Nova Supplementa Entomologica, SDEI, Cover

Senckenberg Deutsches Entomologisches Institut

Nova Supplementa Entomologica (NSE)



Editorial Staff

Prof. Dr. Thomas Schmitt

Scientific Interests

  • Molecular and classic biogeography
  • Classic and molecular ecology
  • •Evolutionary biology
  • Conservation biology and conservation genetics including landscape genetics
  • Systematics of Lepidoptera

Expertise Skills

  • •DNA analytics (microsatellites, mtDNA sequencing, AFLP, etc.); allozyme electrophoresis
  • Morphometry and morphometrics
  • Transect counts, light trapping, etc.
  • GPS based mark-release recapture
  • Landscape parametrisation (e.g. Ellenberg’s indicator values)
  • GIS applications

Organisms of Interest

  • Butterflies (main group)
  • •Odonates, beetles, caddisflies, tipulids, scorpions
  • Amphibians, reptiles
  • •Montane plants

Martin-Luther-Universität Halle-Wittenberg
Universitäts Professor (W3) für Entomologie, ab 2014

Universität Trier, 2001-2014
Universitäts Professor (W2) für Molekular Biogeografie, 2009-2014
Junior Professor (W1)  für Molekular Biogeografie, 2003-2009
Postdoc, 2001-2003

Johannes Gutenberg-Universität Mainz, 1996-2001
Postdoc, 2000-2001
Dissertation 1996-1999; Verteidigung 1999

Universität des Saarlandes, 1989-1996; Universitdade de Lisboa, Portugal, 1993
Diplom in Biologie, 1996

(Mai 2020)

ISI peer-reviewed Articles

Lundhagen, A., Schmitt, T., Wahlberg, N., Sarvarova, L., Konvička, M., Ryrholm, N. & Kanuch, P. (2020) Wing morphology of the butterfly Coenonympha arcania in Europe: traces of both historical isolation in glacial refugia and current adaptation. — Journal of Zoological Systematics and Evolutionary Research: in press. DOI: 10.1111/jzs.12360

Habel, J.C., Gossner, M.M. & Schmitt, T. (2020) What makes a species a priority for nature conservation? — Animal Conservation 23: 28–35.

Habel, J.C., Ulrich, U. & Schmitt, T. (2020) Butterflies in corridors: quality matters for specialists. — Insect Conservation and Diversity 13: 91–98.

Habel, J.C., Trusch, R., Schmitt, T., Ochse, M. & Ulrich, W. (2019) Long-term large-scale decline in relative abundances of butterfly and burnet moth species across south-western Germany. — Scientific Reports 9: 14921.

Habel, J.C., Samways, M. & Schmitt, T. (2019) Mitigating the precipitous decline of terrestrial European insects: Requirements for a new strategy. — Biodiversity and Conservation 28: 1343–1360.

Habel, J.C., Biburger, N., Seibold, S., Ulrich, W. & Schmitt, T. (2019) Agricultural intensification drives butterfly decline. — Insect Conservation and Diversity 12: 289–295.

Habel, J.C., Segerer, A., Ulrich, W. & Schmitt, T. (2019) Succession matters: Community shifts in moths over three decades increases multifunctionality in intermediate successional stages. — Scientific Reports 9: 5586.

Ehl, S., Holzhauer, S.I.J., Ryrholm, N. & Schmitt, T. (2019) Phenology, mobility, and behaviour of the arcto-alpine species Boloria napaea in its arctic habitat. — Scientific Reports 9: 3912.

Ehl, S., Böhm, N., Wörner, M., Rakosy, L. & Schmitt, T. (2019) Dispersal and adaptation strategies of the high mountain butterfly Boloria pales in the Romanian Carpathians. — Frontiers in Zoology 16: 1.

Bartonova, A., Konvicka, M., Korb, S., Kramp, K., Schmitt, T. & Faltynek Fric, Z. (2018) Range dynamics of Palaearctic steppe species under glacial cycles: the phylogeography of Proterebia afra (Lepidoptera: Nymphalidae: Satyrinae). — Biological Journal of the Linnean Society 125: 867–884.

