Evolving Earth and Environment

The Research Activity “Evolving Earth and Environment” reconstructs how the earth’s evolutionary history is reflected in geological and mineralogical processes in the earth’s crust, thereby reinforcing the geobiodiversity approach of the entire Senckenberg research. Different topics from the geosciences, climate and biodiversity research are integrated here, enabling us to understand how the evolution of organisms and their ecosystem structure is linked to geological processes on various spatial-temporal scales. Special focus is given to evolutionary key events (e.g., mass extinctions, followed by recovery phases). From these, we can derive what past processes played a role in mass extinction events, and what we can learn from this in regard to the current, Anthropocene mass extinction. The work of this RA greatly benefits from the outstanding fossils of the Messel Pit, which are renowned for their globally unique level of preservation.


Prof. Dr.  André Freiwald
Director Senckenberg am Meer, Head of Department Marine Research, Head of Department Marine Geology

Functions within Senckenberg:

  • Head of Division Marine Geology (SaM)
  • Speaker of RA 4.1: Evolving Earth and Environment

Research profile

  • Non-tropical biosedimentary systems
  • Cold-water coral ecosystems
  • Bioerosion
  • Paleoecology

Research interests

My research focuses on geobiological studies of recent and fossil cold-water corals, with a focus on occurrences along the increasingly oxygen-poor upwelling areas of West Africa (Mauritania and Angola-Namibia). Another topic is research on the quantitative degradation of carbonate by biological processes (bioerosion). Finally, we are developing a global database on habitat-forming cold-water coral and associated communities across all phyla. In this database, the current state of knowledge of the composition of the reef inhabitants from the unicellular organisms to the fish, their geographical changes and their biological functions will be depicted for the first time.


Professional positions
2018-2022 Director Senckenberg am Meer, Wilhelmshaven
Since 2010 Head of Department  Marine Research at Senckenberg am Meer
Since 2010 Professor, University of Bremen
2002-2010 Professor, Friedrich-Alexander-Universität Erlangen-Nürnberg
2000-2002 Professor, University of Tübingen

2000 Heisenberg Fellow
1999 Habilitation University of Bremen
1993-1999 Researcher at University of Bremen
1993 PhD, Kiel University

Relevant scientific projects  2017-2023
2018-2019 Atlantic Thermocline Ocean and Ecosystems Dynamic during Natural Climate Change (ATHENA) (DFG) (externally administered through University of Heidelberg)
2019-2022 Cold-water Coral sensitivity (MARUM Excellence Cluster)
2019-2022 West-African Biodiversity under Pressure (WASP) (GIZ)
2022-2024 RestoreSeas: Restoration of Marine Forests (Cold-water corals, seagrass and coralline algae) (JPI OCEANS -BioDIVERSA)

Professional service, membership associations, cooperation in national and intl. committees
Since 2023 Member of the International Scientific Council of IMROP (Institut Mauritanien de Recherches Océanographiques et des Pêches, Nouadhibou, Mauritania)
Since 2023 Member of the International Scientific Council of the PNBA (Parc National Banc d’Arguin, Mauritania)
Since 2017 Advisory Board of the Northwest German University Society
Since 2015 -2022 Chairman of the DFG-BMBF Advisory Councils of the research vessels METEOR
2019 Foundation member of the Deutsche Allianz Meeresforschung
2015-2022 Member of the BMBF advisory board research vessel SONNE
2011-2015 Supervisory Board at the Center for Tropical Marine Ecology
Since 2010 Member of the MARUM board of the University of Bremen
​2008-2011 Speaker of DFG Geoforum
2004-2008 Member of the DFG Review Board Geology-Paleontology
2001-2006 Member of the DFG Senate Commission for Oceanography
2000-2003 Coordinator of the EU-FP5 Atlantic Coral Ecosystem Study (ACES) project

Editorial activities
2005-2010 Publisher of the Springer book series “Erlangen Earth Conference Series”
2004-2010 Publisher of the ISI journal FACIES together with Springer

Awards and awarded memberships
2016 Gustav-Steinmann-Medal of the Geological Association
2015 Northern German Science Award for innovative research on cold-water corals



Bridges AEH, Howell KL, Amaro T, Atkinson L, Barnes DKA, Bax N, Bell JB, Bernardino AF, Beuck L, Braga-Henriques A, Brandt A, Bravo ME, Brix S, Butt S, Carranza A, Doti BL, Elegbede IO, Esquete P, Freiwald A, Gaurdron SM, Guilhon M, Hebbeln D, Horton T, Kainge P, Kaiser S, Lauretta D, Limongi P, Mcquaid KA, Milligan RJ, Milosavich P, Narayanaswamy BE, Orejas C, Paulus S, Pearman TRR, Perez JAA, Ross RE, Saeedi H, Shimabukuro M, Sink K, Stevenson A, Taylor M, Titschack J, Vieira RP, Vinha B, Wienberg C (2023) Review of the Central and South Atlantic shelf and deep-sea benthos: Science, policy, and management. Oceanography and Marine Biology: An Annual Review 61:127-218

Caballero Herrera JA, Hoffman L, Freiwald A, Gofas S (2023) The dispersal capacity of Mollusca—a test on the South Azorean Seamount Chain. Marine Biodiversity 53:59

Gofas S, Freiwald A, Hoffman L (2023) New species and new records in Cerithiopsidae and Newtoniellidae (Triphoroidea, Gastropoda) from the South Azorean Seamount Chain Iberus 41:111-148

Hoffman L, Freiwald A (2023) Species in Fissurellidae (Gastropoda) from the North Atlantic with a focus on the Azorean seamounts. Basteria 87:77-96

Knorrn AH, Beuck L, Barros-García, Fernández-Peralta, Freiwald A (accepted) Gaidropsarus mauritanicus (Gadiformes, Gaidropsaridae) a new three-bearded rockling species from a deep-water coral ecosystem with a genetically verified biogeographical distribution of the genus. Spixiana

Wienberg C, Freiwald A, Frank N, Mienis F, Titschack J, Orejas C, Hebbeln D (accepted) Cold-water coral reefs in the oxygen minimum zones off West Africa. In: Cordes EE, Mienis F (eds) Coral Reefs of the World, Springer

Vinha B, Rossi S, Gori A, Hanz U, Pennetta A, De Benedetto GE, Mienis F, Huvenne VAI, Hebbeln D, Wienberg C, Titschack J, Freiwald A, Piraino S, Orejas C (2023) Trophic ecology of Angolan cold‑water coral reefs (SE Atlantic) based on stable isotope analyses. Scientific Reports 13:9933



Freiwald A (2022) Leon der Profi. Natur, Forschung und Museum 152:86-89

Herrán N, Narayan GR, Doo SS, Klicpera A, Freiwald A, Westphal H (2022) High-resolution imaging sheds new light on a multi-tier symbiotic partnership between a “walking” solitary coral, a sipunculan, and a bivalve from East Africa. Ecology and Evolution 12:e8633

Hesemann M, Hoffman L, Ottway B, Freiwald A (2022) Benthic foraminiferal assemblages from the Mauritanian shelf and upper slope and their association with cold-water coral habitats. Natur im Fokus 53:21-153

Hoffman L, Freiwald A (2022) Anatoma occidentalis n. sp. (Vetigastropoda, Anatomidae) from upper bathyal coral habitats off western Africa. Iberus 40:355-362

Hoffman L, Freiwald A (2022) A review of Atlantic deep-water species in the genus Talassia (Caenogastropoda, Vanikoridae). European Journal of Taxonomy 819:140-157

Meyer N, Wisshak M, Edinger EN, Azetsu-Scott K, Freiwald A (2022) Ichnodiversity in the eastern Canadian Arctic in the context of polar microbioerosion patterns. Polar Research 41:8083

Sampaio I, Beuck L, Freiwald A (2022) A new octocoral species of Swiftia (Holaxonia, Plexauridae) from the upper bathyal off Mauritania (NE Atlantic). ZooKeys 1106:121-140

Sampaio I, Beuck L, Menezes GM, Freiwald A (2022) The Mauritanian slope (NE Atlantic) has no desert: Swiftia phaeton (Holaxonia: Plexauridae) shaping coral gardens. In: Chimienti G (ed) Corals – Habitat formers from the shallow to the deep. InTech Open, pp 1-27

Wisshak M, Meyer N, Kuklinski P, Rüggeberg A, Freiwald A (2022) ‘Ten Years After’—a long-term settlement and bioerosion experiment in an Arctic rhodolith bed (Mosselbukta, Svalbard). Geobiology 20:112-136

Zettler ML, Hendrycks EA, Freiwald A (2022) A new amphipod species of the bathyal genus Dautzenbergia Chevreux, 1900 (Amphipoda, Calliopioidea, Pontogeneiidae) associated with cold-water corals off Angola. Zootaxa 5213:49-63



Freiwald A (2021) Wettlauf mit der Zeit. Natur, Forschung und Museum 151:126-131

Gil M, Freiwald A, Ramil F (2021) Two new species of Rosalinda (Cnidaria, Hydrozoa, Anthoathecata) from West African cold-water coral mounds. Deep-Sea Research 168:103436

Hoffman L, Freiwald A (2021) Bathyal species in Rissoidae (Gastropoda) from Azorean seamounts. Biodiversity Journal 12:777-792

Hoffman L, Gofas S, Freiwald A (2021) The genus Anatoma Woodward, 1859 (Gastropoda, Anatomidae) from Azorean seamounts. Iberus 39:139-152

Meyer N, Wisshak M, Freiwald A (2021) Bioerosion ichnodiversity in barnacles from the Ross Sea, Antarctica. Polar Biology 44:667-682

Orejas C, Wienberg C, Titschack J, Tamborrino L, Freiwald A, Hebbeln D (2021) Madrepora oculata forms large frameworks in hypoxic waters off Angola (SE Atlantic). Scientific Reports 11:15170

Petersen J, Titschack H, Groeneveld J, Wehrmann A, Hebbeln D, Freiwald A (2021) Reef-building Pacific oysters record seasonal variations in water mass-properties of tidal basins from the Central Wadden Sea (North Sea). Palaeogeography, Palaeoclimatology, Palaeoecology 577:110534

Sanna G, Freiwald A (2021) Deciphering the composite morphological diversity of Lophelia pertusa, a cosmopolitan deep-water ecosystem engineer. Ecosphere 12:e03802

Schätzle PK, Wisshak M, Bick A, Freiwald A, Kieneke A (2021) Exploring confocal laser scanning microscopy (CLSM) and fluorescence staining as a tool for imaging and quantifying traces of marine microbioerosion and their trace-making microendoliths. Journal of Microscopy 284:118-131

Schleinkofer N, Evans D, Wisshak M, Büscher JV, Fiebig J, Freiwald A, Härter S, Marshall HR, Voigt S, Raddatz J (2021) Host-influenced geochemical signature in the parasitic foraminifera Hyrrokkin sarcophaga. Biogeosciences 18:4733-4753



Hebbeln D, Wienberg C, Dullo WC, Freiwald A, Mienis F, Orejas C, Titschack J (2020) Cold-water coral reefs thriving under hypoxia. Coral Reefs 39:853-859

Hoffman L, Freiwald A (2020) Bathyal Eulimidae (Gastropoda: Vanikoroidea) from the Azorean seamounts collected during the R/V Meteor Cruise M151 Athena. Miscellanea Malacologica 8:81-99

Hoffman L, Freiwald A (2020) Cantrainea mauritanea n. sp. (Gastropoda: Colloniidae) from deep-water coral habitats off Mauritania. Miscellanea Malacologica 8:69-73

Hoffman L, Gofas S, Freiwald A (2020) A large biodiversity of “skeneimorph” (Gastropoda: Vetigastropoda) species from the South Azorean Seamount Chain, with the description of seventeen new species. Iberus Supplement 9:1-82

Hoffman L, Gofas S, Freiwald A (2020) New and little-known Seguenziidae (Vetigastropoda, Gastropoda) from the South Azorean Seamount Chain. Iberus 38:1-18

Hoffman L, Gofas S, Freiwald A (2020) Ten new species in Papuliscala de Boury, 1911 (Gastropoda, Epitoniidae) from the South Azorean Seamount Chain. Iberus 38:29-53

Morato T, González-Irusta JM, Dominguez-Carrió C, Wei CL, Davies A, Sweetman AK, Taranto GH, Beazley L, García-Alegre A, Grehan A, Laffargue P, Murillo FJ, Sacau M, Vaz S, Kenchington E, Arnaud-Haond S, Callery O, Chimienti G, Cordes E, Egilsdottir H, Freiwald A, Gasbarro R, Gutiérrez-Zárate C, Gianni M, Gilkinson K, Wareham Hayes VE, Hebbeln D, Hedges K, Henry LA, Johnson D, Koen-Alonso M, Lirette C, Mastrototaro F, Menot L, Molodtsova T, Muñoz PD, Orejas C, Pennino MG, Puerta P, Ragnarsson SA, Ramiro-Sánchez B, Rice J, Rivera J, Roberts JM, Ross SW, Rueda JL, Sampaio Í, Snelgrove P, Stirling D, Treble MA, Urra J, Vad J, Van Oevelen D, Watling L, Walkusz W, Wienberg C, Woillez M, Levin LA, Carreiro-Silva M (2020) Climate-induced changes in the suitable habitat of cold-water corals and commercially important deep-sea fishes in the North Atlantic. Global Change Biology 26:2181-2202

Raddatz J, Titschack J, Frank N, Freiwald A, Conforti A, Osborne A, Skornitzke S, Stiller W, Rüggeberg A, Voigt S, Albuquerque ALS, Vertino A, Schröder-Ritzrau A, Bahr A (2020) Solenosmilia variabilis-bearing cold-water coral mounds off Brazil. Coral Reefs 39:69-83



Freiwald A (2019) Messinian Salinity Crisis: What happened to cold-water corals? In: Orejas C, Jiménez C (eds) Mediterranean Cold-Water Corals: Past, Present and Future. Springer, pp 47-50

George KH, Wehrmann A, Freiwald A (2019) Gipfelstürmer der Tiefe. Natur, Forschung und Museum 149:72-77

Hanz U, Wienberg C, Hebbeln D, Duineveld G, Lavaleye M, Juva K, Dullo WC, Freiwald A, Tamborrino L, Reichart GJ, Flögel S, Mienis F (2019) Environmental factors influencing cold-water coral ecosystems in the oxygen minimum zones on the Angolan and Namibian margins. Biogeosciences 16:4337-4356

Hoffman L, Beuck L, Van Heugten B, Lavaleye M, Freiwald A (2019) Last snails standing since the Pleistocene, a tale of Calliostomatidae (Gastropoda) living in deep-water coral habitats in the Northeast Atlantic. Zootaxa 4613:93-110

Hoffman L, Freiwald A (2019) Neocardia kandeli n. sp. (Bivalvia: Philobryidae) from Namibia. Miscellanea Malacologica 8:55-58

Hoffman L, Freiwald A (2019) Vexillum monscorallum n. sp. (Gastropoda: Costellariidae) from deep-water coral habitats off Mauritania. Miscellanea Malacologica 8:37-40

Hoffman L, Monsecour K, Freiwald A (2019) Columbellidae (Gastropoda) from deep-water coral habitats off Mauritania. Miscellanea Malacologica 8:41-49

Hoffman L, Von Cosel R, Freiwald A (2019) New and little-known Pliocardiinae (Bivalvia, Vesicomyidae) from the continental slope off Mauritania. Basteria 83:151-157

Sampaio I, Carreiro-Silva M, Freiwald A, Menezes G, Grasshoff M (2019) Natural history collections as a basis for sound biodiversity assessments: Plexauridae (Octocorallia, Holaxonia) of the Naturalis CANCAP and Tyro Mauritania II expeditions. ZooKeys 870:1-32

