For research enquiries, contact Dr Julia Sigwart, for specimen photographs or queries about type material, contact Sigrid Hof.

Mala-FFM_Julia Sigwart
Prof. Dr.  Julia D. Sigwart
Head of Section

Research interests

My group works on the evolution of diversity in molluscs and other marine invertebrates. Morphology shapes how each species responds to environmental change, in the modern world and in the fossil record. We often focus on very strange body forms, like bivalved gastropods, or apparently conservative forms, like chitons. We use tomography (right) to visualise animal adaptations in 3D and understand the world from other organisms‘ points of view.

Understanding evolutionary radiations, survivorship, and the way animals respond to environmental change, require high resolution data from multiple disciplines, and the research infrastructure in museum collections. The work in my research group crosses genetics, morphology, anatomy, neurobiology, physiology, computational modelling, and experimental approaches, to understand species diversity.

Some of these are discussed in my recent book, What Species Mean: A User’s Guide to the Units of Biodiversity, published by CRC/Taylor & Francis in early 2018.

 Teaching & Student opportunities

 We welcome interest from prospective PhD, MSc, BSc, or volunteer and internship projects. Topics include

  • Genomics
  • Systematics of polyplacophoran molluscs
  • Red List assessments of deep sea invertebrates
  • Neural architechture
  • 3D morphology
  • Collections data and collections care

Taxa described

Identifying and naming species is a fundamental aspect of understanding biodiversity. These are the species and genus names of polyplacophoran molluscs named by Dr Sigwart:

Fossil genera

  • Kulindroplax Sutton, Briggs, Siveter, Siveter & Sigwart, 2012
  • Phthipodochiton Sutton & Sigwart, 2012

Living species

  • Leptochiton subrugatus Sirenko & Sigwart, 2021
  • Crassostrea (Magallana) saidii Wong & Sigwart in Sigwart & al., 2021
  • Sakuraeolis arcana Ellis-Diamon, Picton, Tiberica et Sigwart, 2021
  • Hanleyella henrici Sigwart & C. Chen, 2018
  • Callochiton stefaniae Dell’Angelo, Renda, Sosso, Sigwart & Giacobbe, 2017
  • Leptochiton cascadiensis Sigwart & C. Chen, 2017
  • Leptochiton rogeri Sigwart & Sirenko, 2015
  • Leptochiton consimilis Sigwart & Sirenko, 2012
  • Leptochiton angustidens Sigwart & Sirenko, 2012
  • Leptochiton dykei Sigwart & Sirenko, 2012
  • Leptochiton samadiae Sigwart & Sirenko, 2012
  • Leptochiton longisetosus Sigwart & Sirenko, 2012
  • Leptochiton clarki Sigwart & Sirenko, 2012
  • Leptochiton schwabei Sigwart & Sirenko, 2012

Fossil species

  • Kulindroplax perissokomos Sutton, Briggs, Siveter, Siveter & Sigwart, 2012
  • Leptochiton faksensis Sigwart, Anderson & Schnetler, 2007
  • 2020 – present, Head of Section, Malacology, Senckenberg Research Institute
  • 2019 – 2021, NERC Advisory Network (National Environment Research Council UK)
  • 2018 – present, Pool of Experts, United Nations World Ocean Assessment
  • 2018 – present, Elected Trustee, Deep Sea Biology Society
  • 2014 – 2020, Elected Secretary, International Society for Invertebrate Morphology
  • 2010 – 2019, Elected Council Member/Officer, Unitas Malacologica (world mollusc society)
  • 2019 – present, Visiting Professor, Yunnan University, Yunnan Key Laboratory for Palaeobiology, Kunming, China
  • 2010 – present, Honorary Associate Professor, School of University College Dublin, Ireland
  • 2009 – present, Associate / Assistant Professor of Marine Biology, Queen’s University Belfast
  • 2009 – 2019, Director, Queen’s University Belfast, Marine Laboratory, Portaferry
  • 2003 – 2009, Researcher, National Museum of Ireland & University College Dublin (head of Collections-based Biology in Dublin)
  • 2004 – 2008, Ph.D. (part time), Queen’s University Belfast
  • 2001 – 2003 ,Collections manager (Mollusca), American Museum of Natural History
  • 1997 – 2001, B.Sc. (hons), Biology & Mathematics, University of Victoria, Canada



Sigwart, JD (2018): What Species Mean: A User’s Guide to the Units of Biodiversity. — CRC Press / Taylor & Francis. 241 pp.

IUCN Red List global assessments:

Sigwart JD,C Chen, & EA Thomas* (2019): Chrysomallon squamiferum. — The IUCN Red List of Threatened Species 2019: e.T103636217A103636261

Journal articles and book chapters:

127. Vončina, K., Mikkelsen, N.T., Morrow, C., Ang, R. & Sigwart, J.D. (2023): Clarification of the taxonomic status of Acanthochitona discrepans (Brown, 1827) with new data for the North-East Atlantic Acanthochitona (PolyplacophoraAcanthochitonidae). Biodiversity Data Journal, 11: e109554

126. Janssen, R., Neubert, E. & Sigwart, J.D. (2023): Obituary of Hartmut Nordsieck (4 January 1940–24 October 2022). – Archiv für Molluskenkunde, 152 (2):  83–91.

125. Liu, X., Sigwart, J.D., Sun, J. (2023): Phylogenomic analyses shed light on the relationships of chiton superfamilies and shell-eye evolution. — Marine Life Science & Technology.

124. Mamat, N.S., Esa, Y., Wong, N.L.W.S., Mohd Nor, S.A., Abdul Kada,r N., Sigwart, J.D., Arshad, A. (2023): The complete mitogenome of Haliotis asinina (Gastropoda, Haliotidae) from Malaysian waters provides further insights into the phylogeny of the abalone. — New Zealand Journal of Marine and Freshwater Research.

123. Sigwart, J.D., Pollom, R. & Böhm, M. (http://https// The IUCN Species Survival Commission launches a new Red List Authority to assess marine invertebrates. — Oryx, 57 (4): 418-419.

122. Sigwart, J.D., Chen, C., Tilic, E., Vences, M. & Riehl, T. (2023): Why is there no service to support taxonomy? — BioEssays, 2023: 2300070

121. Chen, C. & Sigwart, J.D. (2023): The lost vent gastropod species of Lothar A. Beck. — Zootaxa, 5270 (3): 401-436.

120. Brandt, A., Chen, C., Tandberg, A.H., Miguez‐Salas, O., Sigwart, J.D. (2023): Complex sublinear burrows in the deep sea may be constructed by amphipods. — Ecology and Evolution, 13(3): e9867.

119. Sigwart, J.D., Ogawa, A. & Chen, C. (2023): “Dreamer holothurians” in the north. — Marine Biodiversity. 53(1): 8.

118. Sigwart ,J.D., Brandt, A., Di Franco, D., Briones, E.E., Gerken, S., Gooday, A.J., Grimes, C.J., Głuchowska, K., Hoffmann, S., Jażdżewska, A.M., Kamyab, E., Kelch, A., Knauber, H., Kohlenbach, K., Miguez-Salas, O., Moreau, C., Ogawa, A., Poliseno, A., Santín Muriel, A.,  Tandberg, A.H.S., Theising*F.I., Walter, T., Wölfl, A.C., Chen, C.  (2023): Heterogeneity on the abyssal plains: A case study in the Bering Sea. – Frontiers in Marine Science, 9: 1037482.

117. Thomas, E. A, Sigwart, J.D., Heylar, S. (2022): New evidence for a globally widespread holothurian species at deep-sea reducing environments. – Marine Biodiveristy, 52: 63.

116. McKenna V., Archibald, J.M., Beinart, R., Dawson, M.N., Hentschel, U., Keeling, P.J., Lopez, J.V., Martin-Duran, J.M., Petersen, J.M., Sigwart, J.D., Simakov, O., Sutherland, K.R., Sweet, M., Talbot, T., Thmopson, A.W., Bender, S., Harrison, P.W., Rajan, J., Cochrane, G., Berriman, M., Lawniczak, M., Blaxter, M. (2022): The Aquatic Symbiosis Genomics Project. – Wellcome Open Research, 6: 254.

115. Colomba, M, Sigwart, J.D., Renda, W., Gregorini, A., Soss, M., Dell’Angelo, B. (2022): Molecular analysis of Lepidopleurus cajetanus from the Mediterranean and near Atlantic. – ZooKeys, 1099: 29-40.

114. Stefanoudis, P.V., Biancani, L.M., Cambronero-Solano,S., Clark, M.R., Copley, J.T., Easton, E., Elmer, F., Haddock, S.H., Herrera, S., Iglesias, I.S., Quattrini, A.M. (2021): Moving conferences online: lessons learned from an international virtual meeting. – Proceedings of the Royal Society, B. 288 (1961): 20211769.

113. Thomas, E.A., Böhm, M., Pollock, C., Chen, C., Seddon, M. & Sigwart, J.D. (2021): Assessing the extinction risk of insular, under-studied species in the deep sea. — Conservation Biology.

112.  Stefanoudis, P.V., Biancani, L.M., Cambronero-Solano, S., Clark, M., Copley, J.T, Easton, E.,  Elmer, F., Haddock, S.H.D., Herrera, S., Iglesias, I.S., Quattrini, A. M., Sigwart, J., Yesson, C. & Glover, A.G. (2021): Moving conferences online: lessons learned from an international virtual meeting. — Proceedings of the Royal Society B., 288: 20211769.  doi: 10.1098/rspb.2021.1769

111. Sigwart, J.D., Wong, N.L.W.S. & Esa, Y. (2021): Global controversy in oyster systematics and a newly described species from SE Asia (Bivalvia: Ostreidae: Crassostreinae). — Marine Biodiversity, 51: 83. doi: 10.1007/s12526-021-01203-x

110.  Sirenko, B.I. & Sigwart, J.D. (2021): Leptochiton subrugatus sp. nov. (Mollusca: Polyplacophora) from low boreal waters of the northern Pacific. — Marine Biodiversity, 51: 64. doi: 10.1007/s12526-021-01190-z

109. Ellis-Diamond, D.C., Picton, B.E., Tibiriçá, Y. & Sigwart, J.D. (2021): A new species of Sakuraeolis from Mozambique, described using 3D reconstruction of anatomy and phylogenetic analysis. —  Journal of Molluscan Studies, 87. eyab010. doi:10.1093/mollus/eyab010

108. Ricart, A.M., Gaylord, B., Hill, T.M., Sigwart, J.D., Shukla, P., Ward, M., Ninokawa, A. & Sanford, E. (2021): Seagrass‑driven changes in carbonate chemistry enhance oyster shell growth. — Oecologia.

107. Sigwart, J.D., Lindberg, D.R., Chen, C. & Sun, J. (2021): Molluscan phylogenomics requires strategically elected genomes. — Phil. Trans. R. Soc., B 376: 20200161.

106. Sigwart, J.D. & Sumner-Rooney, L. (2021): Continuous and Regular Expansion of a Distributed Visual System in the Eyed Chiton Tonicia lebruni. — Biol. Bull., 240. DOI: 10.1086/712114

105. Sun, J., Li, R., Chen, C., Sigwart, J.D. & Kocot, K.M. (2021): Benchmarking Oxford Nanopore read assemblers for high-quality molluscan genomes. — Phil. Trans. R. Soc.,  B 376: 20200160.

104. Sigwart, J.D., Blasiak, R., Jaspers, M., Jouffray, J.-B. & Tasdemir, D. (2020): Unlocking the potential of marine biodiscovery. — Natural Product Reports. DOI: 10.1039/d0np00067a

103. Howell K, A Hilario, AL Allcock, D Bailey, M Baker, M Clark, A Colaço, J Copley, E Cordes, R Danovaro, A Dissanayake, E Escobar, P Esquete, A Gallagher, A Gates, S Gaudron, C German, K Gjerde, N Higgs, N Le Bris, L Levin, E Manea, C McClain, L Menot, N Mestre, A Metaxas, R Milligan, A Muthumbi, B Narayanaswamy, S Ramalho, E Ramirez-Llodra, L Robson, A Rogers, J Sellanes, J Sigwart, K Sink, P Snelgrove, P Stefanoudis, P Sumida, M Taylor, A Thurber, R Vieira, H Watanabe, L Woodall, J Xavier (2020): Blueprint for a Decade to Study Deep-Sea Life. — Nature Ecology and Evolution.

