elsevier.

Fuentes, R. and Pawlik, A., 2023. Barely scratched the surface: Development and future directions of lithic use-wear analysis in Island Southeast Asia. Archaeological Research in Asia, 33, p.100413.

Ono, R., Fuentes, R., Nakatani, A., Kanetani, M., and Ueba, Y., 2022. “Bamboo Hypothesis” Revisited: Functions of Prehistoric Stone Tools and Plant Use in Wallacea (in Japanese). Bulletin of the National Museum of Ethnology 46(3): 1–124. https://doi.org/10.15021/00009890

Fuentes, R., Ono, R., Aziz, N., Alamsyah, N., Sofian, H.O., Miranda, T. and Pawlik, A., 2021. Inferring human activities from the Late Pleistocene to Holocene in Topogaro 2, Central Sulawesi through use-wear analysis. Journal of Archaeological Science: Reports, 37, p.102905 https://doi.org/10.1016/j.jasrep.2021.102905

Ono, R., Fuentes, R., Amano, N., Sofian, H.O., Aziz, N. and Pawlik, A., 2021. Development of bone
and lithic technologies by anatomically modern humans during the late Pleistocene to Holocene in
Sulawesi and Wallacea. Quaternary International. https://doi.org/10.1016/j.quaint.2020.12.045

Ono, R., Pawlik, A. and Fuentes, R., 2020. Island Migration, Resource Use, and Lithic Technology by Anatomically Modern Humans in Wallacea. In Pleistocene Archaeology-Migration, Technology, and Adaptation. IntechOpen. DOI: 10.5772/intechopen.93819

Ono, R., Fuentes, R., Pawlik, A., Sofian, H.O., Aziz, N., Alamsyah, N. and Yoneda, M., 2020. Island migration and foraging behaviour by anatomically modern humans during the late Pleistocene to Holocene in Wallacea: New evidence from Central Sulawesi, Indonesia. Quaternary International. https://doi.org/10.1016/j.quaint.2020.03.054

Fuentes, R., Ono, R., Carlos, J., Kerfant, C., Miranda, T., Aziz, N., Sofian, H.O. and Pawlik, A., 2020. Stuck within notches: Direct evidence of plant processing during the last glacial maximum to Holocene in North Sulawesi. Journal of Archaeological Science: Reports, 30, p.102207. https://doi.org/10.1016/j.jasrep.2020.102207

Fuentes, R., and Pawlik, A., 2020. Not formal but functional: Traceology and the Lithic Record in the Philippines. In Gibaja-Bao, J. Marreiros, J., & Mazzucco, N (Eds.) Hunter-Gatherers tool kit: a functional perspective (pp. 290-308). Cambridge Scholars Publishing.

Fuentes, R., Ono R., Nakajima, N., Nishizawa, H., Siswanto, J., Aziz, N., Sriwigati, Sofian, H.O., Miranda, T., and Pawlik, A.F., 2019. Technological and behavioural complexity in expedient industries: the importance of use-wear analysis for understanding flake assemblages. Journal of Archaeological Science, 112, p. 105031. https://doi.org/10.1016/j.jas.2019.105031

Pawlik, A., Crozier, R., Fuentes, R., Wood, R. and Piper, P., 2019. Early to Mid-Holocene Burial Traditions of Island Southeast Asia and a 5th millennium BP flexed inhumation from Bubog-1, Ilin Island, Mindoro Occidental. Antiquity. Volume 93, Issue 370, pp. 901-918. https://doi.org/10.15184/aqy.2018.190