Sampaio I, Freiwald A, Porteiro FM, Menezes G, Carreiro-Silva M (2019) Census of Octocorallia (Cnidaria: Anthozoa) of the Azores (NE Atlantic) with a nomenclature update. Zootaxa 4550:451-498

Schleinkofer N, Raddatz J, Freiwald A, Evans D, Beuck L, Rüggeberg A, Liebetrau V (2019) Environmental and biological controls on Na/Ca ratios in scleractinian cold-water corals. Biogeosciences 16:3565-3582

Tamborrino L, Wienberg C, Titschack J, Wintersteller P, Mienis F, Schröder-Ritzrau A, Freiwald A, Orejas C, Dullo WC, Haberkern J, Hebbeln D (2019) Mid-Holocene extinction of cold-water corals on the Namibian shelf steered by the Benguela oxygen minimum zone. Geology 47:1185-1188



Hoffman L, Fraussen K, Freiwald A (2018) The genus Chauvetia (Gastropoda, Buccinoidea) in deep-water coral habitats on the continental slope of Mauritania. Miscellanea Malacologica 7:127-138

Hoffman L, Freiwald A (2018) Last snails standing: a tale of Fissurellidae (Gastropoda) from deep-water coral habitats off Mauritania since the Pleistocene. Miscellanea Malacologica 7:115-126

Hoffman L, Freiwald A (2018) A new genus and two new species in Calliotropidae (Mollusca: Gastropoda) from NE Atlantic cold-water coral habitats. Miscellanea Malacologica 7:89-96

Hoffman L, Freiwald A, van Heugten B, Lavaleye M (2018) New information on Ganesa, Lopheliella and Trochaclis (Gastropoda) in cold-water coral habitats in the temperate and tropical North Atlantic  Miscellanea Malacologica 8:27-34

Hoffman L, Lavaleye M, van Heugten B, Freiwald A (2018) Lepetella (Gastropoda: Lepetellidae) in deep-water coral habitats in the North Atlantic. Miscellanea Malacologica 8:19-26

Ramos A, Ramil F, Freiwald A, Beuck L, Moctar M, Bouzouma ME, Khallahi B, Kloff S (2018) Une deuxieme vie pour la zone d’exclusivite du puits Chinguetti – Un réseau d’Aires Marines Protégées sur le talus Mauritanien pour une pêche plus riche et une meilleure maitrise du milieu marin. Présenté par le panel d’experts mandaté par Le Ministère des Pêches et de l’Economie Maritime, Le Ministère de l’Environnement et du Développement Durable et Le Ministère de l’Education nationale

Rosso A, Beuck L, Vertino A, Sanfilippo A, Freiwald A (2018) Cribrilinids (Bryozoa, Cheilostomata) associated with deep-water coral habitats at the Great Bahama Bank slope (NW Atlantic), with description of new taxa. Zootaxa 4524:401-439

Wienberg C, Titschack J, Freiwald A, Frank N, Lundälv T, Taviani M, Beuck L, Schröder-Ritzrau A, Krengel T, Hebbeln D (2018) The giant Mauritanian cold-water coral mound province: Oxygen control on coral mound formation. Quaternary Science Reviews 185:135-152

Zettler ML, Freiwald A, Guerra-García JM (2018) Cold-water corals off Angola as refuge for a new Aeginella species (Crustacea: Amphipoda: Caprellidae). Zootaxa 4462:535-546



Freiwald A, Rogers A, Hall-Spencer J, Guinotte JM, Davies AJ, Yesson C, Martin CS, Weatherdon LV (2017) Global distribution of cold-water corals (version 5.0). Fifth update to the dataset in Freiwald et al. (2004) by UNEP-WCMC, in collaboration with Andre Freiwald and John Guinotte. . URL: http://data.unep-wcmc.org/datasets/3 edn., Cambridge, UK

Hoffman L, Freiwald A (2017) A unique and diverse amalgamated mollusk shell assemblage from the Coral Patch Seamount, eastern Atlantic. Miscellanea Malacologica 7:55-73

Milker Y, Weinkauf MFG, Titschack J, Freiwald A, Krüger S, Jorissen FJ, Schmiedl G (2017) Testing the applicability of a benthic foraminiferal-based transfer function for the reconstruction of paleowater depth changes in Rhodes (Greece) during the early Pleistocene. PlosOne 12:e0188447



Beuck L, Aguilar R, Fabri M, Freiwald A, Gofas S, Hebbeln D, López Correa M, Ramos Matos A, Ramil F, Sánchez Delgado F, Taviani M, Wienberg C, Wisshak M, Zibrowius H (2016) Biotope characterisation and compiled geographical distribution of the deepwater oyster Neopycnodonte zibrowii in the Atlantic Ocean and Mediterranean Sea. Rapport Commission International Mer Méditerranée 41:462

Cordes EE, Arnaud-Haond S, Bergstad OA, Da Costa Falcao AP, Freiwald A, Roberts JM, Bernal P (2016) Cold-water corals. In: Inniss L, Simcock A, Ajawin AY et al. (eds) World Ocean Assessment I. United Nations, pp 1-28

Färber C, Titschack J, Schönberg CHL, Ehrig K, Boos K, Baum D, Illerhaus B, Asgaard U, Bromley RG, Freiwald A, Wisshak M (2016) Long-term macrobioerosion in the Mediterranean Sea assessed by micro-computed tomography. Biogeosciences 13:3461-3474

Göcke C, Hestetun JT, Uhlir C, Freiwald A, Beuck L, Janussen D (2016) Cladorhiza corallophila sp. nov., a new carnivorous sponge (Cladorhizidae, Demospongiae) living in close association with Lophelia pertusa and Madrepora oculata (Scleractinia). Zootaxa 4168:512-524

Macpherson E, Beuck L, Freiwald A (2016) Some species of Munidopsis from the Gulf of Mexico, Florida Straits and Caribbean Sea (Decapoda: Munidopsidae), with the description of two new species. Zootaxa 4137:405-416

Titschack J, Fink HG, Baum D, Wienberg C, Hebbeln D, Freiwald A (2016) Mediterranean cold-water corals – an important regional carbonate factory? Depositional Record 2:74-96

Welte C, Wacker L, Hattendorf B, Christl M, Fohlmeister J, Breitenbach SFM, Robinson LF, Andrews AH, Freiwald A, Farmer JR, Yeman C, Synal HA, Günther D (2016) Laser ablation − accelerator mass spectrometry: An approach for rapid radiocarbon analyses of carbonate archives at high spatial resolution. Analytical Chemistry 88:8570-8576



Färber C, Wisshak M, Pyko I, Bellou N, Freiwald A (2015) Effects of water depth, seasonal exposure, and substrate orientation on microbial bioerosion in the Ionian Sea (Eastern Mediterranean). Plos One 10:e0126495

Freiwald A (2015) Kaltwasserriffe als Geobiodiversitäts-Hotspots. Natur, Forschung und Museum 145:6-13

Freiwald A (2015) Tiefseeforschungsschiff „Sonne“ nimmt Fahrt auf. Natur, Forschung und Museum 145:34-37

Matsuyama K, Titschack J, Baum D, Freiwald A (2015) Two new species of erect Bryozoa (Gymnolaemata: Cheilostomata) and the application of non-destructive imaging methods for quantitative taxonomy. Zootaxa 4020:81-100

Sakai K, Türkay M, Beuck L, Freiwald A (2015) A collection of the Infraorder Callianassidea (Decapoda, Pleocyemata) with one new genus and five new species from the Eastern Atlantic off Mauritania (R/V Maria S. Merian cruise MSM 16/3 “PHAETON”). Marine Biodiversity 45:113-133

Taviani M, Angeletti L, Beuck L, Campiani E, Canese S, Foglini F, Freiwald A, Montagna P, Trincardi F (2015) On and off the beaten track: Megafaunal sessile life and Adriatic cascading processes. Marine Geology 369:273-287

Titschack J, Baum D, De Pol-Holz R, López Correa M, Forster N, Flögel S, Hebbeln D, Freiwald A (2015) Aggradation and carbonate accumulation of Holocene Norwegian cold-water coral reefs. Sedimentology 62:1873-1898

Wisshak M, Berning B, Jakobsen J, Freiwald A (2015) Temperate carbonate production: biodiversity of calcareous epiliths from intertidal to bathyal depths (Azores). Marine Biodiversity 45:87-112



Fabri MC, Pedel L, Beuck L, Galgani F, Hebbeln D, Freiwald A (2014) Megafauna of vulnerable marine ecosystems in French Mediterranean submarine canyons: Spatial distribution and anthropogenic impacts. Deep-Sea Research II 104:184-207

Freiwald A (2014) Kaltwasserkorallen – aus der Grundlagenforschung auf die politische Agenda. Natur, Forschung, Museum 144:6-17

Hebbeln D, Wienberg C, Wintersteller P, Freiwald A, Becker M, Beuck L, Dullo C, Eberli GP, Glogowski S, Matos L, Forster N, Reyes-Bonilla H, Taviani M (2014) Environmental forcing of the Campeche cold-water coral province, southern Gulf of Mexico. Biogeosciences 11:1799-1815

Matsuyama K, Janssen A, Martínez Arbizu P, Martha SO, Freiwald A (2014) Bryozoans from RV Sonne deep-sea cruises SO 167 ‘Louisville’ and SO 205 ‘Mangan’. Zootaxa 3856:100-116

Montagna P, McCulloch M, Douville E, López Correa M, Trotter J, Rodolfo-Metalpa R, Dissard D, Ferrier-Pagès C, Frank N, Freiwald A, Goldstein S, Mazzoli C, Reynaud S, Rüggeberg A, Russo S, Taviani M (2014) Li/Mg systematics in scleractinian corals: Calibration of the thermometer. Geochimica et Cosmochimica Acta 132:288-310

Savini A, Vertino A, Marchese F, Beuck L, Freiwald A (2014) Mapping cold-water coral habitats at different scales within the northern Ionian Sea (Central Mediterranean): An assessment of coral coverage and associated vulnerability. Plos One 9:e87108

Teichert S, Freiwald A (2014) Polar coralline algal CaCO3-production rates correspond to intensity and duration of the solar radiation. Biogeosciences 11:833-842

Teichert S, Woelkerling W, Rüggeberg A, Wisshak M, Piepenburg D, Meyerhöfer M, Form A, Freiwald A (2014) Arctic rhodolith beds and their environmental controls (Spitsbergen, Norway). Facies 60:15-37

Wisshak M, Schönberg CHL, Form A, Freiwald A (2014) Sponge bioerosion accelerated by ocean acidification across species and latitudes? Helgoland Marine Research 68:253-262



Barnich R, Beuck L, Freiwald A (2013) Scale worms (Polychaeta: Aphroditiformia) associated with cold-water corals in the eastern Gulf of Mexico. Journal of the Marine Biological Association of the United Kingdom 93:2129-2143

Freiwald A, Beuck L, Wisshak M (2013) Korallenriffe im kalten Wasser des Nordatlantiks. In: Beck E (ed) Die Vielfalt des Lebens: Wie hoch, wie komplex, warum? Wiley-VCH, pp 88-98

Joseph N, López Correa M, Schönfeld J, Rüggeberg A, Freiwald A (2013) Subarctic Holocene climatic and oceanographic variability in Stjernsund, northern Norway: evidence from benthic foraminifera and stable isotopes. Boreas 42:511-531

Marali S, Wisshak M, López Correa M, Freiwald A (2013) Skeletal microstructure and stable isotope signature of three bathyal solitary cold-water corals from the Azores. Palaeogeography, Palaeoclimatology, Palaeoecology 373:25-38

Raddatz J, Liebetrau V, Rüggeberg A, Hathorne E, Krabbenhöft A, Eisenhauer A, Böhm F, Vollstaedt H, Fietzke J, López Correa M, Freiwald A, Dullo WC (2013) Stable Sr-isotope, Sr/Ca, Mg/Ca, Li/Ca and Mg/Li ratios in the scleractinian cold-water coral Lophelia pertusa. Chemical Geology 352:143-152

Sanfilippo R, Vertino A, Rosso A, Beuck L, Freiwald A, Taviani M (2013) Serpula aggregates and their role in deep-sea coral communities in the southern Adriatic Sea. Facies 59:663-677

Titschack J, Joseph N, Fietzke J, Freiwald A, Bromley RG (2013) Record of a tectonically-controlled regression captured by changes in carbonate skeletal associations on a structured island shelf (mid-Pleistocene, Rhodes, Greece). Sedimentary Geology 283:15-33

Wisshak M, Schönberg CHL, Form A, Freiwald A (2013) Effects of ocean acidification and global warming on reef bioerosion—lessons from a clionaid sponge. Aquatic Biology 19:111-127



Addamo AM, Reimer JD, Taviani M, Freiwald A, Machordom A (2012) Desmophyllum dianthus (Esper, 1794) in the scleractinian phylogeny and its intraspecific diversity. Plos One 7:e50215

Fink HG, Wienberg C, Hebbeln D, McGregor HV, Schmiedl G, Taviani M, Freiwald A (2012) Oxygen control on Holocene cold-water coral development in the eastern Mediterranean Sea. Deep-Sea Research I 62:89-96

Hebbeln D, Freiwald A (2012) Manche mögen’s kalt und dunkel. In: Wefer G, Schmieder F, Freifrau von Neuhoff S (eds) Tiefsee Expeditionen zu den Quellen des Lebens. Rosenheim, pp 160-167

Kuechler RR, Birgel D, Kiel S, Freiwald A, Goedert J, Thiel V, Peckmann J (2012) Miocene methane-derived carbonates from southwestern Washington, USA and a model for silification at seeps. Lethaia 45:259-273

López Correa M, Montagna P, Joseph N, Rüggeberg A, Fietzke J, Flögel S, Dorschel B, Goldstein SL, Wheeler A, Freiwald A (2012) Preboreal onset of cold-water coral growth beyond the Arctic Circle revealed by coupled radiocarbon and U-series dating and neodymium isotopes. Quaternary Science Reviews 34:24-43

McCulloch M, Trotter J, Montagna P, Falter J, Dunbar R, Freiwald A, Försterra G, López Correa M, Maier C, Rüggeberg A, Taviani M (2012) Resilience of cold-water scleractinian corals to ocean acidification: Boron isotopic systematics of pH and saturation state up-regulation. Geochimica et Cosmochimica Acta 87:21-34

Teichert S, Woelkerling W, Rüggeberg A, Wisshak M, Piepenburg D, Meyerhöfer M, Form A, Büdenbender J, Freiwald A (2012) Rhodolith beds (Corallinales, Rhodophyta) and their physical and biological environment at 80°31’N in Nordkappbukta (Nordaustlandet, Svalbard Archipelago, Norway). Phycologia 51:371-390

Wisshak M, Schönberg CHL, Form A, Freiwald A (2012) Ocean acidification accelerates reef bioerosion. Plos One 7:e45124



Eisele M, Frank N, Wienberg C, Hebbeln D, López Correa M, Douville E, Freiwald A (2011) Productivity controlled cold-water coral growth periods during the last glacial off Mauritania. Marine Geology 280:143-149

Fabri MC, Pedel L, Freiwald A, Madurell T (2011) Habitats particuliers des étages bathyal et abyssal (Med). SRM MO Initial Assessment for the Water Marine Framework Strategy

Frank N, Freiwald A, López Correa M, Wienberg C, Eisele M, Hebbeln D, van Rooij D, Henriet JP, Colin C, van Weering T, De Haas H, Buhl-Mortensen P, Roberts JM, De Mol B, Douville E, Blamart D, Hatte C (2011) Northeastern Atlantic cold water coral reefs and climate. Geology 39:743-746

Freiwald A (2011) Cold-water coral reefs. Encyclopedia of modern coral reefs. Springer, Heidelberg