102. Thomas, E.A., Liu, R., Amon, D., Copley, J.T., Glover, A.G., Heylar, S.J., Olu, K., Wiklund, H., Zhang, H., & Sigwart, J.D. (2020): Chiridota heheva – the cosmopolitan holothurian. — Marine Biodiversity. 50:110

101. Sigwart J.D., Lundy, M., Dick, J.T.A., Becker, C. (2020): Declining female size at onset of maturity in Nephrops norvegicus in long term surveys (1997-2016). — ICES Journal of Marine Science. doi: 10.1093/icesjms/fsaa185

100. Howell, K.L., Hilário, A., Allcock, A.L., Bailey, D., Baker, M., Clark, M.R., Colaço, A., Copley, J., Cordes, E., Danovaro, R., Dissanayake, A., Escobar-Briones, I.G., Esquete Garrote, P., Gallagher, A., Gates, A.R., Gaudron, S.M., German, C., Gjerde, K., Higgs, N.D., Le Bris, N., Levin, L.A., Manea, E., McClain, C.R., Menot, L., Mestre, N.C., Metaxas, A., Milligan, R.J., Muthumbi, A.W., Narayanaswamy, B.E., Ramalho, S., Ramirez-Llodra, E., Robson, L., Rogers, A.D., Sellanes, J., Sigwart, J.D., Sink, K.J., Snelgrove, P.V., Stefanoudis, P.V., Sumida, P.Y., Taylor, M.L., Thurber, A.R., Vieira, R., Watanabe, H.K., Woodall, L.C. & Xavier, J.R. (2020): A blueprint for an inclusive, global deep-sea Ocean Decade field programme.  ─ Frontiers in Marine Science; doi: 10.3389/fmars.2020.584861

99. Penney B, JD Sigwart, DY Parkinson (2020): Spicules and skeletons: Mantle musculature of two species of dorid nudibranchs (Gastropoda: Nudibranchia: Doridina). — Canadian Journal of Zoology, 98: 777–786, doi:10.1139/cjz-2019-0295

98. Cunningham, E.M., Ehlers, S.M., Dick, J.T.A., Sigwart, J.D., Linse, K., Dick, J.J., Kiriakoulakis, K. (2020): Deep-sea sediments as a sink for microplastic pollution: Evidence from Antarctica and the Southern Ocean. — Environmental Science & Technology.

97. Becker, C., Dick, J.T.A., Cunningham, E.M., Lundy, M., Bell, E., Eagling, L. & Sigwart, J.D. (2020): Ovary resorption in the Norway lobster (Nephrops norvegicus) and its possible causes with special reference to sperm storage. — Helgoland Marine Research, 74: 12.

96. Linse, K. & Sigwart, J.D. & Chen, C. & Krylova, E. (2020): Ecophysiology and ecological limits of symbiotrophic vesicomyid bivalves (Pliocardiinae) in the Southern Ocean. — Polar Biology. 10.1007/s00300-020-02717-z.

95. Ibáñez CM, C Pardo-Gandarillas, J Sellanes, JD Sigwart, BI Sirenko (2020): Phylogenetic position and morphological descriptions of Chiton species from the Southeastern Pacific. — Zoological Journal of the Linnean Society.

94. Sun, J., Chen, C., Miyamoto, N., Li, R., Sigwart, J.D., Xu, T., Sun, Y., Wong, Y.C., Ip, J.C.H., Zhang, W., Lan, Y., Bissessur, D., Watsuji, T., Watanabe, H.K., Takaki, Y., Qiu, J.-W., Takai, K., Qian, P.Y. (2020): The Scaly-foot Snail genome and the ancient origins of biomineralized armour. — Nature Communications, 11: 1657

93. Ashton, E.C., Guist*, S., Roberts, D., Sigwart, J.D. (2020): Effects of environmental factors and husbandry practices on summer mortality events in the cultivated Pacific Oyster Crassostrea gigas in the North of Ireland. — Journal of Shellfish Research, 39(1): 13-20.

92. Cunningham, E.M., Kiriakoulakis, K., Dick, J.T., Kregting, L., Schuchert, P., Sigwart, J.D. (2020): Driven by speculation, not by impact–the effects of plastic on fish species. — Journal of Fish Biology. doi: 10.1111/jfb.14303

91 . Linse, K., Bohrmann, G., Sigwart, J.D. (2019): Eualus amandae (Decapoda: Caridea: Thoridae) is an indicator of active venting sites in the Southern Ocean. — Marine Biodiversity, 49: 2937-2942

90 . Sigwart JD, GJ Vermeij, P Hoyer* (2019): Why do chitons curl into a ball? — Biology Letters, 15(10): 20190429. 

89. Ibáñez CM, Waldisperg M, Torres FI, Carrasco SA, Sellanes J, Pardo-Gandarillas MC, Sigwart JD (2019): Environmental and ecological factors mediate taxonomic composition and body size of polyplacophoran assemblages along the Peruvian Province. — Scientific Reports. 9: 15934.

88 . Cunningham EM*, JD Sigwart (2019): Environmentally accurate microplastic levels and their absence from exposure studies. — Integrative and Comparative Biology.

87 . Morrow C, P Cárdenas, N Boury-Esnault, B Picton, G McCormack, R Van Soest, N Redmond, A Collins, C Maggs, JD Sigwart, AL Allcock (2019): Integrating morphological and molecular taxonomy with the revised concept of Stelligeridae (Porifera: Demospongiae). — Zoological Journal of the Linnean Society.

86 . Sigwart JD, C Chen, EA Thomas*, AL Allcock, M Böhm, M Seddon (2019): Red Listing can protect deep-sea biodiversity. — Nature Ecology & Evolution. 

85 . Wong NLWS, JD Sigwart (2019): Natural history clues to the evolution of bivalved gastropods (Mollusca: Gastropoda: Sacoglossa: Juliidae). — Marine Biodiversity.

84 . Kingston ACN, JD Sigwart, DR Chappell, DI Speiser (2019): Monster or multiplacophoran: A teratological specimen of the chiton Acanthopleura granulata with a valve split into independent and symmetrical halves. — Acta Zoologica.

83 . Sigwart JD, C Chen (2018): A new deep water chiton (Mollusca: Polyplacophora) from hydrothermal vent ecosystems in the Okinawa Trough, Japan. Zootaxa. 4531: 430–436

82. Becker C, EM Cunningham*, JTA Dick, LE Eagling, JD Sigwart (2018): A unified scale for female reproductive stages in Nephrops norvegicus: evidence from macroscopic and microscopic characterisation. — Journal of Morphology.

81. Sigwart JD, C Chen (2018): Comparative oxygen consumption of gastropod holobionts from deep-sea hydrothermal vents in the Indian Ocean. — Biological Bulletin.

80. Sigwart JD, KD Bennett, SM Edie, L Mander, B Okamura, K Padian, Q Wheeler, J Winston, N Yeung (2018): Measuring biodiversity and extinction – present and past. — Integrative and Comparative Biology.

79. Becker C, JTA Dick, EM Cunningham*, C Schmitt, JD Sigwart (2018): The crustacean cuticle does not record chronological age: new evidence from the gastric mill ossicles. — Arthropod Structure and Development.

78. Chen C, K Uematsu, K Linse, JD Sigwart (2018): Cryptic niche switching in a chemosymbiotic gastropod. — Proceedings of the Royal Society B.

77. Sigwart JD, A Garbett* (2018): Biodiversity assessment, DNA barcoding, and the minority majority. — Integrative and Comparative Biology.

76. Sumner-Rooney LH, JD Sigwart (2018): Do chitons have a brain? New evidence for diversity and complexity in the polyplacophoran central nervous system. — Journal of Morphology.

75. Sigwart JD, MK Wicksten, MG Jackson, S Herrera. 2018. Deep-sea video technology tracks a monoplacophoran to the end of its trail. — Marine Biodiversity.

74. Sumner-Rooney LH, I Rahman, JD Sigwart, E Ullrich-Lüter (2018): Whole-body photoreceptor networks are independent of crystalline ‘lenses’ in brittle stars. — Proceedings of the Royal Society B. 285: 20172590.

73. Eagling* LE, EC Ashton, AC Jensen, JD Sigwart, D Murray, D Roberts (2017): Spatial and temporal differences in gonad development, sex ratios and reproductive output influence the sustainability of exploited populations of the European oyster, Ostrea edulis. — Aquatic Conservation.

72. South* J, D Welsh, A Anton, JD Sigwart, JTA Dick (2017): Effects of increasing temperature to the functional response of the intertidal shanny Lipophrys pholis. — Journal of Fish Biology.

71. Dell’Angelo B, W Renda, M Sosso, JD Sigwart, S Giacobbe (2017): A new species of Callochiton (Mollusca: Polyplacophora) from the Strait of Messina (central Mediterranean). — Archiv für Molluskenkunde, 146: 243–250.

70. Ibáñez CM, RD Sepúlveda, JD Sigwart (2017): Comparative allometric variation in intertidal chitons (Polyplacophora: Chitonidae). — Zoomorphology.

69. Sigwart JD, MD Sutton, KD Bennett (2017): How big is a genus? Towards a nomothetic systematics. — Zoological Journal of the Linnean Society.

68. Sigwart JD, Schwabe E (2017): Anatomy of the many feeding types in polyplacophoran molluscs. — Invertebrate Zoology, 14: 205–216

67. Sigwart JD. (2017): Molluscs all beneath the sun, one shell, two shells, more, or none. — Current Biology. 27: R708–R710.

66. Sumner-Rooney LH, JD Sigwart (2017): Lazarus in the museum: resurrecting historic specimens through new technology. — Invertebrate Zoology.

65. Sigwart JD, Chen C. (2017): Life history, patchy distribution, and patchy taxonomy in a shallow-water invertebrate (Mollusca: Polyplacophora: Lepidopleurida). — Marine Biodiversity.

64. Sigwart JD, L McLean* (2017): Contribution of Caulerpa spp. to marine mollusc species richness in the megadiverse Philippines. — Visaya.

63. Sigwart JD, C Chen, L Marsh (2017): Is mining the seabed bad for molluscs? — The Nautilus.

62. Chen C, K Uematsu, K Linse, JD Sigwart (2017): By more ways than one: Rapid convergence in adaptations to hydrothermal vents shown by 3D anatomical reconstruction of Gigantopelta (Mollusca: Neomphalina). — BMC Evolutionary Biology.

61. Geraldi N, C Bertollini*, MC Emmerson, D Roberts, JD Sigwart, NE O’Connor (2017): Aggregations of brittle stars can provide similar ecological roles as mussel reefs. — Marine Ecology Progress Series.

60. Sigwart JD, LH Sumner-Rooney, J Dickey*, N Carey (2017): The scaphopod foot is ventral: more evidence from the anatomy of Rhabdus rectius (Dentaliida: Rhabdidae). — Molluscan Research.

59. Carey N, S Dupont, JD Sigwart (2016): Sea hare Aplysia punctata (Mollusca: Gastropoda) can maintain shell calcification under extreme ocean acidification. — Biological Bulletin.

58. Sigwart JD (2016): The chiton stripe tease. — Marine Biodiversity.

57. Sumner-Rooney* LH, JD Sigwart, L Smith, J McAfee*, ST Williams (2016): Repeated eye reduction events reveal multiple pathways to degeneration in a family of marine snails. — Evolution.

56. Sigwart JD (2016): Deep trees: woodfall biodiversity dynamics in present and past oceans. — Deep Sea Research Part II.

55. Sirenko BI, JD Sigwart, B Dell’Angelo (2016): Hanleya hanleyi Bean in Thorpe, 1844 (Mollusca, Polyplacophora) and the influence of the Gulf Stream System on its distribution. — Ruthenica.

54. Calderwood* J, JD Sigwart (2016): Broken pieces: Can variable ecological interactions be deduced from the remains of crab attacks on bivalve shells? — Lethaia.

53. Sigwart, JD, LH Sumner-Rooney* (2016): Mollusca: Caudofoveata, Monoplacophora, Polyplacophora, Scaphopoda, Solenogastres. — In: Schmidt-Rhaesa, Harzsch, and Purschke, eds., Structure and Evolution of Invertebrate Nervous Systems. Oxford University Press. [book chapter]

52. Green, DS, B Boots, JD Sigwart, S Jiang, C Rocha (2016): Effects of conventional and biodegradable microplastics on a marine ecosystem engineer and nutrient cycling. — Environmental Pollution Bulletin.

51. Sigwart, JD, G Lyons*, A Fink, MA Gutowska, D Murray, F Melzner, JDR Houghton, MY Hu(2016): Elevated pCO2 drives lower growth and yet increased calcification in the early life history of the cuttlefish Sepia officinalis (Mollusca: Cephalopoda). — ICES Journal of Marine Science.

50. Sigwart, JD, PA Green*, SB Crofts (2015): Functional morphology in chitons (Mollusca, Polyplacophora): influences of environment and ocean acidification. — Marine Biology.

49. Sumner-Rooney*,LH, M Schrödl, E Lodde-Bensch, DR Lindberg, M Heß, GP Brennan, JD Sigwart (2015): A neurophylogenetic approach provides new insight to the evolution of Scaphopoda. — Evolution & Development.

48. Cledón, M, JD Núñeza, EH Ocampoa, JD Sigwart (2015): Sexual traits plasticity of the potentially invasive limpet Bostrycapulus odites (Gastropoda: Calyptraeidae) within its natural distribution in South America. — Marine Ecology.