Drag, L., Hauck, D., Rican, O., Schmitt, T., Shovkoon, D.F., Godunko, R.J., Curletti, G. & Cizek, L. (2018) Phylogeography of the endangered saproxylic beetle Rosalia longicorn, Rosalia alpina (Coleoptera, Cerambycidae), corresponds with its main host, the European beech (Fagus sylvatica, Fagaceae). — Journal of Biogeography 45: 2631–2644. cover paper

Habel, J.C., Seibold, S., Ulrich, W. & Schmitt, T. (2018) Seasonality overrides differences in butterfly species composition between natural and anthropogenic forest habitats. — Animal Conservation 21: 405-413.

Ehl, S., Hostert, K., Korsch, J., Gros, P. & Schmitt, T. (2018) Sexual dimorphism in the alpine butterflies Boloria pales and Boloria napaea: Differences in movement and foraging behaviour (Lepidoptera: Nymphalidae). — Insect Science 25: 1089–1101. cover paper

Kramp, K., Schmitt, T., Lang, P., Jeschke, M., Schäfer, P., Kulanek, D., Zhang, X., Yu, R. & Thomas, F.M. (2018) Clones or no clones: genetic structure of riparian Populus euphratica forests in Central Asia. — Journal of Arid Land 10: 750-766.

Rangel López, J.Á., Husemann, M., Schmitt, T., Kramp, K. & Habel, J.C. (2018) Mountain barriers and trans-Saharan connections shape genetic structures of Pimelia darkling beetles (Coleoptera, Tenebrionidae). — Biological Journal of the Linnean Society 124: 547-556.

Hochkirch, A., Beninde, J., Fischer, M., Krahner, A., Lindemann, C., Matenaar, D., Rohde, K., Wagner, N., Wesch, C., Wirtz, S., Zink, A., Lötters, S., Schmitt, T., Proelss, A. & Veith, M. (2018) License to Kill? – Disease eradication programs may not be in line with the Convention on Biological Diversity. — Conservation Letters 11: e12470.

Ehl, S., Dalstein, V., Tull, F., Gros, P. & Schmitt, T. (2018) Specialised or opportunistic – How does the high mountain endemic butterfly Erebia nivalis survive in its extreme habitats? — Insect Science 25: 161–171. cover paper

Habel, J.C. & Schmitt, T. (2018) Vanishing of the common species: Empty habitats and the role of genetic diversity. — Biological Conservation 218: 211–216.

Ehl, S., Ebertshäuser, M., Gros, P. & Schmitt, T. (2017) Population demography of alpine butterflies: Boloria pales and Boloria napaea (Lepidoptera: Nymphalidae) and their specific adaptations to high mountain environments. — Acta Oecologica 85: 53–61.

Vitali, F. & Schmitt, T. (2017) Ecological patterns strongly impact the biogeography of western Palaearctic longhorn beetles (Coleoptera: Cerambycoidea). — Organisms, Diversity & Evolution 17: 163–180.

Habel, J.C., Vila, R., Voda, R., Husemann, M., Schmitt, T. & Dapporto, L. (2017) Differentiation in the marbled white butterfly species complex driven by multiple evolutionary forces. — Journal of Biogeography 44: 433–445.

Kühne, G., Kosuch, J., Hochkirch, A. & Schmitt, T. (2017) Extra-Mediterranean glacial refugia in a Mediterranean faunal element: the phylogeography of the chalk-hill blue Polyommatus coridon (Lepidoptera, Lycaenidae). — Scientific Reports 7: 43533.

Habel, J.C., Lens, L., Eggermont, H., Githiru, M., Mulwa, R.K., Suleiman Shauri, H., Lewinsohn, T.M., Weisser, W.W. & Schmitt, T. (2017) More topics from the tropics: additional thoughts to Mammides et al. — Biodiversity and Conservation 26: 237–241.

Kramp, K., Cizek, O., Madeira, P.M., Ramos, A.A., Konvicka, M., Castilho, R. & Schmitt, T. (2016) Genetic implications of the phylogeographical patterns in the conservation of the boreal wetland butterfly Colias palaeno (Pieridae). — Biological Journal of the Linnean Society 119: 1068–1081.