Freiwald A, Beuck L (2011) Sind Kaltwasserkorallen durch den Klimawandel gefährdet? In: Lozán JL, Graßl H, Karbe L, Reise K (eds) Warnsignal Klima: Die Meere – Änderungen und Risiken. Hamburg, pp 220-224

Rüggeberg A, Flögel S, Dullo WC, Hissmann K, Freiwald A (2011) Water mass characteristics and sill dynamics in a subpolar cold-water coral reef setting at Stjernsund, northern Norway. Marine Geology 282:5-12

Schönfeld J, Dullo WC, Pfannkuche O, Freiwald A, Rüggeberg A, Schmidt S, Weston J (2011) Recent benthic foraminiferal assemblages from cold-water coral mounds in the Porcupine Seabight. Facies 57:187-213

Wisshak M, Tribollet A, Golubic S, Jakobsen J, Freiwald A (2011) Temperate bioerosion: ichnodiversity and biodiversity from intertidal to bathyal depths (Azores). Geobiology 9:492-520



Beuck L, Freiwald A, Taviani M (2010) Spatiotemporal bioerosion patterns in deep-water scleractinians from off Santa Maria di Leuca (Apulia, Ionian Sea). Deep-Sea Research II 57:458-470

Copard K, Colin C, Douville E, Freiwald A, Gudmundsson G, De Mol B, Frank N (2010) Nd isotopes in deep-sea corals in the North-eastern Atlantic. Quaternary Science Reviews 29:2499-2508

Hebbeln D, Freiwald A (2010) Manche mögen’s kalt. Der europäische Korallengürtel von Norwegen bis ins Mittelmeer. In:  Klima und Leben. pp 6-13

Heindel K, Titschack J, Dorschel B, Huvenne VAI, Freiwald A (2010) The sediment composition and predictive mapping of facies on the Propeller Mound – A cold-water coral mound (Porcupine Seabight, NE Atlantic). Continental Shelf Research 30:1814-1829

Taviani M, Freiwald A, Beuck L, Angeletti L, Remia A, Vertino A, Dimech M, Schembri PJ (2010) The deepest known occurrence of the precious red coral Corallium rubrum (L., 1758) in the Mediterranean Sea. Proceedings of the International Workshop on Red Coral Science, Management, and Trade: Lessons from the Mediterranean. NOAA Technical Memorandum CRCP-13,

Trincardi F, Angeletti L, Asioli A, Campiani E, Ceregato A, Foglini F, Freiwald A, Remia A, Taviani M, Vertino A (2010) South West Adriatic margin morphology and deep-sea macrobenthic ecosystems. Rapport de la Commission International pour l’Exploration Scientifique de la Mer Méditerranée 39:70

Westphal H, Halfar J, Freiwald A (2010) Heterozoan carbonates in subtropical to tropical settings in the present and in the past. International Journal of Earth Sciences 99 Supplement 1:S153-S169

Wisshak M, Form A, Jakobsen J, Freiwald A (2010) Temperate carbonate cycling and water mass properties from intertidal to bathyal depths (Azores, N-Atlantic). Biogeosciences 7:2379-2396



Freiwald A, Beuck L, Rüggeberg A, Taviani M, Hebbeln D (2009) The white coral community in the central Mediterranean Sea revealed by ROV surveys. Oceanography 22:58-74

Roberts JM, Wheeler AJ, Freiwald A, Cairns SD (2009) Cold-water corals – The biology and geology of deep-sea coral habitats. Cambridge University Press,

Taviani M, Angeletti L, Dimech M, Mifsud C, Freiwald A, Harasewych MG, Oliverio M (2009) Coralliophilinae (Gastropoda: Muricidae) associated with deep-water coral banks in the Mediterranean. The Nautilus 123:106-112

Titschack J, Radtke U, Freiwald A (2009) Dating and characterization of polymorphic transformation of aragonite to calcite in Pleistocene bivalves from Rhodes (Greece) by combined shell microstructure, stable isotope, and electron spin resonance study. Journal of Sedimentary Research 79:332-346

Titschack J, Thierens M, Dorschel B, Schulbert C, Freiwald A, Kano A, Takashima C, Kawagoe N, Li X (2009) Carbonate budget of a cold-water coral mound (Challenger Mound, IODP Exp. 307). Marine Geology 259:36-46

Weaver PPE, Boetius A, Danovaro R, Freiwald A, Gunn V, Heussner S, Morato T, Schewe I, van den Howe S (2009) The future of integrated deep-sea research in Europe—The HERMIONE Project. Oceanography 22:177-191

Wienberg C, Hebbeln D, Fink HG, Mienis F, Dorschel B, Vertino A, López Correa M, Freiwald A (2009) Scleractinian cold-water corals in the Gulf of Cádiz—First clues about their spatial and temporal distribution. Deep-Sea Research I 56:1873-1893

Wisshak M, López Correa M, Gofas S, Salas C, Taviani M, Jakobsen J, Freiwald A (2009) Shell architecture, element composition, and stable isotope signature of the giant deep-sea oyster Neopycnodonte zibrowii sp. n. from the NE Atlantic. Deep-Sea Research I 66:374-407

Wisshak M, López Correa M, Zibrowius H, Jakobsen J, Freiwald A (2009) Skeletal reorganisation affects geochemical signals, exemplified in the stylasterid hydrocoral Errina dabneyi (Azores Archipelago). Marine Ecology Progress Series 397:197-208

Wisshak M, Neumann C, Jakobsen J, Freiwald A (2009) The ‘living-fossil community’ of the cyrtocrinoid Cyathidium foresti and the deep-sea oyster Neopycnodonte zibrowii (Azores Archipelago). Palaeogeography, Palaeoclimatology, Palaeoecology 271:77-83



Beuck L, López Correa M, Freiwald A (2008) Biogeographical distribution of Hyrrokkin (Rosalinidae, Foraminifera) and its host-specific morphological and textural trace variability. In: Wisshak M, Tapanila L (eds) Current Developments in Bioerosion. Springer, Heidelberg, pp 329-360

Beuck L, Wisshak M, Munnecke A, Freiwald A (2008) A giant boring in a Silurian stromatoporoid analysed by computer tomography. Acta Palaeontologica Polonica 53:149-160

Birgel D, Himmler T, Freiwald A, Peckmann J (2008) A new constraint on the antiquity of anaerobic oxidation of methane: Late Pennsylvanian seep limestones from southern Namibia. Geology 36 (7):543-546

Himmler T, Freiwald A, Stollhofen H, Peckmann J (2008) Late Carboniferous hydrocarbon-seep carbonates from the glaciomarine Dwyka Group, southern Namibia. Palaeogeography, Palaeoclimatology, Palaeoecology 257:185-197

Montagna P, López Correa M, Rüggeberg A, McCulloch M, Rodolfo-Metalpa R, Dullo WC, Ferrier-Pagès C, Freiwald A, Henderson GM, Mazzoli C, Russo S, Silenzi S, Taviani M (2008) Coral Li/Ca in micro-structural domains as a temperature proxy. Geochimica et Cosmochimica Acta 72:A645

Reveillaud J, Freiwald A, Van Rooij D, Le Guilloux E, Altuna A, Foubert A, Vanreusel A, Olu-Le Roy K, Henriet JP (2008) The distribution of scleractinian corals in the Bay of Biscay, NE Atlantic. Facies 54:317-331

Rüggeberg A, Fietzke J, Liebetrau V, Eisenhauer A, Dullo WC, Freiwald A (2008) Stable strontium isotopes (d88/86Sr) in cold-water corals— a new proxy for reconstruction of intermediate ocean water temperatures. Earth and Planetary Science Letters 269:569-574

Titschack J, Nelson CS, Beck T, Freiwald A (2008) Sedimentary evolution of a Late Pleistocene temperate red algal reef (Coralligène) on Rhodes, Greece: correlation with global sea-level fluctuations. Sedimentology 55:1747-1776

Wienberg C, Beuck L, Heidkamp S, Hebbeln D, Freiwald A, Pfannkuche O, Monteys X (2008) Franken Mound: facies and biocoenoses on a newly-discovered “carbonate mound” on the western Rockall Bank, NE Atlantic. Facies 54:1-24



Dullo WC, Freiwald A, Rüggeberg A, Taviani M (2007) Cold-water coral reefs around Europe – Hotspots from the central Mediterranean. Rapport Commission International Mer Méditerranée 38:84

Kiriakoulakis K, Freiwald A, Fisher E, Wolff G (2007) Organic matter quality and supply to deep-water coral/mound systems of the NW European continental margin. International Journal of Earth Sciences 96:159-170

Lopez Correa M, Bednorz A, Freiwald A (2007) Cold-water corals as climate archives in the ocean depths. SOMATOM Sessions 11/07:52-54

Noé S, Lembke-Jene L, Reveillaud J, Freiwald A (2007) Microstructure, growth banding and age determination of a primnoid gorgonian skeleton (Octocorallia) from the late Younger Dryas to earliest Holocene of the Bay of Biscay. Facies 53:177-188

Wheeler AJ, Beyer A, Freiwald A, de Haas H, Huvenne VAI, Kozachenko M, Olu-Le Roy K, Opderbecke J (2007) Morphology and environment of cold-water coral carbonate mounds on the NE European margin. International Journal of Earth Sciences 96:37-57



Guinotte JM, Orr J, Cairns S, Freiwald A, Morgan L, George R (2006) Will human-induced changes in seawater chemistry alter the distribution of deep-sea scleractinian corals? Frontiers in the Ecology and the Environment 4:141-146

Noé S, Titschack J, Freiwald A, Dullo WC (2006) From sediment to rock: diagenetic processes of hardground formation in deep-water carbonate mounds of the NE Atlantic. Facies 52:183-208

Roberts JM, Wheeler AJ, Freiwald A (2006) Reefs of the deep: the biology and geology of cold-water coral ecosystems. Science 312:543-547

Wisshak M, Connor D, Hain S, Golding N, Freiwald A (2006) Lophelia pertusa in the Northeast Atlantic: the current record. UNEP Data Set



Beuck L, Freiwald A (2005) Bioerosion patterns in a deep-water Lophelia pertusa (Scleractinia) thicket (Propeller Mound, northern Porcupine Seabight). In: Freiwald A, Roberts JM (eds) Cold-water corals and ecosystems. Springer, pp 915-936

Costello MJ, McCrea M, Freiwald A, Lundälv T, Jonsson L, Bett BJ, van Weering TCE, de Haas H, Roberts JM, Allen D (2005) Role of cold-water Lophelia pertusa coral reefs as fish habitat in the NE Atlantic. In: Freiwald A, Roberts JM (eds) Cold-water corals and ecosystems. Springer, Heidelberg, pp 771-805

Dorschel B, Hebbeln D, Rüggeberg A, Dullo WC, Freiwald A (2005) Growth and erosion of a cold-water coral covered carbonate mound in the Northeast Atlantic during the Late Pleistocene and Holocene. Earth and Planetary Science Letters 233:33-44

Försterra G, Beuck L, Häussermann V, Freiwald A (2005) Shallow-water Desmophyllum dianthus (Scleractinia) from Chile: characteristics of the biocoenoses, the bioeroding community, heterotrophic interactions and (Paleo)-bathymetric implications. In: Freiwald A, Roberts JM (eds) Cold-water corals and ecosystems. Springer, Heidelberg, pp 937-977

Freiwald A, Rogers A, Hall-Spencer J (2005) Global distribution of cold-water corals (version 2). UNEP World Conservation Monitoring Centre,

Kiriakoulakis K, Fisher E, Wolff GA, Freiwald A, Grehan A, Roberts JM (2005) Lipids and nitrogen isotopes of two deep-water corals from the North-East Atlantic: initial results and implications for their nutrition. In: Freiwald A, Roberts JM (eds) Cold-water corals and ecosystems. Springer, Heidelberg, pp 715-729

Könnecker G, Freiwald A (2005) Plectroninia celtica n.sp. (Calcarea, Minchinellidae), a new species of “pharetronid” sponge from bathyal depths in the northern Porcupine Seabight, NE Atlantic. Facies 51:53-59

López Correa M, Freiwald A, Hall-Spencer J, Taviani M (2005) Distribution and habitats of Acesta excavata (Bivalvia: Limidae) with new data on its shell ultrastructure. In: Freiwald A, Roberts JM (eds) Cold-water corals and ecosystems. Springer, Heidelberg, pp 173-205

Roberts JM, Freiwald A (2005) Integrated European research into cold-water coral reefs. Current 21:41-45

Schröder-Ritzrau A, Freiwald A, Mangini A (2005) U/Th-dating of deep-water corals from the eastern North Atlantic and the western Mediterranean Sea. In: Freiwald A, Roberts JM (eds) Cold-water corals and ecosystems. Springer, Heidelberg, pp 157-172

Taviani M, Freiwald A, Zibrowius H (2005) Deep coral growth in the Mediterranean Sea: an overview. In: Freiwald A, Roberts JM (eds) Cold-water corals and ecosystems. Springer, Heidelberg, pp 137-156

Taviani M, Remia A, Corselli C, Freiwald A, Malinverno E, Mastrototaro F, Savini A, Tursi A (2005) First geo-marine survey of living cold-water Lophelia reefs in the Ionian Sea (Mediterranean basin). Facies 50:409-417

Titschack J, Bromley RG, Freiwald A (2005) Plio-Pleistocene cliff-bound, wedge-shaped, warm-temperate carbonate deposits from Rhodes (Greece): Sedimentology and facies. Sedimentary Geology 180:29-56

Titschack J, Freiwald A (2005) Growth, deposition, and facies of Pleistocene bathyal coral communities from Rhodes, Greece. In: Freiwald A, Roberts JM (eds) Cold-water corals and ecosystems. Springer, Heidelberg, pp 41-59

Wisshak M, Freiwald A, Lundälv T, Gektidis M (2005) The physical niche of the bathyal Lophelia pertusa in a non-bathyal setting: environmental controls and palaeoecological implications. In: Freiwald A, Roberts JM (eds) Cold-water corals and ecosystems. Springer, Heidelberg, pp 979-1001

Wisshak M, Gektidis M, Freiwald A, Lundälv T (2005) Bioerosion along a bathymetric gradient in a cold-temperate setting (Kosterfjord, SW Sweden): an experimental study. Facies 51:93-117



Freiwald A (2004) Kaltwasser-Karbonate und Tiefwasser-Riffe – eine geobiologische Herausforderung. Gmit 18:8-14

Freiwald A, Fosså JH, Grehan A, Koslow T, Roberts JM (2004) Cold-water coral reefs. UNEP-WCMC Biodiversity Series 22:1-84

Mienert J, Weaver PPE, Berné S, Dullo WC, Evans D, Freiwald A, Henriet JP, Joergensen BB, Lericolas G, Lykousis V, Parkes J, Trincardi F, Westbrook G (2004) Dynamics and evolution of European margins. Oceanography 17:12-29

Weaver PPE, Billett DSM, Boetius A, Danovaro R, Freiwald A, Sibuet M (2004) Hotspot ecosystem research on Europe’s deep-ocean margins. Oceanography 17:132-143

Wisshak M, Volohonsky E, Seilacher A, Freiwald A (2004) A trace fossil assemblage from fluvial Old Red deposits (Wood Bay Formation; Lower to Middle Devonian) of NW-Spitsbergen, Svalbard. Lethaia 37:149-163



Beck T, Metzger T, Freiwald A (2003) BIAS Biodiversity Inventorial Atlas of macrobenthic seamount animals. OASIS Deliverable 25 1-127

Blomeier D, Wisshak M, Dallmann W, Volohonsky E, Freiwald A (2003) Facies analysis of the Old Red sandstone of Spitsbergen (Wood Bay Formation): Reconstruction of the depositional environments and implications of basin development. Facies 49:151-174