47. Chen*, C, JT Copley, K Linse, AD Rogers, JD Sigwart (2015):. The heart of a dragon: 3D anatomical reconstruction of the ‘scaly-foot gastropod’ (Mollusca: Gastropoda: Neomphalina) reveals its extraordinary circulatory system. — Frontiers in Zoology.

46. Sigwart, JD, BI Sirenko (2015): A new name for the deep-sea chiton Leptochiton clarki Sigwart & Sirenko non Berry (Lepidopleurida: Leptochitonidae). — Zootaxa.

45. Sumner-Rooney*, LH, JD Sigwart (2015): Is the Schwabe organ a retained larval eye? Anatomical and behavioural studies of a novel sense organ in adult Leptochiton asellus (Mollusca, Polyplacophora) indicate links to larval photoreceptors. — PLoS ONE.

44. Lindberg, DR, JD Sigwart (2015): What is the molluscan osphradium? A reconsideration of homology. — Zoologischer Anzeiger.

43. Chen*, C, JT Copley, K Linse, AD Rogers, JD Sigwart (2015): How the mollusc got its scales: convergent evolution of the molluscan scleritome. — Biological Journal of the Linnean Society.

42. Sigwart, JD, DR Lindberg (2015): Consensus and confusion in molluscan trees: Evaluating morphological and molecular phylogenies. — Systematic Biology.

41. Sigwart, JD, C Todt, A Scheltema (2014): Who are the “Aculifera”? Journal of Natural History.

40. Carey*, N, JD Sigwart (2014): Size matters: body size modulates responses to ocean acidification and temperature. — Biology Letters.

39. Kunc, HP, GN Lyons*, JD Sigwart, KE McLaughlin, JDR Houghton (2014): Anthropogenic noise affects behaviour across sensory modalities. — American Naturalist.

38. Sigwart, JD, N Carey* (2014): Grazing under experimental hypercapnia and elevated temperature does not affect the radula of a chiton (Mollusca, Polyplacophora, Lepidopleurida). — Marine Environmental Research.

37. Calderwood*, J, NE O’Connor, JD Sigwart, D Roberts (2014): Determining optimal duration of seed translocation periods for benthic mussel (Mytilus edulis) cultivation using physiological and behavioural measures of stress. — Aquaculture.

36. Carey*, N, S Dupont, B Lundve, JD Sigwart (2014): One size fits all: stability of metabolic scaling under warming and ocean acidification in echinoderms. — Marine Biology.

35. Sumner-Rooney*,LH , JA Murray, SD Cain, JD Sigwart (2014): Do chitons have a compass? Evidence for magnetic sensitivity in Polyplacophora. — Journal of Natural History.

34. Sumner-Rooney*, LH, SD Cain, GP Brennan, JD Sigwart (2014): A test for mucus removal in the chiton Lepidochitona cinerea (Linnaeus, 1767) (Polyplacophora: Chitonida: Ischnochitonidae). — The Veliger.

33. Sigwart, JD, N Carey*, PJ Orr (2014): How subtle are the biases that shape the fidelity of the fossil record? A test using marine molluscs. — Palaeogeography, Palaeoclimatology, Palaeoecology.

32. Sigwart, JD, LH Sumner-Rooney*, E Schwabe, M Hess, G Brennan, M Schroedl (2014): A new sensory organ in “primitive” molluscs (Polyplacophora: Lepidopleurida), and its context in the nervous system of chitons. — Frontiers in Zoology.

31. Sigwart, JD, I Stoeger*, T Knebelsberger, E Schwabe (2013): Chiton phylogeny (Mollusca: Polyplacophora) and placement of the enigmatic species Choriplax grayi (H Adams & Angas, 1864). — Invertebrate Systematics.

30. Stöger*, I, JD Sigwart, Y Kano, T Knebelsberger, BA Marshall, E Schwabe, M Schrödl (2013): The continuing debate on deep molluscan phylogeny: Evidence for Serialia (Mollusca, Monoplacophora + Polyplacophora). — BioMed Research.

29. Ávila, SP, JD Sigwart (2013): New records for the shallow-water chiton fauna (Mollusca, Polyplacophora) of the Azores. — ZooKeys.

28. Morrow*, CC, NE Redmond, BE Picton, RW Thacker, AG Collins, CA Maggs, JD Sigwart, AL Allcock (2013): Molecular phylogenies support homoplasy of multiple morphological characters used in the taxonomy of Heteroscleromorpha (Porifera: Demospongiae). — Integrative and Comparative Biology.

27. Duperron, S, M-A Pottier, N Léger, SM Gaudron, N Puillandre, S Le Prieur, JD Sigwart, J Ravaux, M Zbinden (2013): A tale of two chitons: is habitat specialisation linked to distinct associated bacterial communities? — FEMS Microbiology Ecology.

26. Carey*, N, JD Sigwart, J Richards (2013): Economies of scaling: allometry of metabolism is linked to activity, metabolic rate and habitat. — Journal of Experimental Marine Biology and Ecology.

25. Sutton, MD, DEG Briggs, David J Siveter, Derek J Siveter, JD Sigwart (2012): A Silurian armoured aplacophoran: implications for molluscan phylogeny. — Nature.

24. Carey*, N, A Galkin, P Henrikkson*, J Richards, JD Sigwart (2012): Variation in oxygen consumption among ‘living fossils'(Mollusca: Polyplacophora). — Journal of the Marine Biology Association of the UK.

23. Sigwart, JD, BI Sirenko (2012): Deep-sea chitons from sunken wood in the West Pacific (Mollusca: Polyplacophora: Lepidopleurida): taxonomy, distribution, and seven new species. — Zootaxa.

22. Doonan*, J, G Beatty, JD Sigwart, J Provan (2012): Extensive local-scale gene flow and long-term population stability in the intertidal mollusc Katharina tunicata (Mollusca: Polyplacophora). — Biological Journal of the Linnean Society.

21. Sutton, MD, JD Sigwart (2012): A chiton without a foot. — Palaeontology.

20. Yearsley, JM, JD Sigwart (2011): Larval transport modelling of deep-sea invertebrates points to potential undiscovered populations. — PLoS ONE.

19. Donovan, S.K., Sutton, M.D., Sigwart, J.D. (2011): The last meal of the Late Ordovician mollusc ‘Helminthochitonthraivensis Reed, 1911, from the Lady Burn Starfish Beds, southwest Scotland. — Geological Journal.

18. Sigwart, J.D., Schwabe, E., Saito, H., Samadi, S., Giribet, G. (2011): Evolution in the deep sea: combined analysis of the earliest-derived living chitons using molecules and morphology (Mollusca, Polyplacophora, Lepidopleurida). — Invertebrate Systematics. [2011-2013: this paper was in the top-10 ‘Most Read’ articles in Invertebrate Systematics]

17. Donovan, S.K., Sutton, M.D. & Sigwart, J.D. (2010): Crinoids for lunch? An unexpected biotic interaction from the Upper Ordovician of Scotland. — Geology.

16. Viscardi, P., Sakumoto, M. & Sigwart, J.D. (2010):. How long is a piece of Strix? Methods in measuring and measuring the measurers. — Zoomorphology.

15. Sigwart, J.D. & Leonard, L.M. (2009): Coming out of its shell: Molluscan collections in the National Museum of Ireland. Irish Naturalists’ Journal.

14. Chritz*, K.L., Dyke, G.J., Zazzo, A., Lister, A.M., Monaghan, N.T., & Sigwart, J.D. (2009): Palaeobiology of an extinct Ice Age mammal: Stable isotope and cementum analysis of giant deer teeth. — Palaeogeography, Palaeoclimatology, Palaeoecology.

13. Sigwart, J.D. (2009): Primitive living chitons: Morphological cladistic analysis as a model for character evaluation. — American Malacological Bulletin. Proceedings of the symposium ‘Molluscs as Models for Evolution’ Antwerp, 2007.

12. Sigwart, J.D. (2009): Parasitic foraminifers on a deep-sea chiton (Mollusca, Polyplacophora, Leptochitonidae) from Iceland. — Marine Biology Research.

11. Sigwart, J.D. (2009): The deep-sea chiton Nierstraszella (Mollusca: Polyplacophora: Lepidopleurida) in the Indo-West Pacific: taxonomy, morphology, and a bizarre ectosymbiont. — Journal of Natural History.

10. Sigwart, J.D. (2008): Gross anatomy and positional homology of gills, gonopores, and nephridiopores in “basal” living chitons (Polyplacophora: Lepidopleurina). — American Malacological Bulletin.

9. Sigwart, J.D. (2008): Crystal Creatures: Context for the Dublin Blaschka Congress. — Historical Biology.

8. Sigwart, J.D. & Sutton, M.D. (2007): Deep molluscan phylogeny: Synthesis of palaeontological and neontological data. — Proceedings of the Royal Society B.

7. Voight, J.R. & Sigwart, J.D. (2007): Scarred limpets at hydrothermal vents: Evidence of predation by deep-sea whelks. — Marine Biology.

6. Sigwart, J.D., Schnetler, K.I., Andersen, S.B. (2007): First record of a chiton from the Palaeocene of Denmark (Polyplacophora: Leptochitonidae) and its phylogenetic affinities. — Journal of Systematic Palaeontology.

5. Sigwart, J.D. (2007): The Irish fossil Polyplacophora. — Irish Journal of Earth Sciences.

4. Sigwart, J.D. (2007): Hidden biodiversity: Chitons in Ireland. pp. 18–20 In R Moles, ed. — Proceedings of ESAI/ENVIRON 2005. University of Limerick.

3. Viscardi, P., Sigwart, J.D., Monaghan, N.T. (2006): Climate control in an uncontrollable building. — Collections Forum.

2. Sigwart, J.D., Callaghan, E., Colla*,A., Dyke, G.J., McCaffrey*, S.L., Monaghan, N.T. (2005): Belatedly hatching ornithology collections at the National Museum of Ireland. — Zoologische Mededelingen.

1. Dyke, G.J. & Sigwart, J.D. (2005): A search for a ‘smoking gun’: No need for an alternative to the Linnaean System of classification. pp. 49–65 — In: A Minelli, G Ortalli and G Sanga, eds. Animal Names, proceedings of the International Conference Animal Names organised by Istituto Veneto di Scienze, Lettere ed Arti, Venice. [Book chapter]



Sigrid Hof, Technische Assistentin in der Sektion Malakologie
Mala-FFM_Karl-Otto Nagel
Dr. Karl-Otto Nagel
Research Associate


* 1959
1978 – 1984 Studies of Biology, Technical University Darmstadt
1984 – 1988 Doctoral studies, University (Gh) Kassel; Scholarship: Evangelisches Studienwerk Villigst
1989 – 1991 Postdoc, University of Turin, Italy (Prof. G. Badino): Scholarship: DFG
2001 – 2005 Consultant in EU projects, Bratislava, Slovakia
2012 – 2015, 2019 scientific assistant, Senckenberg Research Institute and Museum of Nature, Frankfurt a/M; funding: DFG, Datz Foundation


chosen publications
(complete directory and downloads on researchgate)


Nagel, K.-O., Schiøtte, T., Gofarov, M.Y., Vikhrev, I.V., Konopleva, E.S. & Bolotov, I.N. (2022):  Re-discovery of the type series of the Indian freshwater mussel Parreysia corrugata (O. F. Müller, 1774) with the designation of the lectotype (Bivalvia: Unionidae:
Parreysiinae). – Journal of Natural History, 56 (5-8): 493-511, DOI: 10.1080/00222933.2022.2063083.

Bolotov, I.N., Pasupuleti, R., Subba Rao, N.V., Unnikrishnan, S.K., Chan, N., Lunn, Z., Win, T., Gofarov, M.Y., Kondakov, A.V., Konopleva, E.S., Lyubas, A.A., Tomilova, A.A., Vikhrev, I.V., Pfenninger, M., Düwel, S.S., Feldmeyer, B., Nesemann, H.F. & Nagel, K.-O. (2022): Oriental freshwater mussels arose in East Gondwana and arrived to Asia on the Indian Plate and Burma Terrane. – Scientific Reports, 12 (1518). DOI: 10.1038/s41598-022-05257-0.

Nagel, K.-O., Bolotov, I. N. & Feldmayer, B. (2021): Die zweite Welle – Sinanodonta lauta – ist (vorerst) noch nicht da. – Mitteilungen der Deutschen Malakozoologischen Gesellschaft, 105: 64-66.