Habel, J.C., Augenstein, B., Schmitt, T. & Ulrich, W. (2016) Managing towards extinction: diverging developments of plant and ground beetle assemblages following restoration of calcareous grasslands. — Basic and Applied Ecology 17: 668–677.

Schmitt, T., Louy, D., Zimmermann, E. & Habel, J.C. (2016) Species radiation in the Alps: Multiple range shifts caused diversification in Ringlet butterflies in the European high mountains. — Organisms, Diversity & Evolution 16: 791–808.

Habel, J.C., Segerer, A., Ulrich, W., Torchyk, O., Weisser, W.W. & Schmitt, T. (2016) Butterfly community shifts over two centuries. — Conservation Biology 30: 754–762.

Habel, J.C., Hillen, J., Schmitt, T. & Fischer, C. (2016) Restricted movements and high site fidelity in three East African cloud-forest birds. — Journal of Tropical Ecology 32: 83–87.

Wakil, W. & Schmitt, T. (2015): Field trials on the efficacy of Beauveria bassiana, diatomaceous earth and Imidacloprid for the protection of wheat grains from four major stored grain insect pests. — Journal of Stored Products Research 64: 160-167.

Junker, M., Zimmermann, M., Ramos, A.A., Gros, P., Konvička, M., Nève, G., Rákosy, L., Tammaru, T., Castilho, R. & Schmitt, T. (2015) Three in one – multiple faunal elements within an endangered European butterfly species. — PLoS One 10: e0142282.

Lechner, R., Kühn, R., Melzer, A., Schmitt, T. & Habel, J.C. (2015) Ecological separation versus geographic isolation: population genetics of the Water-lily leaf beetle Galerucella nymphaeae (Coleoptera: Chrysomelidae). — Biological Journal of the Linnean Society 116: 423-431.

Filz, K.J. & Schmitt, T. (2015) Niche overlap and host specificity in parasitic Maculinea butterflies (Lepidoptera: Lycaenidae) as a measure for potential extinction risks under climate change. — Organisms, Diversity and Evolution 15: 555-565.

Habel, J.C., Zachos, F.E., Dapporto, L., Rödder, D., Radespiel, U., Tellier, A. & Schmitt, T. (2015) Population genetics revisited – towards a multidisciplinary research field. Biological Journal of the Linnean Society 115: 1-12.

Besold, J. & Schmitt, T. (2015) More northern than ever thought: Refugia of the Woodland Ringlet butterfly Erebia medusa (Nymphalidae: Satyrinae) in Northern Central Europe. — Journal of Zoological Systematics and Evolutionary Research 53: 67-75.

Weyer, J., Jorgensen, D, Schmitt, T., Maxwell, T.J & Anderson, C.D. (2014) Lack of detectable genetic differentiation between den populations of the Prairie Rattlesnake (Crotalus viridis) in a fragmented landscape. — Canadian Journal of Zoology 92: 837-846.

Ashoff, M. & Schmitt, T. (2014) Are different allozyme genotypes of the butterfly Polyommatus coridon adapted to resist cold and heat shocks? — Annales Zoologici Fennici 51: 413-422.

Habel, J.C. & Schmitt, T. (2014) Towards more evidence-based nature conservation: Additional thoughts to Müller & Opgenoorth. The science-practitioners gap in nature conservation. — Basic and Applied Ecology 15: 379-384.

Schmitt, T., Habel, J.C., Rödder, D. & Louy, D. (2014) Effects of recent and past climatic shifts on the genetic structure of the high mountain Yellow-spotted ringlet butterfly Erebia manto (Lepidoptera, Satyrinae): a conservation problem. — Global Change Biology 20: 2045-2061.

Pennekamp, F., Pereira, P. & Schmitt, T. (2014) Habitat requirements and dispersal ability of the Spanish Fritillary (Euphydryas desfontainii) in southern Portugal: evidence-based conservation suggestions for an endangered taxon. — Journal of Insect Conservation 18: 497-508.

Konvicka, M., Mihali, C.V., Rakosy, L., Benes, J., Schmitt, T. (2014) Survival of cold-adapted species in isolated mountains: the population genetics of the Sudeten ringlet, Erebia sudetica sudetica, in the Jeseník Mts., Czech Republic. — Journal of Insect Conservation 18: 153-161.