Blomeier D, Wisshak M, Joachimski M, Freiwald A, Volohonsky E (2003) Calcareous, alluvial and lacustrine deposits in the Old Red Sandstone of central north Spitsbergen (Wood Bay Formation, Early Devonian). Norwegian Journal of Geology 83:281-298

Freiwald A (2003) Korallengärten in kalten Tiefen. Spektrum der Wissenschaft 2/2003:666-673

Schäfer P, Freiwald A, Meyer J, Miketta P, Nadzeika U (2003) Ice-floe induced sedimentation during extreme winters: a case study from the Schleswig-Holstein Wadden Sea. Meyniana 55:73-90



Freiwald A (2002) Reef-forming cold-water corals. In: Wefer G, Billett D, Hebbeln D, Jørgensen BB, Schlüter M, van Weering TCE (eds) Ocean Margin Systems. Springer, pp 365-385

Freiwald A, Hühnerbach V, Lindberg B, Wilson JB, Campbell J (2002) The Sula Reef complex, Norwegian Shelf. Facies 47:179-200

Sorauf JE, Freiwald A (2002) Skeletal structure in the rugosan genus Calophyllum from Permian strata of Timor: Comparison with the living deep water coral Lophelia pertusa. Coral Research Bulletin 7:191-207



Grehan AJ, Freiwald A (2001) The Atlantic Coral Ecosystem Study (ACES): Forging a new partnership between scientists and principal stakeholders. In: Willison JHM, Hall J, Gass S, Kenchington ELR, Butler M, Doherty P (eds) Proceedings of the 1st International Symposium on Deep-Sea Corals. Ecology Action Centre and Nova Scotia Museum, pp 95-105

Nelson CS, Freiwald A, Titschack J, List S (2001) Lithostratigraphy and sequence architecture of temperate mixed siliciclastic-carbonate facies in a new Plio-Pleistocene section at Plimiri, Rhodes Island (Greece). Occasional Report Department of Earth Sciences, University of Waikato 25:1-50



Freiwald A, Mortensen PB (2000) The first record of the deep-water coral Stenocyathus vermiformis (Pourtalès, 1868) (Scleractinia, Guyniidae) from Norwegian waters. Sarsia 85:275-276



Cuif JP, Dauphin Y, Freiwald A, Gautret P, Zibrowius H (1999) Biochemical markers of zooxanthellae symbiosis in soluble matrices of skeleton of 24 Scleractinia species. Comparative Biochemistry and Physiology, Part A: Physiology 123:269-278

Freiwald A, Wilson JB, Henrich R (1999) Grounding Pleistocene icebergs shape recent deep-water coral reefs. Sedimentary Geology 125:1-8



Freiwald A (1998) Microbial maceration and carbonate dissolution on cold-temperate shelves. Historical Biology 13:27-35

Freiwald A (1998) Modern nearshore cold-temperate calcareous sediments in the Troms District, northern Norway. Journal of Sedimentary Research 68:763-776

Freiwald A, Mostafawi N (1998) Ostracods in a cold-temperate coastal environment, western Troms, northern Norway: sedimentary aspects and assemblages. Facies 38:255-274

Freiwald A, Wilson JB (1998) Taphonomy of modern deep, cold-temperate water coral reefs. Historical Biology 13:37-52



Freiwald A, Henrich R, Pätzold J (1997) Anatomy of a deep-water coral reef mound from Stjernsund, West-Finnmark, northern Norway. In: James NP, Clarke JAD (eds) Cool-water carbonates, vol 56. SEPM, Special Publication, pp 141-161

Freiwald A, Reitner J, Krutschinna J (1997) Microbial alteration of the deep-water coral Lophelia pertusa: early postmortem processes. In: Neuweiler F, Reitner J, Monty C (eds) Biosedimentology of microbial buildups IGCP Project No. 380 – Proceedings of 2nd Meeting, 1997. Facies, pp 223-226

Gautret P, Cuif JP, Freiwald A (1997) Composition of soluble mineralizing matrices in zooxanthellate and non-zooxanthellate scleractinian corals: biochemical assessment of photosynthetic metabolism through the study of a skeletal feature. Facies 36:189-194

Henrich R, Freiwald A, Bickert T, Schäfer P (1997) Evolution of an Arctic open-shelf carbonate platform, Spitsbergen Bank (Barents Sea). In: James NP, Clarke JAD (eds) Cool-water carbonates, vol 56. SEPM, Special Volume, pp 163-181



Andruleit H, Freiwald A, Schäfer P (1996) Bioclastic carbonate sediments on the southwestern Svalbard Shelf. Marine Geology 134:163-182

Freiwald A, Schönfeld J (1996) Substrate pitting and boring pattern of Hyrrokkin sarcophaga Cedhagen, 1994 (Foraminifera) in a modern deep-water coral reef mound. Marine Micropaleontology 28:199-207

Henrich R, Freiwald A, Wehrmann A, Schäfer P, Samtleben C, Zankl H (1996) Nordic cold-water carbonates: occurrences and controls. Göttinger Arbeiten zur Geologie und Paläontologie Sb 2:35-52



Freiwald A (1995) Bacteria-induced carbonate degradation: a taphonomic case study of Cibicides lobatulus from a high-boreal carbonate setting. Palaios 10:337-346

Freiwald A (1995) Sedimentological and biological aspects in the formation of branched rhodoliths in northern Norway. Beiträge zur Paläontologie 20:7-19

Henrich R, Freiwald A, Betzler CB, Bader B, Schäfer P, Samtleben C, Brachert TC, Wehrmann A, Zankl H, Kühlmann DHH (1995) Controls on modern carbonate sedimentation on warm-temperate to Arctic coasts, shelves and seamounts in the northern Hemisphere: implications for fossil counterparts. Facies 32:71-108

Wehrmann A, Freiwald A, Zankl H (1995) Formation of cold-temperate water multispecies rhodoliths in intertidal gravel pools from northern Brittany, France. Senckenbergiana maritima 26:51-71



Freiwald A, Henrich R (1994) Reefal coralline algal build-ups within the Arctic Circle: morphology and sedimentary dynamics under extreme environmental seasonality. Sedimentology 41:963-984



Freiwald A (1993) Coralline algal maerl frameworks-Islands within the phaeophytic kelp belt. Facies 29:133-148

Freiwald A (1993) Subarktische Kalkalgenriffe im Spiegel hochfrequenter Meeresspiegelschwankungen und interner biologischer Steuerungsprozesse. PhD Thesis, Kiel University

Schmidt H, Freiwald A (1993) Rezente gesteinsbohrende Kleinorganismen des norwegischen Schelfs. Natur und Museum 123:149-155



Freiwald A, Willmann R (1992) lnsekten aus der Fur-Formation von Dänemark (Moler, ob. Paleozän I unt. Eozän ?). 7. Ptychopteridae (Diptera). Meyniana 44:179-187

Henrich R, Hartmann M, Reitner J, Schäfer P, Freiwald A, Steinmetz S, Dietrich P, Thiede J (1992) Facies belts and communities of the Arctic Vesterisbanken Seamount (Central Greenland Sea). Facies 27:71-104



Freiwald A (1991) lnsekten aus der Fur-Formation von Dänemark (Moler, ob. Paleozän / unt. Eozän 3). 5. Cylindrotomidae (Diptera: Tipulomorpha). Meyniana 43:97-123

Freiwald A, Henrich R, Schäfer P, Willkomm H (1991) The significance of high-boreal to subarctic maerl deposits in northern Norway to reconstruct Holocene climatic changes and sea level oscillations. Facies 25:315-340

Freiwald A, Krzeminski W (1991) Cylindrotomidae (Diptera, Tipulomorpha) from the Paleogene of Bolshaya Svetlovodnaya (Eastern Asiatic USSR). Paläontologische Zeitschrift 65:339-344

Krzeminski W, Freiwald A (1991) Toxorhina (Ceratocheilus) caucasiensis, a new species from the Middle Miocene of Stavropol (northern Caucasus, USSR) (Diptera, Limoniidae). Paläontologische Zeitschrift 65:153-155



Freiwald A (1990) Insekten aus der Fur-Formation von Dänemark (Moler, ob. Paleozän / unt. Eozän ?) 4. Tipulidae. Meyniana 42:47-63



Team Dieter Uhl
Prof. Dr. Dieter Uhl
Head of Department Palaeontology and Historical Geology, Head of Section Palaeoclimate and Palaeoenvironmental Research

Main research interests

  • Evolution of terrestrial ecosystems and biodiversity in relation to abiotic factors during the last 300 million years (e.g. climate, fire and tectonics).
  • Qualitative and quantitative palaeoclimate and palaeoenvironmental reconstruction based on palaeobotanical proxies (i.e. palynomorphs and leaves).
  • Taphonomy of plant macro-remains.

Curriculum  Vitae

1990-1994 Studies in Biology / Kaiserslautern University
1994 Diploma in Biology (Plant physiology / Botany)
1995 Biology Department / Kaiserslautern University
1995 Working contract / Pfalzmuseum für Naturkunde Bad Dürkheim (Permocarboniferous palaeontology)
1996-2000 Palaeontology Department / Tübingen University
Nov. 1999 Dr. rer. nat. in Geology/Palaeontology
2000-2002 Post-Doc, Forschungsstelle für Paläobotanik / Münster University
2003-2004 Post-Doc, Institut of Geosciences / Tübingen University
Jan. 2005 Habilitation in Palaeontology / Tübingen University
Jan. 2005 – Jan. 2010 Privatdozent at Tübingen University
2005-2006 Feodor-Lynen Research Fellow (Alexander von Humboldt Foundation), Department of Paleoecology / Utrecht University
Mar. 2005 NWO – Von Humboldt Stiftung Prijs, awarded by Nederlandse Organisatie voor Wetenschappelijk Onderzoek (NWO)
2007 Freelance palaeobotanist
since 09/2007 Scientist, Senckenberg Forschungsinstitut und Naturmuseum Frankfurt am Main; (from 10/2007 – 12/2008 delegated to the Hessian Ministry of Science and Fine Arts)
since 01/2010 apl. Prof. at Tübingen University and head of section for Palaeoclimate and Palaeoenvironmental Research
05/2011 – 01/2014 Coordinator Senckenberg Centre for Human Evolution and Palaeoenvironment
01/2013 – 10/2016 temporary head of the Dept. of Palaeontology and Historical Geology
12/2013 – 11/2016 Visiting guest researcher (“Pesquisador Visitante Especial”) at „Programa de Pós-Graduação em Ambiente e Desenvolvimento“ at UNIVATES (Lajeado, RS / Brazil) financed by “CAPES/CNPq – Ciências Sem Fronteiras”
since 11/2016 head of the Dept. of Palaeontology and Historical Geology

SS – Vorlesung Biologie für Geowissenschaftler (Übernahme einzelner Themen)
WS – Vorlesung und Übungen Paläobotanik/Palynologie (Kompaktkurs im Anschluss an das Semester, zusammen mit A. Bruch)
WS – Vorlesung und Übungen Topics in Continental Paleobiology and -climatology (zusammen mit M. Bohme)