Riccardi, N., Froufe, E., Bogan, A.E., Zieritz, A., Texeira, A., Vanetti, I., Varandas, S., Zaccara, S., Nagel, K.-O. & Lopes-Lima, M. (2020): Phylogeny of European Anodontini (Bivalvia: Unionidae) with a redescription of Anodonta exulcerata. – Zoological Journal of the Linnean Society, zlz136,

Ferreira-Rodríguez, N., Akiyama, Y.B., Aksenova, O.V., Araujo, R., Barnhart, M.C., Bespalaya, Y.V., Bogan, A.E., Bolotov, I.N., Budha, P.B., Clavijo, C., Clearwater, S.J., Darrigran, G., Do, V.T., Douda, K., Froufe, E., Gumpinger, C., Henrikson, L., Humphrey, C.L., Johnson, N.A., Klishko, O., Klunzinger, M.W., Kovitvadhi,S., Kovitvadhi, U., Lajtner, J., Lopes-Lima, M., Moorkens, E.A., Nagayama, S., Nagel, K.-O., Nakano, M., Negishi, J.N., Ondina, P., Oulasvirta, P., Prié, V., Riccardi, N., Rudzīte, M., Sheldon, F., Sousa, S., Strayer, D.L., Takeuchi, M., Taskinen, J., Teixeira, A., Tiemann, J.S., Urbańska, M., Varandas, S., Vinarski, M.V., Wicklow, B.J., Zając, T., Vaughn, C.C. (2019): Research priorities for freshwater mussel conservation assessment. – Biological Conservation 231: 77-87. ISSN 0006-3207 (Available online: 16 January 2019)

Araujo, R., Buckley, D., Nagel, K.-O., García-Jiménez, R. & Machordom, A. (2018): Species boundaries, geographic distribution and evolutionary history of the Western Palaeartic freshwater mussels Unio (Bivalvia, Unionidae). – Zoological Journal of the Linnean Society 182: 275-299, London.

Lopes-Lima M., Sousa R., Geist J., Aldridge D., Araujo R., Bergengren J., Bespalaja Y., Bodis E., Burlakova L., Van Damme D., Douda K., Froufe E., Georgiev D., Gumpinger C., Karatayev, A., Kebapçi Ü., Killeen I., Lajtner J., Larsen B., Lauceri R., Legakis A., Lois S., Lundberg S., Moorkens E., Motte G., Nagel K.-O., Ondina P., Outeiro A., Paunovic M., Prie V., Von Proschwitz T., Riccardi N., Rudzīte M., Rudzītis M., Scheder C., Seddon M., Şereflişan H., Simić V., Sokolova S., Stoeckl K., Taskinen J., Teixeira A., Thielen F., Trichkova T., Varandas S., Vicentini H., Zajac K., Zajac T. & Zogaris S. (2017): Conservation status of freshwater mussels in Europe: state of the art and future challenges. – Biological Reviews 92: 572-607.

Araujo, R., Buckley, D., Nagel, K.-O. & Machordom, A. (2017): Potomida littoralis (Bivalvia, Unionidae) evolutionary history: slow evolution or recent speciation? – Zoological Journal of the Linnean Society 179: 277-299

Nagel, K.-O., Dümpelmann, C. & Pfeiffer, M. (2015): Effective growth cessation in adult Unio crassus Philipsson, 1788 (Bivalvia: Unionidae) from Germany. – Folia Malacologica 23: 309-313, Poznan.

Nagel, K.-O., Castagnolo, L., Cencetti, E. & Moro, G.A. (2007): Notes on reproduction, growth and habitat of Microcondylaea bonellii (Mollusca: Bivalvia: Unionidae) in the torrente Versa (Italy). — Mollusca, 25(1): 41-49, Dresden.

Nagel, K.-O., Šteffek, J. & Vavrová, L’. (2006): Distribution of Unionidae in Slovakia, with notes on their ecology and conservation. — Lauterbornia, 58: 83-96, Dinkelscherben.

Nagel, K.-O. (2004): Observations on the reproductive period of the freshwater mussel Potomida littoralis (Unionidae). — Iberus, 22 (2): 1-8, Gijón.

Nagel, K.-O. & Badino, G. (2001): Population genetics and systematics of European Unionoidea. — In: Ecology and evolutionary biology of the freshwater mussels Unionoida (G. Bauer & K. Wächtler, eds.). — Ecological studies, 145: 51-80. Springer Verlag Berlin, Heidelberg.

Nagel, K.-O. (2000): Testing hypotheses on the dispersal and evolutionary history of freshwater mussels (Mollusca: Bivalvia: Unionidae). — Journal of Evolutionary Biology, 13: 854-865, Oxford.

Nagel, K.-O., Badino, G. & Celebrano, G. (1998): Systematics of European naiades (Bivalvia: Margaritiferidae and Unionidae): a review and some new aspects. — Malacological Review – Supplement 7 (Bivalvia I): 83-104, Ann Arbor, Mich.

Nagel, K.-O., Badino, G. & Alessandria, B. (1996): Population genetics of European Anodontinae (Bivalvia: Unionidae). — Journal of Molluscan Studies, 62: 343-357, London.


Dr. Zeyuan Chen

Research interests

Molluscs are among nature’s greatest innovators, representing some of the most ancient and evolutionary successful animal radiations, with tremendous  diversity in morphology, behavior, and lifestyle. What molecular mechanisms and driving forces produce such great adaptability and diversity? How are novel genes and functions generated, and how do different reproductive strategies and selection affect genetic diversity (genomic heterozygosity), adaptations, and population genomics? These questions fascinate me, and I look forward to finding answers from the Book of Life by ATGC.

  • 03.2023 – present. Bioinformatician. Senckenberg Research Institute and Museum of Natural History Frankfurt am Main
  • 10.2018 – 02.2023. Ph.D. Ludwig-Maximilians-Universität München; Zoologische Staatssammlung München. Studied Molluscan genomics under the supervision of Prof. Dr. Michael Schrödl. Funded by Marie Skłodowska-Curie Actions.
  • 09.2015 – 07.2018. MS. Sichuan University. Studied genomics and population genetics under the supervision of Prof. Dr. Tao Ma.
  • 09.2011 – 07.2015. BS. Quanzhou Normal University. Major Biotechnology.

8. Chen, Z., Doğan, Ö., Guiglielmoni, N., Guichard, A., Schrödl, M. (2022). Pulmonate slug evolution is reflected in the de novo genome of Arion vulgaris Moquin-Tandon, 1855. Sci Rep 12:14226.

7. Chen, Z., Schrödl, M. (2022). How many single-copy orthologous genes from whole genomes reveal deep gastropod relationships? PeerJ 10:e13285181.

6. Doğan, Ö., Schrödl, M. & Chen, Z. (2020). The complete mitogenome of Arion vulgaris Moquin-Tandon, 1855 (Gastropoda: Stylommatophora): Mitogenome architecture, evolution and phylogenetic considerations within Stylommatophora. PeerJ 8:e8603.

5. Chen, Z., Ai, F., Zhang, J., Ma, X. …& Ma, T. (2020). Survival despite isolation, inbreeding and asexual reproduction in the Tropics: insights from the genome of the world’s southernmost poplar (Populus ilicifolia). Plant J 103:430-442.

4. Su, Y., Bai, X., Yang, W., Wang, W., Chen, Z., Ma, J., Ma, T. (2018). Single-base-resolution methylomes of Populus euphratica reveal the association between DNA methylation and salt stress. Tree Genet Genomes 14:86.

3. Wang, W., Liu, H., He, Q., Yang, W., Chen, Z., Wang, M., Su, Y., Ma, T. (2017). Characterization of the complete chloroplast genome of Camptotheca acuminata. Conservation Genet Resour 9:241-243.

2. Chen, Z., Wang, W., Yang, W., Ma, T. (2016). Characterization of the complete chloroplast genome of Populus ilicifolia. Conservation Genet Resour 8:391-393.

1. Yang, W., Wang, W., Zhang, L., Chen, Z., Guo, X., Ma, T. (2016). Characterization of the complete chloroplast genome of Idesia polycarpa. Conservation Genet Resour 8:271-273.

Katarzyna Vončina
PhD Student - SOSA Project
  • 2022 – present. PhD student Senckenberg Research Institute and Museum of Natural History Frankfurt.
  • 2021–2022. Internship in the Department of Invertebrate Zoology in the Institute of Systematics and Evolution of Animals.
  • 2019–2021. M.Sc. Biology, Jagiellonian University, Poland.
  • 2016–2019. B.Sc. Biology, Jagiellonian University, Poland.

13. Vončina, K., Mikkelsen, N.T., Morrow, C., Ang, R. & Sigwart, J.D. (2023): Clarification of the taxonomic status of Acanthochitona discrepans (Brown, 1827) with new data for the North-East Atlantic Acanthochitona (PolyplacophoraAcanthochitonidae). Biodiversity Data Journal, 11: e109554 

12. Stec, D., Vončina, K., Kristensen, R.M. & Michalczyk, Ł. (2022): The Macrobiotus ariekammensis species complex provides evidence for parallel evolution of claw elongation in macrobiotid tardigrades. — Zoological Journal of the Linnean Society, XX: 1–33.

11. Gąsiorek, P., Vončina, K., Nelson, D.R. & Michalczyk, Ł. (2021): The importance of being integrative: a remarkable case of synonymy in the genus Viridiscus (Heterotardigrada: Echiniscidae). — Zoological Letters, 7: 13.

10. Gąsiorek, P., Vončina, K., Kristensen, R.M. & Michalczyk, Ł. (2021): High mountain echiniscid (Heterotardigrada) fauna of Taiwan. — Zoological Studies, 60: 70.

9. Gąsiorek, P., Bochnak, M., Vončina, K. & Michalczyk, Ł. (2021): Phenotypically exceptional Echiniscus species (Heterotardigrada: Echiniscidae) from the Neotropics (Argentina). — Zoologischer Anzeiger, 294: 210–228.

8. Gąsiorek, P., Vončina, K., Ciosek, J., Veloso, M., Fontoura, P. & Michalczyk, Ł. (2021): New Indomalayan Nebularmis species (Heterotardigrada: Echiniscidae) provoke a discussion on its intrageneric diversity. — Zoological Letters, 7: 6.

7. Kiosya, Y., Vončina, K. & Gąsiorek, P. (2021): Echiniscidae in the Mascarenes: the wonders of Mauritius. — Evolutionary Systematics, 5(1): 93–120.

6. Vončina, K., Kristensen, R.M. & Gąsiorek, P. (2020): Pseudechiniscus in Japan: re-description of Pseudechiniscus asper Abe et al., 1998 and description of Pseudechiniscus shintai sp. nov. — Zoosystematics and Evolution, 96(2): 527–536.

5. Bochnak, M., Vončina, K., Kristensen, R.M. & Gąsiorek, P. (2020): Continued exploration of Tanzanian rainforests reveals a new echiniscid species (Heterotardigrada). — Zoological Studies, 59: 18.

4. Gąsiorek, P., Vončina, K., Degma, P. & Michalczyk, Ł. (2020): Small is beautiful: the first phylogenetic analysis of Bryodelphax Thulin, 1928 (Heterotardigrada: Echiniscidae). — Zoosystematics and Evolution, 96(1): 217–236.

3. Gąsiorek, P., Vončina, K. & Michalczyk, Ł. (2020): An overview of the sexual dimorphism in Echiniscus (Heterotardigrada, Echiniscoidea), with the description of Echiniscus masculinus sp. nov. (the virginicus complex) from Borneo. — Zoosystematics and Evolution, 96(1): 103–113.

2. Gąsiorek, P. & Vončina, K. (2019): New Echiniscidae (Heterotardigrada) from Amber Mountain (Northern Madagascar). — Evolutionary Systematics, 3(1): 29–39.

1. Gąsiorek, P., Vončina, K. & Michalczyk, Ł. (2019): Echiniscus testudo (Doyère, 1840) in New Zealand: anthropogenic dispersal or evidence for the ‘Everything is Everywhere’ hypothesis? — New Zealand Journal of Zoology, 46(2): 174–181.

Andre Ampuero-Leon
PhD Student - PHENOME Project

Research interests

Molluscan species have always fascinated me, from the gastropods that inhabit our gardens to the remarkably intelligent and agile cephalopods. The vast array of body morphologies within this group has evolved to suit various marine, freshwater, and terrestrial environments. Delving into their world, one can encounter a broad range of creatures, such as bivalved gastropods, tubular bivalves, worm snails, and numerous other peculiarities.

It is precisely due to this diversity that my primary passion lies in studying the anatomical adaptations of these remarkable organisms. To explore their intricacies, I employ cutting-edge techniques such as 3D reconstruction based on histology and tomography, along with the traditional method of gross dissection.

Taxa described

  • Pomacea penai Ampuero & Ramirez 2023
  • Pomacea reevei Ampuero & Ramirez 2023


  • 2023 – present. PhD student. Senckenberg Research Institute and Museum of Natural History Frankfurt am Main.
  • 2015 – 2017. MS. Universidad Peruana Cayetano Heredia. Marine Sciences.
  • 2009 – 2022. Research assistant. Museum of Natural History – Universidad Nacional Mayor de San Marcos
  • 2006 – 2010. BS. Universidad Nacional Mayor de San Marcos. Biological Sciences with Major in Zoology.