Louy, D., Habel, J.C., Abadjev, S., Rakosy, L., Varga, Z., Rödder, D. & Schmitt T. (2014) Molecules and models indicate diverging evolutionary effects from parallel altitudinal range shifts in two mountain Ringlet butterflies. — Biological Journal of the Linnean Society 112: 569-583.

Heiser, M., Dapporto, L. & Schmitt, T. (2014) Coupling impoverishment analyses and partitioning of beta diversity allows a comprehensive description of Odonata biogeography in the Western Mediterranean. — Organisms, Diversity and Evolution 14: 203-214.

Louy, D., Habel, J.C., Ulrich, W. & Schmitt, T. (2014) Out of the Alps: The biogeography of a disjunctly distributed mountain butterfly, the Almond eyed ringlet Erebia alberganus (Lepidoptera, Satyrinae). — Journal of Heredity 105: 28-38.

Husemann, M., Schmitt, T., Zachos, F.E., Ulrich, W. & Habel, J.C. (2014) Palaearctic biogeography revisited: Evidence for the existence of a North African refugium for western Palaearctic biota. — Journal of Biogeography 41: 81-94.

Hochkirch, A., Schmitt, T., Beninde, J., Hiery, M., Kinitz, T., Kirschey, J., Matenaar, D., Rohde, K., Stoefen, A., Wagner, N., Zink, A., Lötters, S., Veith, M., Proelss, A. (2013) How much biodiversity does Natura 2000 cover? — Conservation Letters 6: 470-471.

Hochkirch, A., Schmitt, T., Beninde, J., Hiery, M., Kinitz, T., Kirschey, J., Matenaar, D., Rohde, K., Stoefen, A., Wagner, N., Zink, A., Lötters, S., Veith, M., Proelss, A. (2013) Europe needs a new vision for a Natura 2020 network. — Conservation Letters 6: 462-467.

Weyer, J. & Schmitt, T. (2013) Knowing the way home: strong philopatry of a highly mobile insect species, Brenthis ino. — Journal of Insect Conservation 17: 1197-1208.

Heiser, M. & Schmitt, T. (2013) Tracking the boundary between the Palaearctic and the Oriental region: New insights from dragonflies and damselflies (Odonata). — Journal of Biogeography 40: 2047-2058.

Habel, J.C., Rödder, D. Lens, L. & Schmitt, T. (2013) The genetic signature of ecologically different grassland lepidopterans. — Biodiversity and Converation 22: 2401-2411.

Filz, K.J., Engler, J.O., Stoffels, J., Weitzel, M., Schmitt, T. (2013) Missing the target? A critical view on butterfly conservation efforts on calcareous grasslands in south-western Germany. — Biodiversity and Conservation 22: 2223-2241.

Louy, D., Habel, J.C., Abadjiev, S. & Schmitt, T. (2013) Genetic legacy from past panmixia: High genetic variability and low differentiation in disjunct populations of the Eastern Large Heath butterfly. — Biological Journal of the Linnean Society 110: 281-290.

Filz, K.J., Wiemers, M., Herrig, A., Weitzel, M. & Schmitt, T. (2013) A question of adaptability – Climate and habitat change lower trait diversity in butterfly communities in south-western Germany. — European Journal of Entomology 110: 633-642.

Filz, K.J., Schmitt, T. & Engler, J.O. (2013) How fine is fine-scale? Questioning the use of fine-scale bioclimatic data in species distribution models used for forecasting abundance patterns in butterflies. — European Journal of Entomology 110: 311-317.

Dieker, P., Drees, C., Schmitt, T., Assmann, T. (2013) Low genetic diversity of a high mountain burnet moth species in the Pyrenees. — Conservation Genetics 14: 231-236.

Habel, J.C., Husemann, M., Schmitt, T., Dapporto, L., Rödder, D. & Vandewoestijne, S. (2013) A forest butterfly in Sahara desert oases: isolation does not matter. — Journal of Heredity 104: 234-247.

Pennekamp, F., Monteiro, E. & Schmitt, T. (2013) The larval ecology of the butterfly Euphydryas desfontainii (Lepidoptera: Nymphalidae) in SW-Portugal: food plant quantity and quality as main predictors of habitat quality. — Journal of Insect Conservation 17: 195-206.