Publications Dieter Uhl
In press/accepted 

de Lima, F.J., Pires, E.F., Jasper, A., Uhl, D., Saraiva, A.A.F., Sayão, J.M., in press. Early Cretaceous (Aptian) wildfires in the Araripe Basin, Northeast Brazil – palaeoclimatic and palaeoenvironmental implications. In: R. Iannuzzi, R. Rößler & L. Kunzmann (eds.), Brazilian Paleofloras, 12 pp., Springer. 
Kunzmann, L., Coiffard, C., de Oliveira Westerkamp, A.P.A., Batista, M.E.P., Uhl, D., Solórzano Kraemer, M.M., Mendes, M., Nascimento, Jr., D.R:, Innuzzi, R. & Silva Filho, W.F., accepted. The Crato flora, a 115 Million years old window into the Cretaceous world of Brazil. In: R. Iannuzzi, R. Rößler & L. Kunzmann (eds.), Brazilian Paleofloras, xx-xx. Springer. 
Spiekermann, R., Jasper, A., Guerra-Sommer, M., Ricardi-Branco, F.S., de Faria, R.S. & Uhl, D., in press. The Permian lycopsids from Brazil. In: R. Iannuzzi, R. Rößler & L. Kunzmann (eds.), Brazilian Paleofloras, 29 pp., Springer. 
Jasper, A., Uhl, D., Spiekermann, R., Benicio, J.S.W., Brugnera, A.S., Rockenbach, C., Carniere, J.S. & Silva, A.P., in press. Wildfires in Late Palaeozoic Strata in Brazil. In: R. Iannuzzi, R. Rößler & L. Kunzmann (eds.), Brazilian Paleofloras, 15 pp., Springer. 
Uhl, D., Spiekermann, R., Wuttke, M., Poschmann, M.J. & Jasper, A., 2022. Wildfires during the Paleogene (late Eocene – late Oligocene) of the Neuwied Basin (W-Germany). Review of Palaeobotany and Palynology, 297, 104565 (12 pp.) 
Cleal, C.J., Pardoe, H.S., Berry, C.M., Cascales-Miñana, B., Davis, B.A.S., Diez, J.B., Filipova-Marinova, M.V., Giesecke, T., Hilton, J.,  Ivanov, D., Kustatscher, E., Leroy, S.A.G., McElwain, J.C., Opluštil, S., Popa, M.E., Seyfullah, L.J., Stolle, E., Thomas, B.A. & Uhl, D., 2021. Palaeobotanical experiences of plant diversity in deep time. 1: How well can we identify past plant diversity in the fossil record? Palaeogeography, Palaeoclimatology, Palaeoecology, 576, 110481 (20 pp.) 
Coiffard, C., Uhl, D., Aba Alkhayl, S.S., & El Atfy, H., 2021. An angiosperm dominated florula from the Cretaceous of Saudi Arabia. Cretaceous Research, 128, 104966 (9 pp.) 
de Lima, F.J, Sayão, J.M., de Oliveira Ponciano, L.C.M., Weinschütz, L.C., Figueiredo, R.G., Rodrigues, T., Bantim, R.A.M., Saraiva, A.A.F., Jasper, A., Uhl, D. & Kellner, A.W.A., 2021. Wildfires in the Campanian of James Ross Island: a new macro-charcoal record for the Antarctic Peninsula. Polar Research, 40, 5487 (10 pp.) 
El Atfy, H., El Beialy,S., El Khoriby, E.M. & Uhl, D., 2021. Continental palynomorphs from the Dabaa Formation, North-Western Desert, Egypt: a contribution to the reconstruction of the vegetation on the southern shores of the Tethys Ocean during the Early Oligocene. Botanical Journal of the Linnean Society, 197, 291–321. 
El Atfy, H. & Uhl, D., 2021. Palynology and palynofacies of sediments surrounding the Edmontosaurus annectens mummy at the Senckenberg Naturmuseum in Frankfurt/Main (Germany). Zeitschrift der Deutschen Gesellschaft für Geowissenschaften, 172, 127-139.  
Jasper, A., Pozzebon-Silva, Â, Siquera Carniere, J. & Uhl, D., 2021. Palaeozoic and Mesozoic palaeo-wildfires – An overview on advances in the 21st century. Journal of Palaeosciences, 70, 159-171. 
Marder, F., Mueller de Lara, D., Jasper, A., Ethur, E.M., Uhl, D., Stülp, S., 2021. Avalição Química de lenhos carbonizados de Araucaria columnaris sob diferentes concentrações de oxigênio como comparativo de análise de charcoal fóssil. Química Nova, 44, 288-294. 
Marder, F., Mueller de Lara, D., Jasper, A., Uhl, D., Stülp, S., 2021. Evaluation of carbonized woods from Araucariaceae in different oxygen concentrations through thermogravimetry and scanning electron microscope. Ciencia e Natura, 43, e77 (23 pp.) 
Murthy, S., Mendhe, V.A., Uhl, D., Mathews, R.P., Mishra, V.K. & Gautam, S., 2021. Palaeobotanical and biomarker evidence for wildfire in the Early Permian (Artinskian) of the Rajmahal Basin, India. Journal of Palaeogeography, 10, 5 (21 pp.) 
Pardoe, H.S., Cleal, C.J., Berry, C.M., Cascales-Miñana, B., David, B.A.S., Diez, J.B., Filipova-Marinova, M.V., Giesecke, T., Hilton, J.,  Ivanov, D., Kustatscher, E., Leroy, S.A.G., McElwain, J.C., Opluštil, S., Popa, M.E., Seyfullah, L.J., Stolle, E., Thomas, B.A. & Uhl, D., 2021. Palaeobotanical experiences of plant diversity in deep time. 2: How to measure and analyse past plant biodiversity. Palaeogeography, Palaeoclimatology, Palaeoecology, 580, 110618 (26 pp.) 
Poschmann, M.J., Nel, A., Schindler, T. & Uhl, D., 2021. Die Niedermoschel-Bank, ein bedeutender Fundhorizont mit einer hoch-diversen Insekten- und Pflanzenassoziation im Rotliegend (Unterperm) des Saar-Nahe-Beckens (Rheinland-Pfalz, SW-Deutschland). Mainzer naturwissenschaftliches Archiv, 58, 17-40 
Spiekermann, R., Dolezych, M., Wuttke, M. & Uhl, D., 2021. Ein Cupressaceen-Holz aus dem pliozänen Kratersee von Ruppach-Goldhausen (Westerwald, Rheinland-Pfalz, SW-Deutschland). Mainzer naturwissenschaftliches Archiv, 58, 41-52. 
Spiekermann, R., Jasper, A., Siegloch, A.M., Guerra-Sommer, M., Uhl, D., 2021. Not a lycopsid but a cycad-like plant: Iratinia australis gen. nov. et sp. nov. from the Irati Formation, Kungurian of the Paraná Basin, Brazil. Review of Palaeobotany and Palynology, 289, 104415 (14 pp.) 
Uhl, D., Bruch, A.A. & Utescher, T., 2021. Introduction to the special issue “Palaeobotanical contributions in honour of Volker Mosbrugger”. In: Uhl, D., Bruch, A.A. & Utescher, T. (eds.) Palaeobotanical contributions in honour of Volker Mosbrugger. Palaeobiodiversity and Palaeoenvironments, 101, 1-8. 
Uhl, D. & Havlik, P., 2021. Edmonds Urzeit – Eine Dinograbung in Frankfurt. Biologie in unserer Zeit, 51, 238-245. 
Uhl, D. & Jasper, A., 2021. Wildfire during deposition of the “Illinger Flözzone” (Heusweiler-Formation; “Stephanian B”, Kasimovian – Ghzelian) in the Saar-Nahe Basin (SW-Germany). In: Uhl, D., Bruch, A.A. & Utescher, T. (eds.) Palaeobotanical contributions in honour of Volker Mosbrugger. Palaeobiodiversity and Palaeoenvironments, 101, 9-18. 
Uhl, D., Paudayal, K., Hervet, S. & El Atfy, H., 2021. Menatanthus mosbruggeri gen. nov. et sp. nov. – A flower with in situ pollen tetrads from the Paleocene maar lake of Menat (Puy-de-Dôme, France). In: Uhl, D., Bruch, A.A. & Utescher, T. (eds.) Palaeobotanical contributions in honour of Volker Mosbrugger. Palaeobiodiversity and Palaeoenvironments, 101, 51-58. 
El Atfy, H., Jasper, A. & Uhl, D., 2020. A new record of Paradoxopteris stromeri Hirmer 1927 (Monilophyta, incertae sedis) from the Cenomanian of Sinai, Egypt. Review of Palaeobotany and Palynology, 273, 104148 (5 pp.)  
Kubik, R., Marynowski, L., Uhl, D. & Jasper, A., 2020. Co-occurrence of charcoal, polycyclic aromatic hydrocarbons and terrestrial biomarkers in an early Permian swamp to lagoonal depositional system, Paraná Basin, Rio Grande do Sul, Brazil. International Journal of Coal Geology, 230, 103590 (15 pp.) 
Lü Xinmiao, Paudayal, K.N., Uhl, D., Zhu Liping, Yao Tandong, Mosbrugger, V., 2020. Phenology and climatic regime inferred from airborne pollen on the northern slope of the Qomolangma (Everest) region. Journal of Geophysical Research – Atmospheres, 125 (24), e2020JD033405 (19 pp.) 
Spiekermann, R., Jasper, A., Benício, J.R.W., Guerra-Sommer, M., Ricardi-Branco, F.S. & Uhl, D., 2020. Late Palaeozoic lycopsids from the Paraná Basin in S-America – an historical overview. Journal of South American Earth Sciences, 101, 102615 (27pp.) 
Uhl, D., 2020. A reappraisal of the ‘stomach’ contents of the Edmontosaurus annectens mummy at the Senckenberg Naturmuseum in Frankfurt (Germany). Zeitschrift der Deutschen Gesellschaft für Geowissenschaften, 171(1), 71-85.  
Uhl, D., 2020. Die Edmontosaurus-Mumie (Naturmuseum Spezial: Das Edmontosaurus Projekt). SENCKENBERG – natur • forschung • museum, 150 (4-6), 84-88. 
Uhl, D., Bocherens, H., El Atfy, H., Havlik, P., Methner, K., Mulch, A. & Solórzano Kraemer, M.M., 2020. Die Umwelt zu Edmonds Lebzeiten (Naturmuseum Spezial: Das Edmontosaurus Projekt). SENCKENBERG – natur • forschung • museum, 150 (4-6), 56-65. 
Uhl, D. & Havlik, P., 2020. Wie sah Edmond aus? (Naturmuseum Spezial: Das Edmontosaurus Projekt). SENCKENBERG – natur • forschung • museum, 150 (4-6), 54-55. 
Uhl, D., Jasper, A. & Wuttke, M., 2020. Anatomical preservation of silicified Corylites J.S.Gardner leaves from the Paleocene maar lake of Menat (Puy-de-Dôme, France). Geodiversitas, 42(26), 485-492.  
Uhl, D. & Kerp, H., 2020. 4.2.3 Die terrestrische Makroflora des Zechsteins. – In: Deutsche Stratigraphische Kommission (Hrsg.; Koordination und Redaktion: J. Paul & H. Heggemann für die Subkommission Perm-Trias): Stratigraphie von Deutschland XII. Zechstein. – In: Schriftenreihe der Deutschen Gesellschaft für Geowissenschaften, 89: 83-91. 
Uhl, D., Wuttke, M. & Jasper, A., 2020. Woody charcoal with traces of pre-charring decay from the Late Oligocene (Chattian) of Norken (Westerwald, Rhineland-Palatinate, W-Germany). Acta Palaeobotanica, 60, 43-50. 
Yun Xu, Uhl, D., Zhang Ning, Zhao Cunliang, Qin Shenjun Liang Handong, Sun Yuzhuang, 2020. Evidence of widespread wildfires in coal seams from the Middle Jurassic of Northwest China and its impact on paleoclimate. Palaeogeography, Palaeoclimatology, Palaeoecology, 559, 109819 (12pp.). 


Abu Hamad, A.M.B., Al-Saqarat, B., Goncalves, C.V., Spiekermann, R., Jasper, A. & Uhl, D., 2019. The first record of Dicroidium from the Triassic palaeotropics based on dispersed cuticles from the Anisian Mukheiris Formation of Jordan. Paläontologische Zeitschrift, 93(3), 487-498.

Badihagh, M.T., Sajjadi, F., Farmani, T. & Uhl, D., 2019. Middle Jurassic palaeoenvironment and palaeobiogeography of the Tabas Block, Central Iran: palynological and palaeobotanical investigations. Palaeobiodiversity and Palaeoenvironments, 99(3), 379-399.

Badihagh, M.T. & Uhl, D., 2019. The first occurrence of Phlebopteris dunkeri and Phlebopteris woodwardii (Matoniaceae) from the Middle Jurassic of Iran. Journal of Palaeogeography, 8: 6 (10 pp).

Benício, J.R.W., Jasper, A., Spiekermann, R., Garavaglia, L., Pires-Oliveira, E.F., Machado, N.T.G. & Uhl, D., 2019. Recurrent palaeo-wildfires in a Cisularian coal seam: A palaeobotanical view on high-inertinite coals from the Lower Permian of the Paraná Basin, Brazil. PLOS One, 14(3): e0213854 (18 pp).

Benício, J.R.W., Jasper, A., Spiekermann, R., Rockenbach, C.I., Cagliari, J., Pires-Oliveira, E.F., & Uhl, D., 2019. The first evidence of palaeo-wildfire from the Itararé Group, southernmost portion of the Paraná Basin, Brazil. Journal of South American Earth Sciences, 93: 155-160.

de Lima, F.J., Pires, E.F., Jasper, A., Uhl, D., Saraiva, A.A.F., Sayão, J.M., 2019. Fire in the paradise: evidence of repeated palaeo-wildfires from the Araripe Fossil Lagerstätte (Araripe Basin, Aptian-Albian), Northeast Brazil. Palaeobiodiversity and Palaeoenvironments, 99(3), 367-378.

El Atfy, H., Anan, T., Jasper, A. & Uhl, D., 2019. Repeated occurrence of palaeo-wildfires during deposition of the Bahariya Formation (Early Cenomanian) of Egypt. Journal of Palaeogeography, 8:28 (14 pp.)

El Atfy, H., Havlik, P., Krüger, P.S., Manfroi, J., Jasper, A. & Uhl, D., 2019. Pre-Quaternary wood decay ‘caught in the act’ by fire – Examples of plant-microbe-interactions preserved in charcoal from clastic sediments. Historical Biology, 31, 952-961.

Uhl, D., Jasper, A., Solorzano Kraemer, M.M. & Wilde, V., 2019. Charred biota from an Early Cretaceous fissure fill in the Sauerland (Rüthen-Kallenhardt, Northrhine-Westphalia, W-Germany) and their palaeoenvironmental implications. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, 293(1): 83-105.

Uhl, D., Klein, E., Schindler, T. & Wuttke, M., 2019. Ein Massenvorkommen von Osmunda (Plenasium) lignitum (Giebel) Stur (Osmundaceae) im Oberoligozän von Guckheim (Westerwald, Rheinland-Pfalz, W-Deutschland). Mainzer naturwissenschaftliches Archiv, 56: 59-70.


Adhikari, P., Srivastava, G., Mehrotra, R.C., Adhikari, D., Shrestha, K., Uhl, D. & Paudayal, K.N., 2018. Leaf impressions of Terminalia (Combretaceae) and Daphnogene (Lauraceae) from the Middle Siwalik of the Chatara-Barahakshetra area, eastern Nepal. Bulletin of Department of Geosciences, Tribhuvan University, Kathmandu, Nepal, 20-21, 21-28.

Agnihotri, D., Pillai, S., Aggarwalli, N., Tewari, R., Jasper, A. & Uhl, D., 2018. Palynomorphs from the Barakar Formation of Dhanpuri Open Cast Mine, Sohagpur Coalfield, Madhya Pradesh, India. The Palaeobotanist, 67, 171-184.

Amler, M.R.W., Kauffmann, G. & Uhl, D., 2018. Chapter 42: GeoArchive Marburg (Geological and Palaeontological Collections of the former Faculty of Geosciences, Philipps University of Marburg). In: Beck, L.A. & Joger, U., (eds.), Paleontological Collections of Germany, Austria and Switzerland; 419-426. Springer.

Cardoso, D., Mizusaki, A. M. P., Guerra-Sommer, M., Menegat, R., Jasper, A. & Uhl, D., 2018. Wildfires in the Triassic of Gondwana Paraná Basin. Journal of South American Earth Sciences, 82, 193-206.

de Lara, D.M., Bresciani, L., Hilgemann, M., Osterkamp, I.C., Jasper, A., Ferrão, M.F., Uhl, D., Stülp, S., 2018. Proposição de metodologia de termodegradação por meio de fragmentos de lenjos da Araucaria columnaris. Periódico Tchê Química, 15, 106-112.

Kunzmann L., Uhl, D., Krüger, P.S., 2018. Whitish leaves in Eocene lignites in central Germany – a brief survey from the viewpoint of palaeobotany. Freiberger Forschungshefte, C 554, 213-223.

Osterkamp, I.C., de Lara, D.M., Gonçalves, T.A.P., Kauffmann, M., Périco, E., Stülp, S., Uhl, D., Jasper, A., 2018. Changes of wood anatomical characters of selected species of Araucaria during artificial charring – implications for palaeontology. Acta Botânica Brasilica, 32, 198-211.

Spiekermann, R., Uhl, D., Benício, J.R.W., Guerra-Sommer, M. & Jasper, A., 2018. A remarkable mass-assemblage of lycopsid stems from the Rio Bonito Formation, Early Permian of the Paraná Basin, Rio Grande do Sul, Brazil. Palaeobiodiversity and Palaeoenvironments, 98, 369-384.

Uhl, D. & Jasper, A., 2018. Charred conifer remains from the Late Oligocene – Early Miocene of Northern Hesse (Germany). Acta Palaeobotanica, 58, 175-184.

Uhl, D., Krüger, P.S, & Wuttke, M., 2018. Epidermal anatomy of Glyptostrobus europaeus (Brongniart) Unger from the Late Oligocene of the Westerwald (Rhineland-Palatinate, W-Germany). Fossil Imprint, 75, 334-340.

Uhl, D., Kunzmann, L., Hertler, C., Kahlke, R.-D., Keiler, J,-A., Königshof, P., Kullmer, A., Schaal, S., Smith, K., Sólorzano Kraemer, M.M., Stebich, M., Tietz, O. & Wilmsen, M., 2018. Chapter 21: Palaeontological collections of the Senckenberg Gesellschaft für Naturforschung. In: Beck, L.A. & Joger, U., (eds.), Paleontological Collections of Germany, Austria and Switzerland; 209-240. Springer.

Uhl, D. & Poschmann, M., 2018. Groenlandia pescheri sp. nov. (Potamogetonaceae) from the Late Oligocene Fossil-Lagerstätte Enspel (Westerwald, Germany). Acta Palaeobotanica, 58, 61-72.

Wedmann, S., Uhl, D., Lehmann, T., Garrouste, R., Nel, A., Gomez, B., Smith, K.T. K., Schaal, S.F.K., 2018. The Konservat-Lagerstätte Menat (Paleocene; France) – an overview and new insights. Geologica Acta, 16, 189-213.


de Lara, D.M., Bresciani, L., Osterkamp, I.C., Hilgemann, M., Ethur, E., Jasper, A., Ferrão, M.F., Uhl, D., Stulp, S., 2017. Avaliação de fragmentos de lenhos carbonizados de Araucariaceae por meio de análise por termogravimetria e infravermelhoe associado á análise multivariada. Química Nova, 40, 895-901.

El Atfy, H., Brocke, R. & Uhl, D., 2017. Palynology of the Miocene Rudeis and Kareem formations (Gharandal Group), GH 404-2A Well, Gulf of Suez, Egypt. Abhandlungen der Senckenberg Gesellschaft für Naturforschung, 573, 1-136.

Jasper, A., Agnihotri, D., Tewari, R., Spiekermann, R., Pires, E.F., da Rosa, Á.A.S. & Uhl, D., 2017. Fires in the mire: repeated fire events in Early Permian ‘peat forming’ vegetation of India. Geological Journal, 52, 955-569.