9. Mendivil, A., Ramírez, R., Morin, J., Ramírez, J.L., Siccha-Ramirez, R., Britzke, R., Rivera, F., Ampuero, A., Oliveros, N., & Congrains, C. (2023): Comparative Mitogenome Analysis of two native apple snail species (Ampullariidae, Pomacea) from Peruvian Amazon. — Genes, 14 (9), 1769.

8. Grández, A., Ampuero, A., & Barahona, S. (2023): Peruvian nudibranchs (Mollusca, Gastropoda): an updated literature review-based list of species. — Zookeys, 1176: 117-163.

7. Ampuero, A., Rivera, F., Olórtegui, S., & C. Martel. (2023): Feeding behaviour and food preferences of native and introduced snail species from the Lomas formations, threatened seasonal fog oases. — Pedobiologia, 99-100: 150893.

6. Ampuero, A., & Ramírez R. (2023): Description of two new species of apple snail (Ampullariidae: Pomacea) from Peruvian Amazonia. — Zootaxa, 5258 (1): 76-98.

5. Ramírez R, Ramirez J.L., Rivera F., Justino, S., Solís M., Morín, J., Ampuero A., Mendívil, A., & Congrains A. (2022): Do not judge a snail by its shell: Molecular identification of Pomacea (Ampullariidae) species, with particular reference to the Peruvian Amazonian Giant Apple Snail. — Archiv fur Molluskenkunde, 151 (1): 7-17.

4. Romero, P. E., Calla-Quispe, E., Castillo-Vilcahuaman, C., Yokoo, M., Fuentes-Rivera, H. L., Ramirez, J. L., Ampuero, A., Ibáñez, A. J., & Wong, P. (2021): From the Andes to the desert: 16S rRNA metabarcoding characterization of aquatic bacterial communities in the Rimac river, the main source of water for Lima, Peru. — PLoS One, 16 (4), e0250401.

3. Ramírez, R., Solís, M., Ampuero, A., Morín, J., Jimenez-Vasquez, V., Ramirez, J. L., Congrains, C., Temoche, H., & Shiga, B. (2020): Molecular identification and evolutionary relationships of Pomacea nobilis, basis for the specific authentication of the black apple snail from Peruvian Amazonia. — Revista Peruana de Biología, 27: 139-148.

2. Ampuero, A. & Ramirez R., (2020): Redescription of Helicina weyrauchi Haas, 1948 (Helicinidae) from the Marañon Dry Forest of Northern Peru. — Malacologia, 63 (1): 21-31.

1. Ampuero, A. & Ramirez R., (2017): Anatomy of a Commercial Apple Snail from the Peruvian Amazon. — In Sant’Anna y Hattori B. Amazonian Apple Snails. Nova Publishers. New York. 21-38 pp.


Dr. Ronald Jannsen, ehemaliger Leiter der Sektion Malakologie Frankfurt
Dr. Ronald Janssen
Curator Emeritus


  • 1951 born in Bremen
  • 1964-1970 voluntary helper and employee at the Übersee-Museum Bremen
  • 1966 Promotional prize at the competition for scientific research into local history in Bremen
  • 1968 Prize winner of the competition for scientific research on local history in Bremen
  • 1970 A-levels at the old grammar school in Bremen
  • 1970-1974 Study of geology, palaeontology and zoology at the Philipps-University in Marburg a. d. Lahn
  • 1974 Diploma in geology/palaeontology
  • 1974-1977 Research associate of Prof. Dr. H.-J. Anderson in the DFG priority program “Geological Correlation Research”, subproject: NW-German Tertiary Basin and in the IGCP project 124 “The NW-European Tertiary Basin
  • 1977 Employment with the Senckenberg Natural Research Society as a scientific assistant in the subdepartment Malacology of the FIS (successor to Dr. A. Zilch)
  • 1978 PhD “summa cum laude” with a revision of Upper Oligocene molluscs
  • 1983 Appointment as curator and head of the section of malacology
  • 1997-2015 Deputy head of the Senckenberg School and head of practical training
  • since 2017 retired, volunteer in the subdepartment Malacology (Senckenberg Research Institute Frankfurt)

Committee and department activities

  • 1980-2009 Elected member of the scientific committee of SGN
  • 1981-2016 sole editor of the “Archiv für Molluskenkunde
  • 1981-2008 Editor of the “Mitteilungen der deutschen Malakozoologischen Gesellschaft
  • since 1984 Voting member of the Regional Committee on Northern Neogene Stratigraphy
  • since 1985 member of the board of the German Malacozoological Society (secretary and editor)
  • since 1986 Member of the editorial board of the international malacological journal “Malacologia” (USA)
  • since 1987 Voting member of the Regional Committee on Northern Paleogene Stratigraphy
  • since 1988 Full member of the Tertiary Stratigraphy Subcommission (1976-1987 Corresponding Member)
  • 1988-1989 Member of the organizing committee of the 10th International Congress of Malacologists in Tübingen 1989
  • 1991-1992 Co-organiser of the Symposium on Molluscan Paleontology at the 11th International Congress of Malacologists in Siena 1992
  • since 2003 Member of the board of trustees “Mollusc of the Year”
  • 2012-2016 Chairman of the Tertiary Stratigraphy Subcommission

107. Janssen, R., Neubert, E. & Sigwart, J.D. (2023): Obituary of Hartmut Nordsieck (4 January 1940–24 October 2022). – Archiv für Molluskenkunde, 152 (2):  83–91.

106. Harzhauser, M., Landau, B. & Janssen, R. (2022): The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera. – Zootaxa, 5123 (1): 1-172.

105. Menning, M., Pieńkowski, G., Käding, K.-C., Maletz, J., Kemnitz, H., Kramer, W., Bittmann, F., Hoselmann, C., Janssen, R., Heggemann, H., Hiß, M. & Nitsch, E. (2020): Korrekturen und Ergänzungen zur Stratigraphischen Tabelle von Deutschland 2016 (STD 2016) / Corrections and supplements on the Stratigraphic Table of Germany 2016 (STG 2016). — Zeitschrift der Deutschen Gesellschaft für Geowissenschaften: 6 S.

104. Janssen, R. (2020): Hartmut Nordsieck zum 80. Geburtstag. — Natur Forschung Museum, 150 (4-6): 89.

103. Janssen, R. (2019): Hans-Joachim Anderson 1931-2018. — Geowissenschaftliche Mitteilungen, 75: 93.

102. Albano, P. G., Schnedl, S. M., Janssen, R. & Eschner, A. (2019): An illustrated catalogue of Rudolf Sturany’s type specimens in the Naturhistorisches Museum Wien, Austria (NHMW): Red Sea bivalves. — Zoosystematics and Evolution, 95 (2): 557-598.

101. Albano, P. G., Bakker, P. A. J., Janssen, R. & Eschner, A. (2017): An illustrated catalogue of Rudolf Sturany’s type specimens in
the Naturhistorisches Museum Wien, Austria (NHMW): Red Sea gastropods. — Zoosystematics and Evolution, 93 (1): 45–94.

100.  Janssen, R.; Doppler, G.; Grimm, K.; Grimm, M.; Haas, U.; Hiss, M.; Kothe, A.; Radtke, G.; Reichenbacher, B.; Salamon, M.; Standke, G.; Teipel, U.; Thomas, M.; Uffenorde, H.; Wielandt-Schuster, U. (2018): The Tertiary in the Stratigraphic Table of Germany 2016 (STG 2016). — Zeitschrift der Deutschen Gesellschaft für Geowissenschaften, 169 (2): 267 – 294.

99.  Janssen, R. & Kadolsky, D. (2016): The date of publication of R. A. Philippi’s “Beiträge zur Kenntnis der Tertiaerversteinerungen des nordwestlichen Deutschlands”. — Mitteilungen der Deutschen Malakozoologischen Gesellschaft, 95: 39-44.

98.  Janssen, R. (2015): Welche Bedeutung hat die Historie einer Sammlung? In: Mahr, S., Müller, T., Walker, B. (Hrsg.): Senckenbergs verborgene Schätze. — Kleine Senckenberg-Reihe, 56: 76-77.

97.  Janssen, R. (2015): Geheimnisse einer Lastenträgerschnecke. In: Mahr, S., Müller, T., Walker, B. (Hrsg.): Senckenbergs verborgene Schätze. — Kleine Senckenberg-Reihe, 56: 48-49.

96.  Janssen, A.W., Janssen, R. & van der Voort, J. (2015): Spaniodontella Andrussow in Goloubiatnikow, 1902 – a critical survey of use and validity of the genus, and its relationship to the genus Alveinus Conrad, 1865 (Mollusca, Bivalvia: Glossoidea, Kelliellidae). — Cainozoic research, 15 (1-2): 127-146.

95.  Janssen, R. & Taviani, M. (2015): Taxonomic, Ecological and Historical Considerations on the Deep-Water Benthic Mollusc Fauna of the Red Sea. — In: Rasul, N.M.A & Stewart, I.C.F. (eds.): The Red Sea: 511-529.

94.  Janssen, R. (2015): A review of the Oligocene Limopsidae of the North Sea Basin (Mollusca: Bivalvia). — Geologica Saxonica, 61 (1): 7-33. (printed version)

93.  Janssen, R., Kantor, Y.I. & Stahlschmidt, P. (2015): Identity and generic position of Pleurotoma erythraea Weinkauff 1875 from the Red Sea. — Archiv für Molluskenkunde, 144 (1): 99-104.

92.  Janssen, A.W., Janssen, R., Tracey, S., Vaessen, L.M.B. & van der Voort, J. (2014): Case 3668. Conus antidiluvianus Bruguière, 1792 (Mollusca, Gastropoda, Conidae): proposed conservation of prevailing usage of specific name by setting aside the unidentifiable lectotype and replacing it with a neotype. — Bulletin of Zoological Nomenclature, 71 (4): 223-229.

91.  Janssen, R. (2014): Buchbesprechung: Liverani, V. (2013): The superfamily Stromboidea. Addenda and corrigenda. — Mitteilungen der deutschen malakozoologíschen Gesellschaft, 91: 58.

90.  Janssen, R. (2014): Buchbesprechung: Geiger, D. (2012): Monograph of the Little Slit Shells. — Mitteilungen der deutschen malakozoologíschen Gesellschaft, 91: 55-56.

89.  Janssen, R. (2014): Buchbesprechung: Fraussen, K. & Stratmann, D. (2013): The family Babyloniidae. — Mitteilungen der deutschen malakozoologíschen Gesellschaft, 91: 55.

88.  Janssen, R. (2014): Buchbesprechung: Engl, W. (2012): Shells of Antarctica. — Mitteilungen der deutschen malakozoologíschen Gesellschaft, 90: 56-58.

87.  Janssen, R. (2014): A review of the Oligocene Limopsidae of the North Sea Basin (Mollusca: Bivalvia). — Geologica Saxonica, 61 (7): 7-33. (online version)

86.  Janssen, R. (2014): Günter Wienrich, 12. April 1946 — + 12. Februar 2014. — Cainozoic research, 14 (2): 95-98.

85.  Janssen, R. (2014): In memoriam Günter Wienrich. — Afzettingen, Werkgroep voor Tertiaire en Kwartaire Geologie, 35 (3): 72-73.

84.  Scarponi, D., Daniele, S., Landau, B., Janssen, R., Morgenroth, H. & Della Bella, G. (2014): Lectotype designation for Murex nebula Montagu 1803 (Mangeliidae) and its implications for Bela Leach in Gray 1847. — Zootaxa, 3884 (1): 45-54.

83.  Janssen, R. & Krylova, E. M. (2014): Deep-sea fauna of European seas: An annotated species check-list of benthic invertebrates living deeper than 2000 m in the seas bordering Europe. Bivalvia. — Invertebrate Zoology, 11 (1): 43–82.

82.  Janssen, A. W., Janssen, R., Tracey, S., Vaessen, L. M. B. & van der Voort, J. (2014): History of a marine, Cainozoic gastropod taxon, Conus antidiluvianus Bruguière, 1792 and its nomenclatural implications. — Cainozoic Research, 14 (1): 73-90.

81.  Dijkstra, H. H. & Janssen, R. (2013): Bathyal and abyssal Pectinoidea from the Red Sea and Gulf of Aden (Bivalvia: Propeamussiidae, Entoliidae, Pectinidae). — Archiv für Molluskenkunde, 142 (2): 181-214.

80.  Sonnewald, M. & Janssen, R. (2012): Recent spreading of Xandarovula patula (Mollusca: Gastropoda: Ovulidae) towards the central North Sea suggests a link to rising water temperatures. — Marine Biodiversity Records, 5: 5 S.

79.  Janssen, R. & Krylova, E. M. (2012): Bivalves of the family Vesicomyidae from the Neogene Mediterranean basin (Bivalvia: Vesicomyidae). — Archiv für Molluskenkunde, 141 (1): 87-113, 5 pls.

78.  Scholz, J. & Janssen, R. (2011): Charles Darwin und Senckenberg. — In: Brogiato, H. P. & Kiedel, K.-P. (Hrsg.): Forschen, Reisen, Entdecken. Lebenswelten in den Archiven der Leibnitz-Gemeinschaft: 28-29.