Rákosy, L., Heiser, M., Manci, C.-O. & Schmitt, T. (2013) Strong divergences in regional distributions in Romania: recent ecological constraints in dragonflies (Odonata) versus ancient biogeographical patterns in butterflies (Lepidoptera: Rhopalocera). — Insect Conservation and Diversity 6: 145-154.

De Keyser, R., Shreeve, T.G., Breuker, C.J., Hails, R.S. & Schmitt, T. (2012) Polyommatus icarus butterflies in the British Isles: evidence for a bottleneck. — Biological Journal of the Linnean Society 107: 123-136.

Schmitt, T. & Varga, Z. (2012) Extra-Mediterranean refugia: The rule and not the exception? — Frontiers in Zoology 9: 22.

Molecular Ecology Resource Primer Development Consortium, Abelló, P., Weiming, A.I., Altmann, C., Bernardi, G., Bonato, O., Burchhardt, K.M., Chen, X., Chen, Z., Cizkova, D., Clouet, C., Cubeta, M.A., Garcia-Merchan, V.H., Gauthier, N., Gibson, S., Halacka, K., Hamdi, F., Hankeln, T., Hochkirch, A., Hrbek, T., Jackson, A.M., Lin, C., Lin, S.-M., Macpherson, E., Macrander, J., Maresova, E., Mendel, J., Nowak, M., Orti, G., Palero, F., Papousek, I., Pascual, M., Schmitt, T., Semmens, B.X., Streito, J.-C., Tian, E.-W., Tseng, S.-P., Veith, M., Svetesnik, L., Wang, H.-Y., Weyer, J., Willis, S., Yu, H. & Zhou, Z. (2012) Permanent Genetic Resources added to Molecular Ecology Resources Database 1 April 2012 – 31 May 2012. — Molecular Ecology Resources 12: 972-974.

Habel, J.C., Engler, J.O., Rödder, D. & Schmitt, T. (2012) Contrasting genetic and morphologic responses on recent population decline in two burnet moths (Lepidoptera, Zygaenidae). — Conservation Genetics 13: 1293-1304.

Habel, J.C., Engler, J.O., Rödder, D. & Schmitt, T. (2012) Landscape genetics of a recent population extirpation in a burnet moth species. — Conservation Genetics 13: 247-255.

Alvarez, N., Manel, S., Schmitt, T. & the IntraBioDiv Consortium (2012) Contrasting diffusion of Quaternary gene pools across Europe: The case of the arctic–alpine Gentiana nivalis L. (Gentianaceae). — Flora 207: 408-413.

Huck, S., Büdel, B., Schmitt, T. (2012) Ice-age isolation, postglacial hybridization and recent population bottlenecks shape the genetic structure of Meum athamanticum in Central Europe. — Flora 207: 399-407.

Habel, J.C. & Schmitt, T. (2012) The burden of genetic diversity. — Biological Conservation 147: 270-274.

Schmitt, T. & Zimmermann, M. (2012) To hybridize or not to hybridize: what separates two genetic lineages of the Chalk-hill Blue Polyommatus coridon (Lycaenidae, Lepidoptera) along their secondary contact zone throughout eastern Central Europe? — Journal of Zoological Systematics and Evolutionary Research 50: 106-115.

Husemann, M., Schmitt, T., Stathi, I. & Habel, J.C. (2012) Evolution and radiation in the scorpion Buthus elmoutaouakili Lourenço & Qi, 2006 (Scorpiones: Buthidae) at the foothills of the Atlas Mountains (North Africa). — Journal of Heredity 103: 221-229.

Habel, J.C., Husemann, M., Schmitt, T., Zachos, F.E., Honnen, A.-C., Petersen, B., Parmakelis, A. & Stathi, I. (2012) Microallopatry caused strong diversification in Buthus scorpions (Scorpiones: Buthidae) in the Atlas Mountains (NW Africa). — PLoS ONE 7: e29403.