Krüger, P., Paudayal, K., Wuttke, M. & Uhl, D., 2017. Ein Beitrag zur oberoligozänen Makroflora von Norken (Westerwald, Rheinland-Pfalz, W-Deutschland). Mainzer naturwissenschaftliches Archiv, 54, 65-81.

Srivastava, G., Adhikari, P., Mehrotra, R.C., Paudel, L., Uhl, D. & Paudayal, K.N., 2017. Dipterocarpus Gaertn. (Dipterocarpaceae) leaf from the Middle Siwalik of eastern Nepal and its phytogeographic and climatic significance. Journal of Nepal Geological Society, 53, 39-46.

Scholze, F., Abu Hamad, A., Schneider, J.W., Golubev, V.K., Sennikov, A.G., Voigt, S. & Uhl, D., 2017. An enigmatic ‘conchostracan’ fauna in the eastern Dead Sea Region of Jordan: First records of Rossolimnadiopsis Novozhilov from the Early Triassic Ma’in Formation. Palaeogeography, Palaeoclimatology, Palaeoecology, 466, 314-325.

Uhl, D., 2017. Neue Funde der fossilen Krebsschere Stratiotes kaltennordheimensis (Zenker) Keilhack (Hydrocharitaceae) aus dem Oberoligozän des Westerwaldes (Rheinland-Pfalz, W-Deutschland). Mitteilungen der POLLICHIA, 98, 41-49.

Uhl, D., 2017. Die Fossillagerstätte Öhningen in Baden-Württemberg. Erdgeschichten. Geologie im Ruhrmuseum. 248-249. Verlag der Buchhandlung Walther König, Essen.


Abu Hamad, A.M.B., Amireh, B., Jasper, A. & Uhl, D., 2016. New palaeobotanical data from the Jarash Formation (Aptian-Albian, Kurnub Group) of NW Jordan. The Palaeobotanist, 65, 19-29.

Abu Hamad, A.M.B., Amireh, B., El Atfy, H., Jasper, A. & Uhl, D., 2016. Fire in a Weichselia-dominated coastal ecosystem from the Early Cretaceous (Barremian) of the Kurnub Group in NW Jordan. Cretaceous Research, 66, 82-93.

Agnihotri, D., Tewari, R., Pillai, S.S.K., Jasper, A., Uhl, D., 2016. Early Permian Glossopteris flora from Sharda Open Cast Mine, Sohagpur Coalfield, Shahdol District, Madhya Pradesh. The Palaeobotanist, 65, 97-107.

Benicio, J.R.W., Pires, E.F., da Rosa, A.A.S., Uhl, D. & Jasper, A., 2016. A new fossil Fabaceae wood from the Pleistocene Touro Passo Formation of Rio Grande do Sul State, Brazil. Fossil Imprint, 72, 251-264.

Benicio, J.R.W., Spiekermann, R., Manfroi, J., Uhl, D., Pires, E.F. & Jasper, A., 2016. Palaeoclimatic inferences based on dendrological patterns of permineralized wood from the Permian of the Northern Tocantins Petrified Forest, Parnaíba Basin, Brazil. Palaeobiodiversity and Palaeoenvironments, 96, 255-264.

Bhandari, S., Momohara, A., Uhl, D. & Paudayal, K.N., 2016. Paleoclimatic significance of the Late Quaternary plant macrofossils from the Gokarna Formation, Kathmandu Valley, Nepal. Review of Palaeobotany and Palynology, 228, 98-112.

Dos Santos, Â.C.S., Celestino Holanda, E., de Souza, V., Guerra-Sommer, M., Manfroi, J., Uhl, D. & Jasper, A., 2016. Evidence of palaeo-wildfire from the late Early Cretaceous (Serra do Tucano Formation, Aptian-Albian) of Roraima (North Brazil). Cretaceous Research, 57, 46-49.

El Atfy, H., Abeed, Q., Uhl, D. & Littke, R., 2016. Palynology, palynofacies analysis, depositional environments and source rock potential of Lower Cretaceous successions in southern Iraq. Marine and Petroleum Geology, 76, 362-376.

El Atfy, H., Sallam, H., Jasper, A. & Uhl, D., 2016. The first evidence of palaeo-wildfire from the Late Cretaceous (Campanian) of North Africa. Cretaceous Research, 57, 306-310.

Jasper, A., Manfroi, J., Uhl, D., Tewari, R., Guerra-Sommer, M., Spiekermann, R., Osterkamp, I.C., Bernardes-de-Oliveira, M.E.C., Pires, E.P., da Rosa, A.A.S., 2016. Indo-Brazilian Late Paleozoic palaeo-wildfires: an overview on macroscopic charcoal remains. Geologia USP, Série Científica, 16, 87-97.

Jasper, A., Uhl, D., Agnihotri, D., Tewari, R., Pandita, S.K., Benicio, J.R.W., Pires, E.F., da Rosa, A.A.S., Bhat, G.D., Pillai, S.S.K., 2016. Evidence of wildfire in the Late Permian (Changsinghian) Zewan Formation of Kashmir, India. Current Science, 110, 419-423.

Kauffmann, M., Jasper, A., Uhl, D., Meneghini, J., Osterkamp, I.C., Zvirtes, G. & Pires, E.F., 2016. Evidence for palaeo-wildfire in the Late Permian palaeotropics -charcoal from the Motuca Formation in the Parnaíba Basin, Brazil. Palaeogeography, Palaeoclimatology, Palaeoecology, 450, 122-128.

Nungesser, K. & Uhl, D., 2016. Neue Nachweise fossiler Wasserpflanzen aus der Schildberg Subformation der Sulzheim Formation (Oligozän, Rupelium) des Mainzer Beckens (Rheinland-Pfalz, SW-Deutschland). Mainzer naturwissenschaftliches Archiv, 53, xx-xx.

Paudayal, K., Paudel, L. & Uhl, D., 2016. A new occurrence of silicified wood from the Early Cretaceous (Berriasian) Kagbeni Formation of north Central Nepal. The Palaeobotanist, 65, 61-66.

Uhl, D. & Jasper, A., 2016. New data on the flora of the basal Rotliegend Group (Remigiusberg Formation; Gzhelian) in the Saar-Nahe basin (SW-Germany). Fossil Imprint, 72, 239-250.

Uhl, D., Jasper, A., Valerius, T., Manfroi, J., Guerra-Sommer, M. & Ricardi-Branco, F.S., 2016. Hepaticites iporangae Ricardi-Branco, Faria, Jasper, and Guerra-Sommer, 2011 from the Early Permian of the Paraná basin, Brazil, is not a liverwort but a tracheophyte. Journal of Paleontology, 90, 632–639.

Uhl, D., 2016. 35 Abgrenzungsfeld naturwissenschaftliche Forschungssammlung. – In: Walz, M. (Hrsg.), Handbuch Museum – Geschichte, Aufgaben, Perspektiven, 163-165. Verlag J.B. Metzler, Stuttgart.


Abu Hamad, A., Scholze, F., Schneider, J.W., Golubev, V.K., Voigt, S., Uhl, D. & Kerp, H., 2015. First occurrence of the Permian-Triassic enigmatic conchostracan Rossolimnadiopsis Novozhilov, 1958 from the Dead Sea, Jordan – preliminary report. Permophiles, 62, 22-25.

Abu Hamad, A.M.B. & Uhl, D., 2015. A new macroflora from the Middle Jurassic (Bathonian) of NW Jordan and its palaeoenvironmental significance. Neues Jahrbuch für Geologie und Paläontologie, Abh., 278, 109-122.

Favre, A, Päckert, M., Pauls, S., Jähnig, S.C., Uhl, D., Michalak, I., Muellner-Riehl, A.N., 2015. The role of the uplift of the Qinghai-Tibetan Plateau for the evolution of Tibetan biotas. – Biological Reviews, 90, 236-253.

Köcke, V. & Uhl, D., 2015. The leaf assemblage from the Early – Middle Miocene locality Sulzigtobel near Werthenstein (Canton Lucerne, Switzerland). Phytologia Balcanica, 21(2), 99-109.

Kubik, R., Uhl, D. & Marynowski, L., 2015. Evidence of wildfires during deposition of the Upper Silesian Keuper succession. Annales Societatis Geologorum Poloniae, 85, 685-696.

Manfroi, J., Lindner Dutra, T., Gnaedinger, S.C., Uhl, D. & Jasper, A., 2015. The first report of a Campanian palaeo-wildfire in the West Antarctic Peninsula. Palaeogeography, Palaeoclimatology, Palaeoecology, 418, 12-18.

Manfroi, J., Uhl, D., Guerra-Sommer, M., Francischin, H., Martinelli, A.G., Soares, M.B. & Jasper, A., 2015. Extending the database of Permian palaeo-wildfire on Gondwana: charcoal remains from the Rio do Rasto Formation (Paraná basin), Middle Permian, Rio Grande do Sul state, Brazil. Palaeogeography, Palaeoclimatology, Palaeoecology, 436, 77-84.

Moraweck, K., Uhl, D. & Kunzmann, L., 2015. Estimation of late Eocene terrestrial palaeoclimate: contributions from megafloral assemblages from central Germany. Palaeogeography, Palaeoclimatology, Palaeoecology, 433, 247-258.

Philippe, M., Pacyna, G., Wawrzyniak, Z., Barbacka, M., Boka, K., Filipiak, P., Marynowski, L., Theveneard, F. & Uhl, D., 2015. News from an old wood – Agathoxylon keuperianum (Unger) nov. comb. in the Keuper of Poland and France. Review of Palaeobotany and Palynology, 221, 83-91.

Poschmann, M., Wedmann, S., Schindler, T., Uhl, D. & Wuttke, M., in press. Eine Arthropoden/Pflanzen-Taphozönose aus einem pliozänen Kratersee bei Ruppach-Goldhausen, Westerwald (Rheinland-Pfalz, SW-Deutschland). Mainzer naturwissenschaftliches Archiv, 52, 103-117.

Uhl, D., 2015. Preliminary note on fossil flowers and inflorescences from the Late Oligocene of Enspel (Westerwald, W-Germany). Palaeobiodiversity & Palaeoenvironments, 95, 47-53.

Uhl, D., 2015. Das Klima der Pfalz und angrenzender Regionen im Tertiär. In: Diehl, P., Imhoff, A. & Möller, L. (Hrsg.): Wissensgesellschaft Pfalz – 90 Jahre Pfälzische Gesellschaft zur Förderung derWissenschaften. Veröffentlichungen der Pfälzischen Gesellschaft zur Förderung derWissenschaften, 116, 445-454.

Uhl, D., 2015. Wälder wie in Ostasien – der Westerwald vor 25 Millionen Jahren. In: Geiger, M. & Helb, H.-W. (Hrsg.), Naturforschung, Naturschutz und Umweltbildung – 175 Jahre POLLICHA. POLLICHIA Sonderveröffentlichung, 23, 64-65.

Uhl, D. & Voigt, S., 2015. Eine weltweit einmalige Fossilplatte und ihre Bergungsgeschichte. In: Geiger, M. & Helb, H.-W. (Hrsg.), Naturforschung, Naturschutz und Umweltbildung – 175 Jahre POLLICHA. POLLICHIA Sonderveröffentlichung, 23, 54-55.


Abu Hamad, A.M.B., Jasper, A. & Uhl, D., 2014. Wood remains from the Late Triassic (Carnian) Abu Ruweis Formation of Jordan and their palaeoenvironmental significance. – Journal of African Earth Sciences, 95, 168-174.

El Atfy, H., Brocke, R., Uhl, D., Ghassal, B., Stock, A.T. &Littke, R., 2014. Source rock potential and paleoenvironment of the Miocene Rudeis and Kareem formations, Gulf of Suez, Egypt: An integrated palynofacies and organic geochemical approach. International Journal of Coal Geology, 131, 326-343.

Köhler, J. & Uhl, D. (2014). Die Blatt- und Karpoflora der oberoligozänen Fossillagerstätte Enspel (Westerwald, Rheinland-Pfalz, W-Deutschland). – Mainzer naturwissenschaftliches Archiv, Beihefte, 35, 1-87.

Marynowski, L., Kubik, R., Uhl, D. & Simoneit, B.R.T., 2014. Molecular composition of fossil charcoals and its relation to incomplete combustion of wood. Organic Geochemistry, 77, 22-31.

Oppl, S., Hirschmann, S., Schindler, E., Uhl, D., Amler, M.R.W. & Jansen, U., 2014. Alte Schätze finden ein neues Zuhause – Das GeoArchiv Marburg zieht ins Frankfurter Senckenberg-Institut ein. SENCKENBERG – natur – forschung – museum, 144 (9/10), 310-315.

Printzen, C., Türkay, M. & Uhl, D., in press. Zentral – dezentral: Labore bei Senckenberg. SENCKENBERG – natur – forschung – museum, 144 (5/6), 146-151.

Schindler, T., Poschmann, M., Schoch, R.R., Uhl, D. & Voigt, S., 2014. Early Permian amphibians from the construction pit of a German wind power station: ecological and biogeographical inferences. Permophiles, 59, 20-23.

Uhl, D., 2014. Paläobotanische Problematika aus dem Oberoligozän von Enspel (Westerwald, Rheinland-Pfalz, W-Deutschland). Mainzer naturwissenschaftliches Archiv, 51, 45-53.

Uhl, D., 2014. Variability of selected leaf morphological traits in European beech (Fagus sylvatica L.) in relation to climatic factors – some implications for palaeoenvironmental studies. Phytologia Balcanica, 20, 145-153.

Uhl, D., Dolezych, M. & Böhme, M., 2014. Taxodioxylon-like charcoal from the Late Miocene of Western Bulgaria. Acta Palaeobotanica, 54(1), 101-111.

Uhl, D., Hartkopf-Fröder, C., Littke, R. & Kustatscher, E., 2014. Wildfires in the Late Palaeozoic and Mesozoic of the Southern Alps – The Anisian and Ladinian (Mid Triassic) of the Dolomites (Northern Italy). Palaeobiodiversity & Palaeoenvironments, 94, 271-278.

Uhl, D. & Voigt, S. (2014). Ausgewählte Aspekte der Paläontologie des Rotliegend (Oberkarbon – Unterperm) im Saar-Nahe-Becken (SW-Deutschland) – (Exkursion F am 24. April 2014). – Jahresberichte und Mitteilungen des Oberrheinischen Geologischen Vereins, N.F., 96, 105-128.


Abu Hamad, A.M.B., Jasper, A. & Uhl, D., (2013). Charcoal remains from the Mukheiris Formation of Jordan – the first evidence of palaeowildfire from the Anisian (Middle Triassic) of Gondwana. – Jordan Journal of Earth and Environmental Sciences, 5, 17-22.

Böhme, M., Aigstorfer, M., Antoine, P.-O., Appel, E., Havlik, P, Métais, G., Phuc, L.T., Schneider, S., Setzer, F., Tappert, R., Tran, D.N., Uhl, D. & Prieto, J. (2013). Na Duong (Northern Vietnam) – an exceptional window into Eocene ecosystems from South-East Asia. – Zitteliana A, 53, 121-167.

El Atfy, H., Brocke, R. & Uhl, D. (2013a). A fungal proliferation near the probable Oligocene/Miocene boundary, Nukhul Formation, Gulf of Suez, Egypt. – Journal of Micropalaeontology, 32, 183-195.

El Atfy, H., Brocke, R. & Uhl, D. (2013b). Age and Paleoenvironment of the Nukhul Formation, Gulf of Suez, Egypt: Insights from Palynology, Palynofacies and Organic Geochemistry. – GeoArabia, 18, 137-174.

El Atfy, H., Uhl, D. & Stribrny, B. (2013). Mesozoic (Late Jurassic – Early Cretaceous) palynomorphs and charcoal from a sandstone pebble from Quaternary glacial deposits of the North Sea. – Geological Journal, 48, 376-384.