77.  Janssen, R., Zuschin, M. & Baal, C.(2011): Gastropods and their habitats from the northern Red Sea (Egypt: Safaga). Part 2: Caenogastropoda: Sorbeoconcha and Littorinimorpha. — Ann. Naturhist. Mus. Wien, 113 A: 373-509.

76.  Janssen, R. (2011): Schnecken und Muscheln im Senckenberg Forschungsinstitut Frankfurt. — Natur Forschung Museum, 141 (3/4): 96-103.

75.  Janssen, R. (2010): Hartwig Schütt (* 15. Juni 1923, + 14. Dezember 2009. — Archiv für Molluskenkunde, 139 (2): 255-268.

74.  Krylova, E. M., Sahling, H. & Janssen, R. (2010): Abyssogena: a new genus of the family Vesicomyidae (Bivalvia) from deep-water vents and seeps. — Journal of Molluscan Studies, 76: 107-132.

73. Nordhaus, I., Hadipudjana, F. A., Janssen, R. & Pamungkas, J. (2009): Spatio-temporal variation of macrobenthic communities
in the mangrove-fringed Segara Anakan lagoon, Indonesia, affected by anthropogenic activities. — Regional Environmental Change,  9: 291–313.

72.  Janssen, R. & von der Hocht, F. (2009): A remarkable new species of Architectonicidae from the Oligocene of the Lower Rhine embayment (Germany) (Gastropoda: Heterobranchia: Architectonicidae). — Archiv für Molluskenkunde, 138 (2): 205-212, 2 figs., 2 tabs.

71.  Dinapoli, A. & Janssen, R. (2009): Polyplacophora (Mollusca) of the Socotra Archipelago – Systematics and Biogeography. — Fauna of Arabia, 24: 1-46.

70.  Zuschin, M., Janssen, R. & Baal, C. (2009): Gastropods and their habitats from the northern Red Sea (Egypt: Safaga). Part 1: Patellogastropoda, Vetigastropoda and Cycloneritimorpha. — Ann. Naturhist. Mus. Wien, 111 A: 73-158. Wien.

69.  Janssen, R. (2008): Dr. Lore Schwengberg. — Mitt. dtsch. malakozool. Ges., 79/80: 1. Frankfurt a. M.

68.  Cosel, R. von & Janssen, R. (2008): Bathymodioline mussels of the Bathymodiolus (s.l.) childressi clade from methane seeps near Edison Seamount, New Ireland, Papua New Guinea (Bivalvia: Mytilidae). — Archiv für Molluskenkunde, 137 (2): 195-224. Frankfurt a. M.

67.  Schneider, S., Hochleitner, R. & Janssen, R. (2008): A new bivalved gastropod, Candinia lakoniae n. sp. (Sacoglossa: Juliidae) from the Upper Pliocene of Greece (Glykovrysi, SE Peloponnese and Lardos, Rhodes), with a short survey of the geological range and geographic distribution of the family. — N. Jb. Geol. Paläont. Abh., 247: 79-91. Stuttgart.

66.  Janssen, R. & Wienrich, G. (2007): Turridae (654-718, pl. 137-154) — In: Wienrich, G.: Die Fauna des marinen Miozäns von Kevelaer (Niederrhein), 4. Gastropoda ab Mitridae: 640-954, Leiden (Backhuys).

65.  Helama, S., Schöne, B., Kirchhefer, A.J., Nielsen, J.K., Rodland, D.C. & Janssen, R. (2007): Compound response of marine and terrestrial ecosystems to varying climate: pre-anthropogenic perspective from bivalve shell growth increments and tree-rings. — Marine Environmental Research, 63 (3): 185-199.

64.  Krylova, E.M. & Janssen, R. (2006): Vesicomyidae from Edison Seamount (South West Pacific: Papua New Guinea: New Ireland fore-arc basin) (Bivalvia: Glossoidea). — Archiv für Molluskenkunde, 135 (2): 231-261, 6 figs., 4 pls., 8 tabs; Frankfurt a. M.

63. Schütt, H., Jungbluth, J.H. & Janssen, R. (2006): In memoriam Adolf Zilch. — Mitt. dtsch. malakozool. Ges., 76: 39-43. Frankfurt a. M.

62. Janssen, R. (2006): Ansprache zum Gedenken an Adolf Zilch. — Mitt. dtsch. malakozool. Ges., 75: 3-4. Frankfurt a. M.

61. Janssen, R. (2006): Zum Gedenken an Adolf Zilch (1911-2006). — Natur und Museum, 136 (5/6): 136-138. Frankfurt a. M.

60. Janssen, R. (2006): Adolf Zilch * 4. März 1991, + 1. Januar 2006. — Archiv für Molluskenkunde, 135 (1): 1-21, 5 figs.; Frankfurt a. M.

59. Krylova, E.M. & Janssen, R. (2005): Bivalves of the family Vesicomyidae from Edison Seamount (Southwest Pacific, Papua New Guinea, New Ireland Fore-Arc). — Abstracts: 3rd International Symposium on Hydrothermal Vent and Seep Biology, La Jolla, 2005. Scripps Institution of Oceanography, La Jolla: 84.

58. Janssen, R. (2005): Tertiär-Mollusken. — In: Steininger, F. (Hrsg.): Senckenberg 2004: 34-35. Frankfurt a. M.

57. Löffler, S.-B., Janssen, R., Gürs, K.R. & Taviani, M. (2005): The genus Axinus Sowerby, 1821 in the Mediterranean Basin (Bivalvia: Thyasiridae). — Journal of Conchology, 38 (5): 581-597.

56. Eguchi, K., Bui, T.V. & Janssen, R. (2005): Gastropod guests (Prosobranchia: Pupinidae, and Pulmonata: Subulinidae) associated with the ponerine ant Diacamma sculpturatum complex (Insecta: Hymenoptera: Formicidae). — Sociobiology, 45 (2): 307-315.

55. Janssen, R. (2004): Dr. Dieter Röckel und das Senckenbergmuseum Frankfurt a. M. (35-36). – In: Groh, K. & Jungbluth, J.H. (2004): Dr. jur. Dieter Röckel, dem Coniden-Forscher zum 80. Geburtstag. — Mitt. dtsch. malakozool. Ges., 71/72: 31-50. Frankfurt a. M.

54. Janssen, R. (2004): Bedrohte einheimische Weichtierarten als Briefmarkenmotiv. — Natur und Museum, 134 (3): 85-90; Frankfurt a. M.

53. Janssen, R. & Neubert, E. (2004): Registration of the primary types in the mollusc collection of the Forschungsinstitut und Naturmuseum Senckenberg (pp. 96-97, 1 fig.). — In: Berendsohn, W. G. & Oehlschlaeger, S. (Eds.): GBIF-D: German participation in the Global biodiversity Information Facility. Projects in the German National Programme for the Global Biodiversity Information Facility 2003-2006. Status Report 2004: 156 pp. Berlin (ISBN 3-921800-53-6).

52. Neubert, E. & Janssen, R. (2004): Die Typen und Typoide des Natur-Museums Senckenberg, 84: Mollusca: Gastropoda: Pulmonata: Orthalicoidea: Bulimulidae (2), Orthalicidae, Placostylidae. — Archiv für Molluskenkunde, 133: 193-295, 24 Taf.; Frankfurt a. M.

51. Janssen, R. (2004): The family Pickworthiidae in the Middle Miocene of the Paratethys (Gastropoda: Caenogastropoda: Littorinoidea ?). — Courier Forschungs-Institut Senckenberg, 246: 169-186, 5 pls.; Frankfurt a. M.

50. Radtke, G., Griessemer, Th., Grimm, K., Hottenrott, M., Janssen, R., Lippmann, R., Kümmerle, E., Martini, E., Reichenbacher, B., Scholz, J., Spiegler, D. & Vavra, N. (2003): First marine ingression in the Upper Rhinegraben system – Sedimentation, biodiversity, paleogeography of the Pechelbronn Formation (Upper Eocene-Lower Oligocene). — Abstracts, 9th Biannual joint meeting of RCNNS/RCNPS, Mainz 2003: 23.

49. Glaubrecht, M., Hausdorf, B., Janssen, R., Knebelsberger, T. Köhler, F., Neubert, E., Schrödl, M. & Wiese, V. (2003): MOTYGER – MOlluscan TYpes in GERman research collections. — In: Sustainable use and conservation of biological diversity. A challenge for society. Symposium Report, Part A. International Symposium 1.-4. December 2003, Berlin: 301-302. Berlin (BMBF; ISBN 3-00- 012577-9)

48. Schöne, B., Oschmann, W., Rössler, J., Freyre Castro, A, Houk, St., Kröncke, I., Dreyer, W., Janssen, R., Rumohr, H. & Dunca, E. (2003): North Atlantic oscillation dynamics recorded in shells of a long-lived bivalve mollusk. — Geology, 31 (12): 1037-1040.

47. Witte, V., Janssen, R., Eppenstein, A. & Maschwitz, U. (2002): Allopeas myrmekophilos (Gastropoda, Pulmonata), the first myrmecophilous mollusc living in colonies of the ponerine army ant Leptogenys distinguenda (Formicidae, Ponerinae). — Insectes sociaux, 49: 301-305, 1 fig.

46. Janssen, R. & Witte, V.(2002): Allopeas myrmekophilos n. sp., the first snail reported as living in army ant colonies (Gastropoda: Pulmonata: Subulinidae). — Archiv für Molluskenkunde, 131: 211-215, 2 figs; Frankfurt a.M.

45. Janssen, R. (2000): Report on intertidal molluscs from Socotra Island. – In: Apel, M. & Hariri, K. I. (eds.): Conservation and sustainable use of biodiversity of Socotra Archipelago. Marine Habitat, Biodiversity and Fisheries Surveys and Management. Progress Report of Phase III: 97-105; Frankfurt a. M.

44. Fiege, D. & Janssen, R. (1998): Ein Fall von Parasitismus bei Psolus tesselatus, einer ungewöhnlichen Holothurie. — Natur und Museum, 128 (10): 341-344, 4 Abb.; Frankfurt a. M.

43. Janssen, R. (1998): Wilhelm Hohorst . — Natur und Museum, 128 (3): 91-93, 1 Bildnis; Frankfurt a. M.

42. Leistikow, A. & Janssen, R. (1997): A record of the bivalve Guianadesma sinuosum Morrison from the central Amazon basin (Bivalvia: Corbulidae). — Basteria, 61(1/3): 17-22, 3 figs.; Leiden.

41. Janssen, R. (1997): Exponat des Monats Mai: Porzellanschnecken. — Natur und Museum, 127 (5): 161-163, 1 Abb.; Frankfurt a. M.

40. Janssen, R. (1994): Zur Taxonomie und Nomenklatur europäischer Tertiär-Mollusken, 2. Die Gattung Bathybembix im deutschen Oligozän (Gastropoda Prosobranchia: Trochidae). — Archiv für Molluskenkunde, 123 (1/6): 151-159; Frankfurt a. M.

39. Janssen, R. (1993): Closing session of the Symposium on molluscan Palaeontology. — Scripta Geol., Spec. Issue, 2: 435-436; Leiden.

38. Janssen, R. (1993): Taxonomy, evolution and spreading of the turrid genus Spirotropis (Gastropoda: Turridae). — Scripta Geol., Spec. Issue, 2: 237-261, 2 figs., 5 pls.; Leiden.

37. Janssen, R. (1993): Die Typen und Typoide des Natur-Museums Senckenberg, 81. Die Typen der von Dunker 1860/1861 beschriebenen japanischen Meeresmollusken. — Archiv für Molluskenkunde, 122 (Zilch-Festschrift): 403-435, 8 Taf.; Frankfurt a. M.

36. Janssen, R. (1993): Evolution and spreading of the Turrid genus Spirotropis in the European Mio- Pliocene. — Abstracts 11th Intern. Malacol. Congress, Siena 1992: 120; Siena.

35. Janssen, R. (1992): Sektion Malakologie. — In: 175 Jahre Senckenbergische Naturforschende Gesellschaft, Jubiläumsband 2: 78-86, 2 Bildnisse; Frankfurt a. M. (Waldemar Kramer).

34. Janssen, R. (1992): Oskar Boettger 1844 – 1910. — In: 175 Jahre Senckenbergische Naturforschende Gesellschaft, Jubiläumsband 1: 308-311, 1 Bildnis; Frankfurt a. M. (Waldemar Kramer).

33. Janssen, R. (1992): Wilhelm Kobelt 1840 – 1916. — In: 175 Jahre Senckenbergische Naturforschende Gesellschaft, Jubiläumsband 1: 297-299, 1 Bildnis; Frankfurt a. M. (Waldemar Kramer).

32. Janssen, R. (1992): Friedrich Kinkelin 1836-1913. — In: 175 Jahre Senckenbergische Naturforschende Gesellschaft, Jubiläumsband 1: 290-292, 1 Bildnis; Frankfurt a. M. (Waldemar Kramer).