Habel, J.C., Husemann, M., Rödder, D. & Schmitt, T. (2011) Biogeographical dynamics of the Spanish Marbled White Melanargia ines (Lepidoptera: Satyridae) in the western Mediterranean: does the Atlanto-Mediterranean refuge exist? — Biological Journal of the Linnean Society 104: 828-837.

Bernard, R., Heiser, M., Hochkirch, A. & Schmitt, T. (2011) Genetic homogeneity of the Sedgling Nehalennia speciosa (Odonata: Coenagrionidae) indicates a single Würm glacial refugium and trans-Palaearctic postglacial expansion. — Journal of Zoological Systematics and Evolutionary Research 49: 292-297.

Wagner, N., Pellet, J., Lötters, S., Schmidt, B.R. & Schmitt, T. (2011) The superpopulation approach for estimating the population size of ‘prolonged’ breeding amphibians: Examples from Europe. — Amphibia-Reptilia 32: 323-332.

Habel, J.C., Lens, L., Rödder, D. & Schmitt, T. (2011) From Africa to Europe and back: refugia and range shifts cause high genetic differentiation in the Marbled White butterfly Melanargia galathea. — BMC Evolutionary Biology 11: 215.

Dapporto, L., Habel, J.C., Dennis, R.L.H. & Schmitt, T. (2011) The biogeography of the western Mediterranean: elucidating contradictory distribution patterns of differentiation in Maniola jurtina (Lepidoptera: Nymphalidae). — Biological Journal of the Linnean Society 103: 571-577.

Dapporto, L., Schmitt, T., Vila, R., Scalercio, S., Biermann, H., Dinca, V., Gayubo, S.F., González, J.A., Lo Cascio, P., Dennis, R.L.H. (2011) Phylogenetic island disequilibrium: evidence for ongoing long-term population dynamics in two Mediterranean butterflies. — Journal of Biogeography 38: 854-867.

Rákosy, L. & Schmitt, T. (2011) Are butterflies and moths suitable ecological indicator systems for restoration measures of semi-natural calcareous grassland habitats? — Ecological Indicators 11: 1040-1045.

Vila, M., Marí-Mena, N., Guerrero, A. & Schmitt, T. (2011) Some butterflies do not care much about topography: a single genetic lineage of Erebia euryale (Nymphalidae) along the northern Iberian mountains. — Journal of Zoological Systematics and Evolutionary Research 49: 119-132.

Sinama, M., Dubut, V., Costedoat, C., Gilles, A., Junker, M., Malausa, T., Martin, J.-F., Nève, G., Pech, N., Schmitt, T., Zimmermann, M. & Meglécz, E. (2011) Challenge of microsatellite development in Lepidoptera: Euphydryas aurinia (Nymphalidae) as a case study. — European Journal of Entomology 108: 261-266.

Habel, J.C., Finger, A., Schmitt, T. & Nève, G. (2011) Survival of the endangered butterfly Lycaena helle in a fragmented environment: Genetic analyses over 15 years. — Journal of Zoological Systematics and Evolutionary Research 49: 25-31.

Habel, J.C., Rödder, D., Schmitt, T. & Nève, G. (2011) Global warming will affect genetic diversity and uniqueness of Lycaena helle populations. — Global Change Biology 17: 194-205.

Junker, M., Wagner, S., Gros, P. & Schmitt, T. (2010) Changing demography and dispersal behaviour: ecological adaptation in an alpine butterfly. — Oecologia 164: 971-980.

Habel, J.C., Ivinskis, P. & Schmitt, T. (2010) On the limit of altitudinal range shifts – population genetics of relict butterfly populations. — Acta Zoologica Academiae Scientiarum Hungaricae 56: 383-393.

Michl, T., Huck, S., Schmitt, T., Liebrich, A., Haase, P. & Büdel, B. (2010) The molecular population structure of the tall-forb Cicerbita alpina (Asteraceae) supports the idea of cryptic glacial refugia in central Europe. — Botanical Journal of the Linnean Society 164: 142-154.

Habel, J.C., Junker, M. & Schmitt, T. (2010) High dispersal ability and low genetic differentiation in the widespread butterfly species Melanargia galathea. — Journal of Insect Conservation 14: 467-478.