Havlik, P., Aiglstorfer, M., El-Atfy, H. & Uhl, D. (2013). A peculiar bone-bed from the Norian Stubensandstein (Löwenstein-Formation, Late Triassic) of southern Germany and its palaeoenvironmental interpretation. – Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, 269, 321–337.

Jasper, A., Guerra-Sommer, M., Abu Hamad, A.M.B., Bamford, M., Bernardes-de-Oliveira, M.E.C., Tewari, R. & Uhl, D., (2013). The Burning of Gondwana: Permian fires on the Southern Continent – a palaeobotanical approach. – Gondwana Research, 24, 148-160.

Kunzmann, L., Wilde, V., Brocke, R., Dolezych, M., Uhl, D. & El Atfy, H. (2013). Die Innovation von Melbourne: eine fossile Pflanze – ein Name. SENCKENBERG – natur – forschung – museum, 143 (7/8), 222-229.

Roller, S., Uhl, D. & Mosbrugger, V. (2013). Eine Forschergruppe am Thema der Zeit: CAME – Klima- und Umweltforschung in Zentralasien. SENCKENBERG – natur – forschung – museum, 143 (1/2), 6-13.

Uhl, D. (2013). Eine kurze Geschichte des Feuers – Vegetationsbrände in der Erdgeschichte. Biologie in unserer Zeit, 43, 228-235..

Uhl, D. (2013). The paleoflora of Frankenberg/Geismar (NW-Hesse, Germany) – a largely unexplored “treasure chest” of anatomically preserved plants from the Late Permian (Wuchiapingian) of the Euramerican floral province. – In: Lucas, S.G., et al. eds., The Carboniferous-Permian Transition. New Mexico Museum of Natural History and Science. Bulletin, 60, 433-443.

Uhl, D. (2013). Ein Beitrag zur Flora des Zechsteins (Wuchiapingium, Oberperm) von Ilmenau in Thüringen (Deutschland). Semana – Veröffentlichungen des Naturhistorischen Museums Schleusingen, 28, 35-44.

Uhl, D., Schindler, T. & Heidtke, U.H.J. (2013). Paläontologie der Pfalz – regionale Forschung mit internationaler Relevanz. – In: Haneke, J. & Kremb, K. (Hrsg.), Beiträge zur Geologie der Pfalz; Schriftenreihe der Pfälzischen Gesellschaft zur Förderung der Wissenschaften, 110, 97-108.


Abu Hamad, A.M.B., Jasper, A. & Uhl, D. (2012). The record of Triassic charcoal and other evidence for palaeo-wildfires: Signal for atmospheric oxygen levels, taphonomic biases or lack of fuel? – International Journal of Coal Geology, 96-97, 60-71.

Böhme, M., Aiglstorfer, M., Uhl, D. & Kullmer, O. (2012). The Antiquity of the Rhine River: Stratigraphic Coverage of the Dinotheriensande (Eppelsheim Formation) of the Mainz Basin (Germany). – PLoS ONE, 7(5): e36817. doi:10.1371/journal.pone.0036817

Cleal, C.J., Uhl, D., Cascales-Miñana, B., Thomas, B.A., Bashforth, A.R., King, S. & Zodrow, E.L. (2012). Plant biodiversity changes in Carboniferous tropical wetlands. – Earth Science Reviews, 114, 124-155.

Jasper, A., Guerra-Sommer, M., Uhl, D., Bernardes-de-Oliveira, M.E.C., Tewari, R. &, Secchi, M.I., (2012). Palaeobotanical evidence of wildfires in the Upper Permian of India: macroscopic charcoal remains from the Raniganj Formation, Damodar Valley Basin. – The Palaeobotanist, 61, 75-82.

Manfroi, J., Jasper, A., Guerra-Sommer, M. & Uhl, D. (2012). Sub-arborescent lycophytes in coal-bearing strata from the Artinskian (Lower Permian) of the Santa Catarina coalfield (Paraná basin, Santa Catarina, Brazil). Revista Brasileira de Paleontologia, 15(2), 135-140.

Stribrny, B., Sonnewald, M., Türkay, M., Uhl, D., Wilmsen, M. & Wolters, S. (2012). Die Doggerbank – Gerölle erzählen eine Klimageschichte. – SENCKENBERG – natur – forschung – museum, 142 (1/2), 36-43.

Thiel, C., Klotz, S. & Uhl, D. (2012). Palaeoclimate estimates for selected leaf-floras from the Middle Pliocene (Reuverian) of Central Europe based on different palaeobotanical techniques. – Turkish Journal of Earth Sciences, 21, 263-287.

Uhl, D. (2012). Feuer in der Erdgeschichte – nicht erst der Mensch hat gezündelt. – SENCKENBERG – natur – forschung – museum, 142 (1/2), 32-35.

Uhl, D., Butzmann, R., Fischer, T.C., Meller, B. & Kustatscher, E. (2012). Wildfires in the Late Palaeozoic and Mesozoic of the Southern Alps – The Late Permian of the Bletterbach-Butterloch area (Northern Italy). – Rivista Italiana di Paleontologia e Stratigrafia, 118, 223-233.

Uhl, D., Jasper, A. & Schweigert, G., (2012). Palaeo-wildfires in the Late Jurassic (Kimmeridgian) of Western and Central Europe – Palaeoclimatic and palaeoenvironmental significance. – Palaeobiodiversity and Palaeoenvironment, 92, 329-341

Uhl, D., Jasper, A. & Schweigert, G. (2012) . Die fossile Holzgattung Agathoxylon Hartig im Nusplinger Plattenkalk (Ober-Kimmeridgium, Schwäbische Alb). – Archaeopteryx, 30, 16-22.


Herrmann, M. & Uhl, D. (2011). Paläoökologische und biostratigraphische Implikationen der Dinoflagellaten-Zysten aus dem Oligozän (Alzey-Formation) von Kirchheimbolanden (Mainzer Becken, SW-Deutschland). – Mitteilungen der POLLICHIA, 95, 49-54.

Jasper, A., Manfroi, J., Ost Schmidt, E., Machado, N.T.G. & Uhl, D. (2011). Evidências paleobotânicas de incêndios vegetacionais no Afloramento Morro Papaléo, Paleozóico Superior do Rio Grande do Sul, Brasil. Geonomos, 19, 18-27.

Jasper, A., Uhl, D., Guerra-Sommer, M., Abu Hamad, A.M.B. & Machado, N.T.G. (2011). Charcoal remains from a tonstein layer in the Faxinal coalfield, Lower Permian, southern Paraná Basin, Brazil. – Anais da Academia Brasileira de Ciência, 83(2), 471-481.

Jasper, A., Uhl, D., Guerra-Sommer, M., Bernardes-de-Oliveira, M.E.C. & Machado, N.T.G. (2011). Upper Paleozoic charcoal remains from South America: Multiple evidences of fire events in the coal bearing strata of the Paraná Basin, Brazil. – Palaeogeography, Palaeoclimatology, Palaeoecology, 306, 205-218.

Jasper, A., Guerra-Sommer, M., Uhl, D., Machado, N.T.G., Secchi, M.I., Manfroi, J. & Goncalves, C.V., 2011. Análise de carvão vegetal como evidência direta de paleoincêndios vegetacionais: registros do Permiano da Bacia do Paraná. In: Ismar DE SOUZA CARVALHO; Narenda Kumar SRIVASTAVA; Oscar STROHSCHOEN Jr.; Cecília CUNHA LANA. (Org.). Paleontologia: cenários da vida. 1 ed. Rio de Janeiro: Interciência, 4, 22-32.

Schindler, T., Uhl, D., Nungesser, K. & Herrmann, M. (2011). Die Fossilfundstelle Röntgenstraße in Kirchheimbolanden: Rekonstruktion des Landschaftsbilds und Folgerungen zur Biostratigrafie und Paläobiogeografie (Alzey-Formation, Oligozän; Mainzer Becken, SW-Deutschland). – Mitteilungen der POLLICHIA, 95, 79-82.

Teodoridis, V., Mazouch, P., Spicer, R.A. & Uhl, D., (2011). Refining CLAMP – investigations towards improving the Climate Leaf Analysis Multivariate Program. – Palaeogeography, Palaeoclimatology, Palaeoecology, 299, 39-48.

Uhl, D. & Herrmann, M. (2011). Die terrestrische Paläoflora aus dem Oligozän (Alzey-Formation) von Kirchheimbolanden (Mainzer Becken, SW-Deutschland). – Mitteilungen der POLLICHIA, 95, 55-64.

Uhl, D. & Jasper, A. (2011). Und es brannte doch…! Fossile Holzkohle als Nachweis von Paläo-Vegetationsbränden in Perm und Trias – Ein Gelehrtenstreit? – Fossilien, 28 (2), 106-111.

Uhl, D. & Montenari, M. (2011). Charcoal as evidence of palaeo-wildfires in the Late Triassic of SW-Germany. – Geological Journal, 46, 34-41.

Uhl, D., Schindler, T. & Wuttke, M. (2011). Paläoökologische Untersuchungen im Oberoligozän von Norken (Westerwald, Rheinland-Pfalz, W-Deutschland) – Erste Ergebnisse. – Mainzer naturwissenschaftliches Archiv, 48, 115-127.


Ashraf, A.R., Sun, Y.W., Sun Ge, Uhl, D., Mosbrugger, V., LiJie, Herrmann, M. (2010). Triassic and Jurassic palaeoclimate development in the Junggar Basin, Xinjiang, NW China – a review and additional lithological data. – Palaeobiodiversity and Palaeoenvironment, 90, 187-201.

Herrmann, M., Ashraf, A.R., Uhl, D. & Mosbrugger, V. (2010). Die oberoligozäne Palynoflora der Bohrung Enspel 1996/1 (Westerwald, W-Deutschland) – Teil 2: Angiospermen und Paläökologie. – Mainzer geowissenschaftliche Mitteilungen, 38, 9-60.

Poschmann, M., Schindler, T. & Uhl, D. (2010). Fossillagerstätte Enspel – a short review of current knowledge, the fossil association, and a bibliography. – Palaeobiodiversity and Palaeoenvironment, 90, 3-20.

Uhl, D. & Cleal, C.J. (2010). Late Carboniferous Vegetation change in lowland and intramontane basins in Germany. – International Journal of Coal Geology, 83, 318-328.

Uhl, D. & Herrmann, M. (2010). Palaeoclimate estimates for the Late Oligocene taphoflora of Enspel (Westerwald, W-Germany) based on palaeobotanical proxies. – Palaeobiodiversity and Palaeoenvironment, 90, 39-47.

Uhl, D., Jasper, A., Schindler, T. & Wuttke, M. (2010). First evidence of palaeo-wildfire in the early Middle Triassic (early Anisian) Voltzia Sandstone Fossil-Lagerstätte – the oldest post-Permian macroscopic evidence of wildfire discovered so far. – PALAIOS, 25, 837-842.

Wuttke, M., Uhl, D. & Schindler, T. (2010). Fossil-Lagerstätte Enspel – exceptional preservation in an Upper Oligocene maar. – Palaeobiodiversity and Palaeoenvironment, 90, 1-2.


UHL, D. & RAISCH, M., 2009. Die fossile Schachtelhalmbeblätterung Calamariophyllum im Rotliegend des Saar-Nahe Beckens – Fundmitteilung. Mitteilungen der POLLICHIA, 94, 1-4.

CLEAL, C. J., OPLUSTIL, S., THOMAS, B. A., TENCHOV, Y., ABBINK, O. A., BEK, J., DIMITROVA, T., DRABKOVA, J., HARTKOPF-FRÖDER, Ch., VAN HOOF, T., KEDZIOR, A., JARZEMBOWSKI, E., JASPER, K., LIBERTIN, M., MCLEAN, D., OLIWKIEWICZ-MIKLANSINSKA, M., PSENICKA, J., PTAK, B., SCHNEIDER, J. W., SCHULTKA, S., SIMUNEK, Z., UHL, D., WAKSMUNDZKA, M. I., VAN WAVEREN, I., ZODROW, E. L. (2009). Late Moscovian terrestrial biotas and palaeoenvironments of Variscan Euramerica. Netherlands Journal of Geosciences – Geologie en Mijnbouw, 88, 181-278.

HERRMANN, M., ASHRAF, A.R., UHL, D. & MOSBRUGGER, V., in press (2009). Die oberoligozäne Palynoflora der Bohrung Enspel 1996/1 (Westerwald, W-Deutschland) – Teil 1: Sporen und Gymnospermen. Mainzer geowissenschaftliche Mitteilungen, 37: 33-76.

SCHINDLER, T., UHL, D., SCHOCH, R. & WUTTKE, M., 2009. Die Fossilgemeinschaften des Buntsandsteins und des basalen Muschelkalks in der Pfalz – Abbilder einer stufenweisen Floren-/Faunen-Erholung nach der Perm-Trias-Krise? Mitteilungen der POLLICHIA, 94, 11-37.

TEODORIDIS, V, KVACEK, Z. & UHL, D., 2009. Late Neogene palaeoenvironment and correlation of the Sessenheim-Auenheim floral complex. Palaeodiversity, 2: 1-17.


UHL, D., 2008. Die Paläoflora aus dem Rotliegend (Oberkarbon – Unterperm; Meisenheim-Formation; M5) von Niedermoschel (Saar-Nahe-Becken, SW-Deutschland). Mainzer geowissenschaftliche Mitteilungen. 36, 7-36.

UHL, D. & LAUSBERG, S., 2008. Land plant diversity in selected Latest Pennsylvanian ? – Permian deposits from the Saar-Nahe basin (SW-Germany) and the German Zechstein Basin. Studia Geologica Polonica, 129, 81-106. Special issue “Pennsylvanian (Carboniferous) biotas and palaeoenvironments”. Edited by A. Kedzior and C.J. Cleal.

UHL, D., JASPER, A., ABU HAMAD, A.M.B. & MONTENARI, M., 2008. Permian and Triassic wildfires and atmospheric oxygen levels. Proceedings of the WSEAS Conferences – Special Issues, 13, 179-187.

JASPER, A., UHL, D., GUERRA-SOMMER, M. & MOSBRUGGER, V., 2008. New evidence of wildfires in the Late Palaeozoic of South America (Gondwana) – Early Permian, Rio Bonito Formation, Paraná Basin, Rio Grande do Sul State, Brazil. Journal of South American Earth Sciences, 26, 435-444.


UHL, D., ABU HAMAD, A.M.B., KERP, H. & BANDEL, K., 2007. Evidence for palaeo-wildfire in the Late Permian palaeotropics – charcoalified wood from the Um Irna Formation of Jordan. Review of Palaeobotany and Palynology, 144, 221-230.

UHL, D., BOY, J.A. & SCHINDLER, T., 2007. Süßwassermuscheln (Bivalven) – Filtrierer am Gewässergrund. In: SCHINDLER, T. & HEIDTKE, U.H.J. (Eds.) (2007): Kohlesümpfe, Seen und Halbwüsten – Dokumente einer rund 300 Millionen Jahre alten Lebewelt zwischen Saarbrücken und Mainz. POLLICHIA Sonderveröffentlichung, 10, 118-123.

UHL, D., KLOTZ, S., TRAISER, C., THIEL, C., UTESCHER, T., KOWALSKI, E.A. & DILCHER, D.L., 2007. Paleotemperatures from fossil leaves – a european perspective. Palaeogeography, Palaeoclimatology, Palaeoecology, 248, 24-31.

UHL, D., TRAISER, C., GRIESSER, U. & DENK, T., 2007. Fossil leaves as palaeoclimate proxies in the Palaeogene of Spitsbergen (Svalbard). Acta Palaeobotanica, 47 (1), 89-107. Krakau. Special issue dedicated to Zlatko Kvaèek on the occasion of his 70th birthday.