31. Taviani, M., Janssen, R. & Sengupta, B. (1991): The Red Sea unique deep-sea benthic biota and their significance for the basin biogenic sedimentary budget. — Abstracts International Symposium on Red Sea Rifting and Sedimentation, Cairo 1991: 1 p.; Cairo.

30. Janssen, R. (1990): Malakozoologie. — Cour. Forsch.-Inst. Senckenberg, 127: 147-153, 2 Abb.; Frankfurt a. M.

29. Janssen, R. (1989): Benthos-Mollusken aus dem Tiefwasser des östlichen Mittelmeeres, gesammelt während der „METEOR“-Fahrt 5 (1987). — Senckenbergiana maritima, 20 (5/6): 265-276, 1 Tab.; Frankfurt a. M.

28. Janssen, R. (1989): Einige neue Scaphopoden aus dem Oligozän und Miozän des Nordseebeckens. — Meded. Werkgr. tert. kwart. Geol., 26 (3): 79-90, 2 pls.; Leiden.

27. Janssen, R. (1989): Preliminary report on the deep water mollusk fauna of the Red Sea. — Abstr. 10th Internat. Malacol. Congress, Tübingen 1989: 115; Tübingen.

26. Janssen, R. (1989): Larval shells and taxonomy in some Tertiary turrids. — Abstr. 10th Internat. Malacol. Congress, Tübingen 1989: 114; Tübingen.

25. Janssen, R. (1988): Konfusion um zwei Arbeiten von Nyst 1861 über Tertiärmollusken. — Mitt. dtsch. malakozool. Ges., 42: 37-40; Frankfurt a. M.

24. Janssen, R. (1988): Fritz Nordsieck (1906-1984). — Archiv für Molluskenkunde, 118 (4/6): 105-128, 1 Bildnis; Frankfurt a. M.

23. Janssen, R. (1987): Evolution and stratigraphical distribution of Oligocene and Miocene Scaphopoda in the North Sea Basin – a preliminary report. — Meded. Werkgr. tert. kwart. Geol., 24 (1-2): 171-180, 2 Figs.; Leiden.

22. Janssen, R. (1987): Molluscs (Mollusca). Identification marking of molluscs. (S. 55-56) — In: Welsch, H. (Ed.): The marking of specimens of animal and plant species included in the Appendices to the Convention on International Trade in Endangered Species of Wild Fauna and Flora. — Cour. Forsch.- Inst. Senckenberg, 90: 1-132; Frankfurt a. M.

21. Janssen, R. (1986): Mollusken-Assoziationen und Biotope im nordwestdeutschen Oberoligozän (Chattium). Mollusc Associations and Biotops of the Northwest German Upper Oligocene. — Nordwestdeutschland im Tertiär, Teil 1 , Beitr. region. Geol. Erde, 18: 318-341, 9 Abb; Berlin u. Stuttgart (Gebrüder Bornträger).

20. Janssen, R. (1986): Mollusken. Zusammenfassender Bericht über Bearbeitungsstand und bisherige Ergebnisse. Molluscs. Comprehensive Report. — Nordwestdeutschland im Tertiär, Teil 1 , Beitr. region. Geol. Erde, 18: 313-317; Berlin u. Stuttgart (Gebrüder Bornträger).

19. Janssen, R. (1986): Remarks on the Oligocene and Miocene Scaphopods of the North Sea Basin. — Regional Committee on Northern Neogene Stratigraphy, 3rd meeting at Leiden, Documents: 24; Leiden.

18. Janssen, R. (1986): Sektion Malakozoologie. — Cour. Forsch.-Inst. Senckenberg, 85: 79-88, 6 Abb.; Frankfurt a. M.

17. Daniels, C. H. von, Harms, F.-J., Janssen, R. & Uffenorde, H. (1985): Zur Fauna (Mollusca, Ostracoda, Foraminiferida) des Ober-Oligozäns von Freden an der Leine (Süd-Niedersachsen). — Ber. naturhist. Ges. Hannover, 128: 153-170, 1 Abb., 3 Tab.; Hannover.

16. Janssen, R. & Taviani, M. (1985): Factors constraining the composition of the Red Sea deep mollusc fauna. — Abstracts 4th Symposium on Deep-Sea Biology, Hamburg 1985: 1 p.; Hamburg.

15. Janssen, R. (1984): Zur Taxonomie und Nomenklatur europäischer Tertiär-Mollusken, 1. Eine Zeidora– Art aus dem Oligozän des Mainzer Beckens (Prosobranchia: Fissurellidae). — Archiv für Molluskenkunde, 115 (1/3): 119-123, 1 Abb.; Frankfurt a. M.

14. Janssen, R. (1982): Sammlungsschwerpunkte und Forschungsprojekte in der Mollusken-Sektion des Senckenberg-Museums. — Mitt. dtsch. malakozool. Ges., 3 (Suppl.): 18-19; Frankfurt a. M.

13. Janssen, R. (1981): Molluskenfauna [des Rupeltons] (S. 57), Die Molluskenfauna von Niederkaufungen (S. 61-65). — In: Erläut. geol. Karte Hessen 1:25.000, Bl. 4723 Oberkaufungen, 2. Aufl.; Wiesbaden.

12. Janssen, R. (1981): Mollusken-Assoziationen und Biotope im norddeutshen Oberoligozän. — Natur und Museum, 111 (3): 70-78, 8 Abb.; Frankfurt a. M.

11. Janssen, R. (1981): Adolf Zilch zum 70. Geburtstag. — Natur und Museum, 111 (3): 58-59; Frankfurt a. M.

10. Janssen, R. (1979): Berthelinia und Julia aus dem Oligozän von SW-Frankreich (Gastropoda: Sacoglossa). — Archiv für Molluskenkunde, 110 (1/3): 75-79, 3 Abb.; Frankfurt a. M.

9. Janssen, R. (1979): Revision der Bivalvia des Oberoligozäns (Chattium, Kasseler Meeressand). — Geol. Abh. Hessen, 78: 181 S., 1 Abb., 4 Taf.; Wiesbaden.

8. Janssen, R. (1979): Die Mollusken des Oberoligozäns (Chattium) im Nordsee-Becken. 2. Neogastropoda, Euthyneura, Cephalopoda. — Archiv für Molluskenkunde, 109 (4/6): 277-376, Taf. 15-18a; Frankfurt a. M.

7. Janssen, R. (1978): Die Mollusken des Oberoligozäns (Chattium) im Nordsee-Becken. 1. Scaphopoda, Archaeogastropoda, Mesogastropoda. — Archiv für Molluskenkunde, 109 (1/3): 137-227, 1 Abb., Taf. 9-14; Frankfurt a. M.

6. Janssen, R. (1978): Revision der Molluskenfauna des Oberoligocäns (Chattium) im Nordseebecken. (Auszug der wesentlichen Ergebnisse) — 1-24, 2 Abb.; Inaugural-Dissertation, FB Geowissenschaften, Philipps-Universität Marburg/Lahn; Marburg/Lahn.

5. Janssen, R. (1978): Revision der Polyplacophora des Oligozäns in Mitteleuropa. — Archiv für Molluskenkunde, 108 (4/6): 215-235, 1 Abb., Taf. 14-16; Frankfurt a. M.

4. Janssen, R. (1978): Die Scaphopoden und Gastropoden des Kasseler Meeressandes von Glimmerode (Niederhessen). — Geol. Jb., A 41: 195 S., 3 Abb., 3 Tab., 7 Taf.; Hannover.

3. Janssen, R. (1977): Mollusken-Assoziationen im norddeutschen Oberoligocän. — Vortrags-Kurzfassungen Geotagung ‘77, Göttingen, Nachr. dtsch. geol. Ges., 17: 43-44; Göttingen.

2. Huckriede, R. & Janssen, R. (1973): Euryhaline Mollusken im älteren Chattium („Kasseler Meeressand“) von Glimmerode in Niederhessen. — Geologica et Palaeontologica, 7: 189-202, 3 Taf.; Marburg/Lahn.

1. Janssen, R. (1972): Beiträge zur Kenntnis der Bryozoa, Vermes, Crustacea und Echinodermata aus dem nordwestdeutschen Mittel- und Obermiozän. — Veröff. Überseemuseum Bremen, (A) 4 (11): 71-108, 56 Abb.; Bremen.

Dr. Adrienne Jochum
Dr. Adrienne Jochum
Research Associate

Research Interests

 The focus of my research is the phylogeny, biogeography, palaeontology, taxonomy and ecology of terrestrial microgastropods, and especially those belonging to the Carychiidae and Hypselostomatidae. My work emphasises an integrative taxonomic approach including CT and SEM investigation of shells, molecular analyses and histology. Ecological investigation delves into their superficial subterranean and subterranean habitats. A large scale revision of the ellobiid family Carychiidae is in progress.

 New Taxa described

Fossil genera

  • Hirsuticyclus Neubauer, Lida & Jochum 2019

 Fossil species

  • Archaeocyclotus brevivillosus Bichain, Jochum, Pouillon & Neubauer 2022
  • Carychium nashuaense Jochum, Lee & Portell 2023
  • Cretatortulosa gignens Jochum, Yu & Neubauer 2021
  • Galba prima Yu, Neubauer & Jochum 2021
  • Hirsuticyclus electrum Neubauer, Lida & Jochum 2019

Extant genera

  • Angustopila Jochum, Slapnik & Páll-Gergely 2014
  • Dentisphaera Páll-Gergely & Jochum 2017
  • Iberozospeum Jochum, Kneubühler, Prieto & Neubert 2022
  • Koreozospeum Jochum & Prozorova 2015
  • Tonkinospira Jochum, Slapnik & Páll-Gergely 2014

 Extant species 

  • Angustopila bidentata Páll-Gergely & Jochum, 2023
  • Angustopila coprologos Páll-Gergely, Jochum & Hunyadi 2022
  • Angustopila huoyani Jochum, Slapnik & Páll-Gergely 2014
  • Angustopila stochi Páll-Gergely & Jochum 2014
  • Angustopila vandevenderi Páll-Gergely & Jochum, 2023
  • Carychium belizeense Jochum & Weigand 2017
  • Carychium hardiei Jochum & Weigand 2017
  • Carychium zarzaae Jochum & Weigand 2017
  • Carychium panamaense Jochum 2018
  • Dentisphaera maxima Páll-Gergely & Jochum 2017
  • Iberozospeum costulatum Prieto & Jochum 2022
  • Iberozospeum praetermissum (Jochum, Prieto & De Winter 2019)
  • Iberozospeum vasconicum (Prieto, De Winter, Weigand, Gómez & Jochum 2015)
  • Iberozospeum zaldivarae (Prieto, De Winter, Weigand, Gómez et Jochum 2015)
  • Koreozospeum nodongense Lee, Prozorova & Jochum 2015
  • Laoennea renouardi Jochum & Wackenheim 2020
  • Tonkinospira tomasini Páll-Gergely & Jochum 2017
  • Zospeum amplioscutum Jochum & Ruthensteiner 2024
  • Zospeum biokovoense Jochum & Ruthensteiner 2024
  • Zospeum bucculentum Inäbnit, Jochum & Neubert 2019
  • Zospeum clathratum Inäbnit, Jochum & Neubert 2019
  • Zospeum constrictum Jochum & Ruthensteiner 2024 
  • Zospeum dubokidoense Jochum & Ruthensteiner 2024
  • Zospeum gittenbergeri Jochum, Prieto & De Winter 2019
  • Zospeum intermedium Jochum & Ruthensteiner 2024
  • Zospeum kolbae Jochum, Inäbnit, Kneubühler & Ruthensteiner 2024
  • Zospeum manitaense Inäbnit, Jochum & Neubert 2019
  • Zospeum neuberti Jochum & Ruthensteiner 2024
  • Zospeum njegusiense Jochum & Ruthensteiner 2024
  • Zospeum njunjicae Jochum, Schilthuizen & Ruthensteiner 2024 
  • Zospeum pagodulum Inäbnit, Jochum & Neubert 2019
  • Zospeum robustum Inäbnit, Jochum & Neubert 2019
  • Zospeum simplex Inäbnit, Jochum & Neubert 2021
  • Zospeum tortuosum Jochum & Ruthensteiner 2024
  • Zospeum tumidum Jochum, Schilthuizen & Ruthensteiner 2024 


  • 2017 – present: Senckenberg Honorary Research Scientist
  • 2015 – present: Research Scientist, Malacology, Natural History Museum Bern, Switzerland
  • 2014 – 2015: Ph.D. (summa cum laude) University of Bern, Bern, Switzerland
  • 2010 – 2014: Ph.D. research studies, Goethe University, Frankfurt am Main, Germany
  • 2010: Palaeontological practicum in Malacology, Senckenberg Research Institute
  • 2007 – 2010: Research Scientist, Goethe University, Frankfurt am Main, Germany
  • 1978 – 1982: B. Sci. Fisheries & Oceanographic Sciences, University of Washington, WA., USA
  • 1977 – 1978: University of San Diego, Ca., USA

42. Jochum, A., Michalik, P., Inäbnit, T., Kneubühler, J., Slapnik, R., Vrabec, M., Schilthuizen, M. & Ruthensteiner, B. (2024): 3D X-ray microscopy (Micro-CT) and SEM reveal Zospeum troglobalcanicum Absolon, 1916 and allied species from the Western Balkans (Ellobioidea: Carychiidae). – European Journal of Taxonomy, 926: 1-62. 