Habel, J.C., Schmitt, T., Meyer, M., Finger, A., Rödder, D., Assmann, T. & Zachos, F. (2010) Biogeography meets conservation: the genetic structure of the endangered lycaenid butterfly Lycaena helle (Denis & Schiffermüller, 1775). — Biological Journal of the Linnean Society 101: 155-168.

Ujvárosi, L., Bálint, M., Schmitt, T., Mészáros, N., Ujvárosi, T. & Popescu, O. (2010) Divergence and speciation in the Carpathians area: patterns of morphological and genetic diversity of the crane fly Pedicia occulta (Diptera:Pediciidae). — Journal of the North American Benthological Society 29: 1075-1088.

Kühne, G. & Schmitt, T. (2010) Genotype shifts along one generation of the blue butterfly Polyommatus coridon without changes in allele frequencies. — Annales Zoologici Fennici 47: 278-286.

Junker, M. & Schmitt, T. (2010) Demography, dispersal and movement pattern of Euphydryas aurinia (Lepidoptera: Nymphalidae) at the Iberian Peninsula: an alarming example in an increasingly fragmented landscape? — Journal of Insect Conservation 14: 237-246.

Bernard, R. & Schmitt, T. (2010) Genetic poverty of an extremely specialized wetland species, Nehalennia speciosa: implications for conservation (Odonata: Coenagrionidae). — Bulletin of Entomological Research 100: 405-413.

Schmitt, T. & Besold, J. (2010) Upslope movements and large scale expansions: the taxonomy and biogeography of the Coenonympha arcania – C. darwiniana – C. gardetta butterfly species complex. — Zoological Journal of the Linnean Society 159: 890-904.

Kadlec, T., Vrba, P., Kepka, P., Schmitt, T. & Konvicka, M. (2010) Tracking the decline of the once-common butterfly: delayed oviposition, demography and population genetics in the hermit Chazara briseis. — Animal Conservation 13: 172-183.

Habel, J.C., Rödder, D., Stefano, S., Meyer, M. & Schmitt, T. (2010) Strong genetic cohesiveness between Italy and North Africa in four butterfly species. — Biological Journal of the Linnean Society 99: 818-830.

Hammouti, N., Schmitt, T., Seitz, A., Kosuch, J. & Veith, M. (2010) Combining mitochondrial and nuclear evidences: a refined evolutionary history of Erebia medusa (Lepidoptera: Nymphalidae: Satyrinae) in Central Europe based on the CO1 gene. — Journal of Zoological Systematics and Evolutionary Research 48: 115-125.

Heiser, M. & Schmitt, T. (2010) Do different dispersal capacities influence the biogeography of the western Palearctic dragonflies (Odonata)? — Biological Journal of the Linnean Society 99: 177-195.

Habel, J.C., Zachos, F.E., Finger, A., Meyer, M., Louy, D., Assmann, T. & Schmitt, T. (2009) Unprecedented long-term genetic monomorphism in an endangered relict butterfly species. — Conservation Genetics 10: 1659-1665.

Habel, J.C., Dieker, P. & Schmitt, T. (2009) Biogeographical connections between the Maghreb and the Mediterranean peninsulas of southern Europe. — Biological Journal of the Linnean Society 98: 693-703.

Habel, J.C. & Schmitt, T. (2009) The genetic consequences of different dispersal behaviours in Lycaenid butterfly species. — Bulletin of Entomological Research 99: 513-523.

Karl, I., Schmitt, T. & Fischer, K. (2009) Genetic differentiation between alpine and lowland populations of a butterfly is related to PGI enzyme genotype. — Ecography 32: 488-496.

Finger, A., Schmitt, T., Zachos, F.E., Meyer, M., Assmann, T. & Habel, J.C. (2009) The genetic status of the violet copper Lycaena helle – a relict of the cold past in times of global warming. — Ecography 32: 382-390.

Habel, J.C., Meyer, M. & Schmitt, T. (2009) The genetic consequence of differing ecological demands of a generalist and a specialist butterfly species. — Biodiversity and Conservation 18: 1895-1908.

Schmitt, T. (2009) Biogeographical and evolutionary importance of the European high mountain systems. — Frontiers in Zoology 6: 9.