BRUCH, A.A., UHL, D. & MOSBRUGGER, V. 2007. Miocene Climate in Europe – patterns and evolution: A first synthesis of NECLIME. In: BRUCH, A.A., UHL, D. & MOSBRUGGER; V. (eds.), Miocene Climate in Europe – Patterns and Evolution. First Synthesis of NECLIME. Palaeogeography, Palaeoclimatology, Palaeoecology, 253, 1-7.

JASPER, A., GUERRA-SOMMER, M., UHL, D., SALVI, J., KAUFFMANN, M., OSTERKAMP, I.C., & GONCALVES, C.V., 2007. A ocorrência de incêndios vegetacionais durante Paleozóico Superior da Bacia do Paraná (The occurrence of wildfires in the Late Paleozoic of the Paraná Basin). In: Ismar de Souza Carvalho. (Eds.). Paleontologia: cenários da vida. 1 ed. Rio de Janeiro: Interciencia, 2007, 1: 15-23.

KERP, H., NOLL, R. & UHL, D., 2007a. Übersicht über Pflanzengruppen und Vegetationsentwicklung im Permokarbon des Saar-Nahe Beckens. In: SCHINDLER, T. & HEIDTKE, U.H.J. (Eds .) (2007): Kohlesümpfe, Seen und Halbwüsten – Dokumente einer rund 300 Millionen Jahre alten Lebewelt zwischen Saarbrücken und Mainz. POLLICHIA Sonderveröffentlichung, 10, 42-74.

KERP, H., NOLL, R. & UHL, D., 2007b. Vegetationsbilder aus dem saarpfälzischen Permokarbon. In: SCHINDLER, T. & HEIDTKE, U.H.J. (Eds .) (2007): Kohlesümpfe, Seen und Halbwüsten – Dokumente einer rund 300 Millionen Jahre alten Lebewelt zwischen Saarbrücken und Mainz. POLLICHIA Sonderveröffentlichung, 10, 76-109.

MARTINETTO, E., UHL, D. & TARABRA, E., 2007. Leaf physiognomic indications for a moist warm-temperate climate in NW Italy during the Messinian (Late Miocene). In: BRUCH, A.A., UHL, D. & MOSBRUGGER; V. (eds.), Miocene Climate in Europe – Patterns and Evolution. First Synthesis of NECLIME. Palaeogeography, Palaeoclimatology, Palaeoecology, 253, 41-55.

MICHEELS, A., BRUCH, A.A., UHL, D., Utescher, T. & MOSBRUGGER, V. 2007. A Late Miocene climate model simulation with ECHAM4/ML and its quantitative validation with terrestrial proxy data. In: BRUCH, A.A., UHL, D. & MOSBRUGGER; V. (eds.), Miocene Climate in Europe – Patterns and Evolution. First Synthesis of NECLIME. Palaeogeography, Palaeoclimatology, Palaeoecology, 253, 267-286.

NOLL, R. & UHL, D., 2007. Eine krautige Konifere aus einem mittelalterlichen Gemäuer – ein Beitrag zur stratigraphischen Verbreitung von Dicranophyllum hallei REMY& REMY1959 (Coniferales). Mitteilungen der POLLICHIA, 93, 1-6.

POSCHMANN, M. & UHL, D., 2007. Syncaride Krebse – Im Häutungshemd mit Fransenlook. In: SCHINDLER, T. & HEIDTKE, U.H.J. (Eds.) (2007): Kohlesümpfe, Seen und Halbwüsten – Dokumente einer rund 300 Millionen Jahre alten Lebewelt zwischen Saarbrücken und Mainz. POLLICHIA Sonderveröffentlichung, 10, 164-168.

SCHINDLER, T. & UHL, D. (mit Zeichnungen von D. HERR-HEIDTKE), 2007. Zeitliche Entwicklung der Landschaft und der Lebewelt. In: SCHINDLER, T. & HEIDTKE, U.H.J. (Eds .) (2007): Kohlesümpfe, Seen und Halbwüsten – Dokumente einer rund 300 Millionen Jahre alten Lebewelt zwischen Saarbrücken und Mainz. POLLICHIA Sonderveröffentlichung, 10, 304-308.

STEPPUHN, A., MICHEELS, A., BRUCH, A.A., UHL, D., UTESCHER, T. & MOSBRUGGER, V., 2007. The sensitivity of ECHAM4/ML to a double CO2 scenario for the Late Miocene and the comparison to terrestrial proxy data. Global and Planetary Change, 57, 189-212.

TRAISER, C., UHL, D., KLOTZ, S. & MOSBRUGGER, V., 2007. Leaf physiognomy and palaeoenvironmental estimates – an alternative technique based on an European calibration. Acta Palaeobotanica, 47 (1), 183-201. Krakau. Special issue dedicated to Zlatko Kvaèek on the occasion of his 70th birthday.


UHL, D., 2006a. Some considerations on the taphonomy of conifer remains from the Late Permian of Europe. Neues Jahrbuch für Geologie & Paläontologie, Monatshefte, 08/2006, 483-502.

UHL, D., 2006b. “Kornähren, Fliegenfittiche und Holzgraupen” – Ein Beitrag zur Bildung pyritisierter Pflanzenreste aus dem Zechstein von Nordwest-Hessen. Philippia, 12 (4), 347-360.

UHL, D., 2006c. Ein Baumfarn der Gattung Caulopteris LINDLEY et W. HUTTON aus dem Rotliegend (Oberkarbon – Unterperm) des Saar-Nahe-Beckens (SW-Deutschland). Mitteilungen der POLLICHIA, 92, 5-8.

UHL, D., 2006d. Fossil plants as palaeoenvironmental proxies – some remarks on selected approaches. Acta Palaeobotanica, 46 (2), 87-100.

UHL, D. & LEPPER, J., 2006. Anatomically preserved plant remains from the Lower Triassic of N-Germany – a contribution to the understanding of permineralisation processes. Neues Jahrbuch für Geologie & Paläontologie, Monatshefte, 09/2006, 539-566.

UHL, D., BRUCH, A.A., TRAISER, C. & KLOTZ, S., 2006. Palaeoclimate estimates for the Middle Miocene Schrotzburg flora (S-Germany) – A multi-method approach. International Journal of Earth Sciences, 95, 1071-1085.

KLOTZ, S., FAUQUETTE, S., COMBOURIEU-NEBOUT, N., UHL, D., SUC, J.-P., MOSBRUGGER, V., 2006. Seasonality intensification and long-term winter cooling as a part of the Late Pliocene climate development. Earth and Planetary Science Letters, 241, 174-187.


UHL, D., 2005. Die Rotliegend-Pflanzen aus dem Saar-Nahe-Becken (SW-Deutschland) in der Sammlung des Naturhistoriska Riksmuseet in Stockholm (Schweden). Mitteilungen der POLLICHIA, 91, 1-11.

UHL, D. & KERP, H., 2005. Variability of stomatal density and index in the Upper Permian conifer Quadrocladus Mädler – a taphonomical case study. Palaeogeography, Palaeoclimatology, Palaeoecology, 218, 205-215.

BURK, D., UHL, D. & WALTHER, H., 2005. Some aspects of the actuotaphonomy of leaves in stagnant ponds with implications for the formation of fossil leaf deposits – preliminary results. Neues Jahrbuch für Geologie & Paläontologie, Monatshefte, 12/2005, 705-728.

HOECHSTETTER, S., WAHR, J., KNAPP, S., KLOTZ, S., UHL, D., THIV, M., MOSBRUGGER, V., 2005. Muster der Biodiversität von Gefäßpflanzen in Baden-Württemberg – eine geostatistische Analyse abiotischer Einflussfaktoren. Stuttgarter Beiträge zu Naturkunde Serie A (Biologie), 676, 1-25.

MONTENARI, M. & UHL, D., 2005. Mikroanalytische und rasterelektronenmikroskopische Untersuchungen zur Diagenese fossiler Arthropoden in “kontaktmetamorphen” Sedimenten – Erste Ergebnisse. Mitteilungen der POLLICHIA, 91, 13-20.

TRAISER, C., KLOTZ, S., UHL, D. & MOSBRUGGER, V., 2005. Environmental signals from leaves – A physiognomic analysis of European vegetation. New Phytologist, 166, 465-484.


UHL, D., 2004a. A new species of Ullmannia (Coniferophytina: Ullmanniaceae) from the Upper Permian of northern England. Neues Jahrbuch für Geologie & Paläontologie, Monatshefte, 1/2004, 53-64.

UHL, D., 2004b. Typification of the fossil conifer Pseudovoltzia liebeana (GEINITZ) FLORIN. Taxon, 53, 185-186.

UHL, D., 2004c. Anatomy and taphonomy of a coniferous wood from the Zechstein (Upper Permian) of NW-Hesse (Germany). Geodiversitas, 26 (3), 391-401.

UHL, D. & BRANDT, S., 2004. Durchwachsene weibliche Zapfen von Pseudovoltzia liebeana (GEINITZ) FLORIN aus dem Zechstein (Oberperm) von Thüringen. Veröffentlichungen Museum für Naturkunde Schleusingen, 19, 49-54.

UHL, D., NOLL, R., LAUSBERG, S. & STAPF, K.R.G., 2004. Wildfires in the Late Palaeozoic of Central Europe – An overview of the Rotliegend (Upper Carboniferous – Lower Permian) of the Saar-Nahe Basin (SW-Germany). Palaeogeography, Palaeoclimatology, Palaeoecology, 207, 23-35.

SCHINDLER, T., UHL, D., NOLL, R., POSCHMANN, M., BACH, T., HÖHN, W., RAHM, B., SCHWEISS, D., WUTTKE, M., 2004. Erstfunde von Sigillaria-Stämmen in situ in Rotliegend-Ablagerungen (Ober-Karbon bis Unter-Perm) der Nordpfalz (Südwestdeutschland). Neues Jahrbuch für Geologie & Paläontologie, Abhandlungen, 233 (1), 1-26.

UHL, D. & HEIDTKE, U.H.J., 2003. Crossotheca ZEILLER und Caulopteris LINDLEY et W. HUTTON aus dem Rotliegend (Ober-Karbon – Unter-Perm) des Saar-Nahe-Beckens (SW-Deutschland). Neues Jahrbuch für Geologie & Paläontologie, Monatshefte, 6/2003, 363-375.

UHL, D. & KERP, H., 2003. Wildfires in the late Palaeozoic of Central Europe – The Zechstein (Upper Permian) of NW-Hesse (Germany). Palaeogeography, Palaeoclimatology, Palaeoecology, 199, 1-15.

UHL, D. & WALTHER, H., 2003. Aderungsdichte und Blattgröße bei vier Arten der Gattung Acer L. aus SW-Deutschland. Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie, 124, 449-460.

UHL, D., MOSBRUGGER, V., BRUCH, A. & UTESCHER, T., 2003. Reconstructing palaeotemperatures using leaf floras – case studies for a comparison of leaf margin analysis and the coexistence approach. Review of Palaeobotany and Palynology, 126, 49-64.

NOLL, R., UHL, D. & LAUSBERG, S., 2003. Brandstrukturen an Kieselhölzern der Donnersberg Formation (Oberes Rotliegend, Unterperm) des Saar-Nahe Beckens (SW-Deutschland). Veröffentlichungen des Museum für Naturkunde Chemnitz, 26, 63-72.


UHL, D. 2002. Uronectes fimbriatus (Syncarida, Malacostraca) aus dem Rotliegend (Ober-Karbon – Unter-Perm) des Saar-Nahe-Beckens (SW-Deutschland). Mitteilungen der POLLICHIA, 89, 43-56.

UHL, D. & KERP, H., 2002. Die Makroflora des ostthüringischen und sächsischen Zechsteins – Stand der Forschung und Ausblick. Veröffentlichungen Museum für Naturkunde der Stadt Gera, 29, 44-63.

UHL, D. & KERP, H., 2002. Preservation of fossil plants from the Zechstein (Upper Permian) of Central Europe. Freiberger Forschungshefte C 497- Paläontologie, Stratigraphie, Fazies – 10, 29-43.

UHL, D. & MOSBRUGGER, V., 2002. Some data on the potential use of leaf venation density as a palaeoclimatic proxy in leaves from the Permocarboniferous. Neues Jahrbuch für Geologie & Paläontologie, Abhandlungen, 223, 405-432.

UHL, D., WALTHER, H. & KRINGS, M., 2002 . The Paleogene Flora of Hochstetten-Dhaun (Nahe-area, SW-Germany, Rhineland-Palatinate). Feddes Repertorium, 113, 477-491.

UHL, D., WALTHER, H. & MOSBRUGGER, V., 2002. Leaf venation density in angiosperm leaves as related to climatic and environmental conditions — problems and potential for palaeoclimatology. Neues Jahrbuch für Geologie & Paläontologie, Abhandlungen, 224, 49-95.


UHL, D., 2001. Die Zechsteinpflanzen in der Sammlung des Museums für Naturkunde der Stadt Gera. Veröffentlichungen Museum für Naturkunde der Stadt Gera, 28, 153-156.

LEPPER, J. & UHL, D., (2001). Farnreste aus dem Buntsandstein von Bad Karlshafen (Weserbergland) – Fundbericht. Berichte der Naturhistorischen Gesellschaft Hannover, 143 (for 2001), 19-26.

ROTH-NEBELSICK, A., UHL, D., MOSBRUGGER, V. & KERP, H., 2001. Evolution and function of leaf venation architecture: a review. Annals of Botany, 87 (5), 553-566.


UHL, D. & WALTHER, H., 2000. Sun leaf or shade leaf? – Known facts in the light of new data with implications for palaeobotany. Feddes Repertorium, 111 (3-4), 165-174.

UHL, D. & WALTHER, H., 2000. Pflanzenfunde aus dem Oligozän des Nahegebiets (SW-Deutschland, Rheinland-Pfalz) – Vorläufige Mitteilung. Mainzer geowissenschaftliche Mitteilungen, 29, 37-46.

UHL, D., MOSBRUGGER, V. & WALTHER, H., 2000. Stress-induzierte Aberrationen der Blattaderung in Rotbuchen (Fagus sylvatica L.) – Kurze Mitteilung. Feddes Repertorium, 111 (7-8), 493-497.

KRINGS, M., KERP, H. & UHL, D., 2000. Epidermale Anatomie von Odontopteris subcrenulata (Rost) ZEILLER aus dem Stefan (Oberkarbon) von Blanzy-Montceau (Zentralfrankreich). Feddes Repertorium, 111 (7-8), 385-398.


UHL, D., 1999. Syncarids (Crustacaea, Malacostraca) from the Stephanian D (Upper Carboniferous) of the Saar-Nahe-Basin (SW-Germany). Neues Jahrbuch für Geologie & Paläontologie, Monatshefte, 11/1999, 679-697.

UHL, D. & MOSBRUGGER, V., 1999. Leaf venation density as a climate and environmental proxy – a critical review and new data. Palaeogeography, Palaeoclimatology, Palaeoecology, 149 (1-4), 15-26.

UHL, D. & MOSBRUGGER, V., 1999. Variabilität der Aderungsdichte in Laubblättern von Fels-Ahorn (Acer monspessulanum L.). Mitteilungen der POLLICHIA, 86, 99-109.

ASHRAF, A. R., SUN GE, WANG XINFU, UHL, D., LI CHE & MOSBRUGGER, V., 1999. The Triassic-Jurassic boundary in the Junggar basin (NW-China) – Preliminary palynostratigraphic results. Acta Palaeobotanica, Supplement 2, 85-91.


UHL, D., 1997. Erstnachweis der Gattung Bourbonnella (Aeduelliformes, Actinopterygii) aus dem Unteren Rotliegend (Unter-Perm) des Saar-Nahe-Beckens (SW-Deutschland). Mitteilungen der POLLICHIA, 84, 7-13.