41. Jochum, A., Bochud, E., Haberthür, D., Lee, H.G., Hlushchuk, R. & Portell, R.W. (2023): Fossil Carychiidae (EupulmonataEllobioidea) from the Lower Pleistocene Nashua Formation of Florida, with the description of a new species. Zookeys 1167: 89-107.

40. Páll-Gergely, B., Hunyadi, A., Vermeulen, J.J., Grego, J., Sutcharit, C., Reischütz, A., Dumrongrojwattana, P., Botta-Dukát, Z., Örstan, A., Fekete, J., & Jochum, A. (2023): Five times over: 42 new Angustopila species highlight Southeast Asia’s rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae). – ZooKeys 1147: 1-177.

39. Bichain, J.-M., Jochum, A., Pouillon, J.-M. & Neubauer, T.A. (2022): Archaeocyclotus brevivillosus sp. nov., a new cyclophorid land snail (Gastropoda: Cyclophoroidea) from mid-Cretaceous Burmese amber. – Cretaceous Research 140, December 2022, 105359.

38. Páll-Gergely, B., Jochum, A., Vermeulen, J. J., Anker, K., Hunyadi, A., Örstan, A., Szabó, A., Dányi, L. & Schilthuizen, M. (2022): The world’s tiniest land snails from Laos and Vietnam (Gastropoda, Pulmonata, Hypselostomatidae). – Contributions to Zoology: 1-17.  doi:10.1163/18759866-bja10025

37. Kneubühler, J., Jochum, A., Prieto, C.E. & Neubert, E. (2022): Molecular investigation and description of Iberozospeum n. gen., including the description of one new species (Eupulmonata, Ellobioidea, Carychiidae). – Organisms Diversity & Evolution 22: 61–92.

36. Jochum, A., Yu, T. & Neubauer, T.A. (2022): Reply to Jiang et al’s comments on the article “Mother snail labors for posterity in bed of mid-Cretaceous amber” by Jochum, Yu and Neubauer (2021) Gondwana Research 97, 68–67. – Gondwana Research 101: 24–25.

35. Parlov, N. & Jochum, A. (2021): A malacofaunistic study of the Wellbach pond, Taunus Nature Park, Hesse. Mitteilungen der Deutschen Malakozoologischen Gesellschaft 106: 37–42.

34. Inäbnit, T. , Jochum, A., Slapnik, R. & Neubert, E. (2021): New genetic data reveals a new species of Zospeum in Bosnia (Gastropoda, Ellobioidea, Carychiinae). – ZooKeys 1071: 175–193.

33. Chen, Z.-Y. & Jochum, A. (2021): Comments on “A new species of land snail from the genus Diplommatina Benson, 1849 (Gastropoda, Caenogastropoda, Diplommatinidae) from Sikkim Himalaya, North East India” by N.K. Das & N.A. Aravind, Molluscan Research 41 (3), 262–268, 2021. – Molluscan Research.

32. Jochum, A., Yu, T. & Neubauer, T.A. (2021): Mother snail labors for posterity in bed of mid-Cretaceous amber. Gondwana Research, 97: 68-72.

31. Yu, T., Neubauer, T.A. & Jochum, A. (2021): First freshwater gastropod preserved in amber suggests long-distance dispersal during the Cretaceous Period. Geological Magazine,

30. Slapnik, R., Jochum, A. (2021): Mollusca: Eupulmonata, 189-197. In: Ozimec, R. (ed.), Vjetrenica: cave biodiversity hotspot of the Dinarides (Vjetrenica: centar špiljske bioraznolikosti Dinarida), JP Vjetrenica – ADIPA, 356 pp., Ravno. 

29. Jochum, A., Bochud, E., Favre, A. Ferrand, M., Wackenheim, Q. (2020): A new species of Laoennea microsnail (Stylommatophora, Diapheridae) from a cave in Laos. Subterranean Biology 36: 1–9. doi: 10.3897/subtbiol.36.58977.

28. Páll-Gergely, B., Grego, J., Vermeulen, J.J., Reischütz, A., Hunyadi, A. & Jochum, A. (2019): New Tonkinospira Jochum, Slapnik & Páll-Gergely, 2014 species from Laos and Vietnam (Gastropoda: Pulmonata: Hypselostomatidae). The Raffles Bulletin of Zoology 67: 517535.

 27. Neubauer T.A., Xing, L. & Jochum A (2019): Land Snail with Periostracal Hairs Preserved in Burmese Amber. iScience 20: 567–574.

 26. Jochum A., Yu T, Neubauer TA (2019). First record of the Paleozoic land snail family Anthracopupidae in the Lower Jurassic of China and the origin of Stylommatophora. Journal of Paleontology: 1–7.

 25. Neubauer TA, Páll-Gergely B, Jochum A, Harzhauser M (2019). Striking case of convergence – Alleged marine gastropods in Cretaceous Burmese amber are terrestrial cyclophoroids. Comment on Yu et al. Palaeoworld 28: 572–575.

 24. Inäbnit T, Jochum A, Kampschulte M, Martels G, Ruthensteiner B, Slapnik R, Nesselhauf C, Neubert E (2019). An integrative taxonomic study reveals carychiid microsnails of the troglobitic genus Zospeum in the Eastern and Dinaric Alps (Gastropoda, Ellobioidea, Carychiinae). Organisms Diversity & Evolution 19: 135–177.

 23. Jochum A, Prieto C, Kampschulte M, Martels G, Ruthensteiner B, Vrabec M, Dörge DD, De Winter AJ (2019). Re-evaluation of Zospeum schaufussi von Frauenfeld, 1862 and Z. suarezi Gittenberger, 1980, including the description of two new Iberian species using Computer Tomography (CT) (Eupulmonata, Ellobioidea, Carychiidae). ZooKeys 835: 65–86.

 22. Jochum A, Ruthensteiner B, Kampschulte M, Martels G, Kneubühler J, Favre A (2018). Fulfilling the taxonomic consequence after DNA Barcoding: Carychium panamaense sp. n. (Eupulmonata, Ellobioidea, Carychiidae) from Panama is described using computed tomographic (CT) imaging. ZooKeys 795: 1–12.

 21. Gittenberger E, Leda P, De Winter AJ, Jochum A (2017). First record of Carychium in Bhutan (Gastropoda, Ellobioidea). Basteria 81 (4-6): 107–110.

 20. Adrienne Jochum & Adrien Favre (2017). First record of the sea slug Stylocheilus striatus (Quoy & Gaimard, 1825) (Anaspidea, Aplysiidae) and swarming behavior for Bazaruto Archipelago, Mozambique with the first record of Pleurobranchus forskalii Rüppel & Leuckart, 1828 (Nudipleaura, Pleurobranchidae) for Bazaruto Island (Gastropoda, Heterobranchia). Check List 13 (5): 435–441.

 19. Adrienne Jochum, Alexander M. Weigand, Estee Bochud, Thomas Inäbnit, Dorian D. Dörge, Bernhard Ruthensteiner, Adrien Favre, Gunhild Martels & Marian Kampschulte (2017). Three new species of Carychium O.F. Müller, 1773 from the Southeastern USA, Belize and Panama are described using computer tomography (CT) (Eupulmonata, Ellobioidea, Carychiidae). ZooKeys 675: 97–127.

 18. Barna Páll-Gergely, Adrienne Jochum & Takahiro Asami (2017). Three new species of Hypselostomatidae (Gastropoda: Pulmonata) from Cong Troi Cave, Northern Vietnam. Acta Zoologica Academiae Scientiarum Hungaricae 63 (3): 327–341.

 17. Jochum A, Slapnik R, Klussmann-Kolb A, Páll-Gergely B, Kampschulte M, Martels, G, Vrabec M, Nesselhauf C & Weigand AM (2015). Groping through the black box of variability: An integrative taxonomic and nomenclatural re-evaluation of Zospeum isselianum Pollonera, 1887 and allied species using new imaging technology (Nano-CT, SEM), conchological, histological and molecular data (Ellobioidea: Carychiidae). Subterranean Biology 16: 123−165.

 16. Jochum A, Prozorova L, Sharyi-ool M & Páll-Gergely B (2015). A new member of troglobitic Carychiidae, Koreozospeum nodongense gen. et sp. nov. (Gastropoda, Eupulmonata, Ellobioidea) is described from Korea. ZooKeys 517: 39−57.

 15. Páll-Gergely B, Hunyadi A, Jochum A & Asami T (2015). Seven new hypselostomatid species from China, including the World’s smallest land snail (Gastropoda: Pulmonata: Orthurethra). ZooKeys 523: 31−62.

 14. Jochum A, Neubauer TA & Harzhauser M (2015). Microstructural details in shells of the gastropod genera Carychiella and Carychium of the Middle Miocene. Lethaia 49: 87−101.

 13. Jochum A, De Winter AJ, Weigand AM, Gómez B, & Prieto C (2015). Two new species of Zospeum Bourguignat 1856 from the Basque-Cantabrian Mountains, Northern Spain (Eupulmonata, Ellobioidea, Carychiidae). ZooKeys 483: 81−96.

 12. Jochum A, Slapnik R, Kampschulte M, Martels G, Heneka M & Páll-Gergely B (2014). A review of the microgastropod genus Systenostoma Bavay & Dautzenberg, 1908 and a new subterranean species from China (Gastropoda, Pulmonata, Hypselostomatidae). ZooKeys 410: 23−40.

11. Weigand AM, Jochum A & Klussmann-Kolb A (2014). DNA barcoding cleans house through the Carychiidae (Eupulmonata, Ellobioidea). American Malacological Bulletin 32:236−245.

10. Malkowsky Y & Jochum A (2014). Three-dimensional reconstructions of pallial eyes in Pectinidae (Mollusca: Bivalvia). Acta Zoologica (Stockholm): 1−7.

9. Weigand AM, Jochum A, Slapnik R, Schnitzler J, Zarza E & Klussmann-Kolb A (2013). Evolution of microgastropods (Ellobioidea, Carychiidae): integrating taxonomic, phylogenetic and evolutionary hypotheses. BMC Evolutionary Biology 13:1−23.

 8. Weigand AW, Pfenninger M, Jochum A & Klussmann-Kolb A (2012). Alpine Crossroads or origin of genetic diversity? Comparative phylogeography of two sympatric microgastropod species. PLoS ONE 7: 1−23.

 7. Weigand AM, Götze M-C & Jochum A (2012). Outdated but established?! Conchologically driven species delineations in microgastropods (Carychiidae, Carychium). Organisms Diversity & Evolution 12: 377–386.

 6. Jochum A, Weigand AM, Slapnik R, Valentinčič & Prieto CE (2012). The microscopic ellobioid, Zospeum Bourguignat, 1856 (Pulmonata, Ellobioidea, Carychiidae) makes a big debut in Basque Country and the province of Burgos (Spain). MalaCo 8: 400−403.   

5. Jochum A (2011). Evolution and diversity of the troglobitic Carychiidae – A morphological and phylogenetic investigation of the terrestrial ellobioid genera, Carychium and Zospeum. The Malacologist 57: 16−18.

 4. Weigand AM, Jochum A, Pfenninger M, Steinke D & Klussmann-Kolb A (2011). A new approach to an old conundrum – DNA barcoding sheds new light on phenotypic plasticity and morphological stasis in microsnails (Gastropoda, Pulmonata, Carychiidae). Molecular Ecology Resources 11: 255−265

3. Weigand AM & Jochum A (2010). Mollusca, Gastropoda, Pulmonata, Carychium minimum O.F. Müller, 1774: Filling gaps. New population record for the State of New York, Northeastern United States. CheckList 6: 517−518.

 2. Weigand AM & Jochum A (2010) Carychium minimum Müller, 1774 Microsnail hitchhikes across the Atlantic. Taxonomy in Action. BioNet 14.

 1. Oetken M, Jagodzinski LS, Vogt C, Jochum A & Oehlmann J (2009). Combined effects of chemical and temperature stress on Chironomus riparius populations with differing genetic variability. Journal of Environmental Science and Health, Part A 44: 955−962.






Dr. Eike Neubert
Research Associate

Kurator für Malakologie am Naturhistorischen Museum Bern

Ian F. Smith
Volunteer - MarInvert

Senckenberg – Marinvert

Photographic guide to the identification and biology of European marine invertebrates

Images and Texts by Ian Frank Smith

Ellen Svensson-Stenbrock
Silvia Kühn-Krämer