Kramp, K., Huck, S., Niketić, M., Tomović, G. & Schmitt, T. (2009) Multiple glacial refugia and complex postglacial range shifts of the obligatory woodland plant Polygonatum verticillatum (Convallariaceae). — Plant Biology 11: 392-404.

Lermen, D., Blömeke, B., Browne, R., Clarke, A., Dyce, P., Fixemer, T., Fuhr, G., Holt, W.V., Jewgenow, K., Lloyd, R.E., Lötters, S., Paulus, M., McGregor Reid, G., Rapoport, D.H., Rawson, D., Ringleb, J., Ryder, O.A., Spörl, G., Schmitt, T., Veith, M. & Müller, P. (2009) Cryobanking of viable biomaterials: Necessities of new strategies for conservation purposes. — Molecular Ecology 18: 1030-1033.

Bálint, M., Barnard, P.C., Schmitt, T., Ujvárosi, L. & Popescu, O. (2008) Differentiation and speciation in mountain streams: a case study in the caddisfly Rhyacophila aquitanica (Trichoptera). — Journal of Zoological Systematics and Evolutionary Research 46: 340-345.

Besold, J., Schmitt, T., Tammaru, T. & Cassel-Lundhagen, A. (2008) Strong genetic impoverishment from the centre of distribution in southern Europe to peripheral Baltic and isolated Scandinavian populations of the pearly heath butterfly. — Journal of Biogeography 35: 2090-2101.

Besold, J., Huck, S. & Schmitt, T. (2008) Allozyme polymorphisms in the small heath Coenonympha pamphilus: recent ecological selection or old biogeographical signal? — Annales Zoologici Fennici 45: 217–228.

Habel, J.C., Meyer, M., El Mousadik, A. & Schmitt, T. (2008) Africa goes Europa: The complete phylogeography of the marbled white butterfly species complex Melanargia galathea / lachesis (Lepidoptera: Satyridae). — Organisms, Diversity & Evolution 8: 121-129.

Karl, I., Schmitt, T. & Fischer, K. (2008) Phosphoglucose isomerase genotype affects life-history traits and cold stress resistance in a Copper butterfly. — Functional Ecology 22: 887-894.

Schmitt, T. & Haubrich, K. (2008) The genetic structure of the mountain forest butterfly Erebia euryale unravels the late Pleistocene and postglacial history of the mountain coniferous forest biome in Europe. — Molecular Ecology 17: 2194–2207.

Schmitt, T., Augenstein, B. & Finger, A. (2008) The influence of changes in viticulture management on the butterfly (Lepidoptera) diversity in a wine growing region of southwestern Germany. — European Journal of Entomology 105: 249-255.

Habel, J.C., Finger, A., Meyer, M., Schmitt, T. & Assmann, T. (2008) Polymorphic microsatellite loci in the endangered butterfly Lycaena helle (Lepidoptera: Lycaenidae). — European Journal of Entomology 105: 361–362.

Varga, Z.S. & Schmitt, T. (2008) Types of oreal and oreotundral disjunction in the western Palearctic. — Biological Journal of the Linnean Society 93: 415-430.

Schmitt, T. (2007) Molecular Biogeography of Europe: Pleistocene cycles and postglacial trends. — Frontiers in Zoology 4: 11. highly accessed

Haubrich, K. & Schmitt, T. (2007) Cryptic differentiation in alpine-endemic, high-altitude butterflies reveals down-slope glacial refugia. — Molecular Ecology 16: 3643-3658.

Schmitt, T. & Müller, P. (2007) Limited hybridization along a large contact zone between two genetic lineages of the butterfly Erebia medusa (Satyrinae, Lepidoptera) in Central Europe. — Journal of Zoological Systematics and Evolutionary Research 45: 39-46.

Schmitt, T., Rákosy, L., Abadjiev, S. & Müller, P. (2007) Multiple differentiation centres of a non-Mediterranean butterfly species in south-eastern Europe. — Journal of Biogeography 34: 939-950.

Louy, D., Habel, J.C., Schmitt, T., Assmann, T., Meyer, M. & Müller, P. (2007) Strongly diverging population genetic patterns of three skipper species: the role of habitat fragmentation and dispersal ability. — Conservation Genetics 8: 671-681.

Habel, J.C., Schmitt, T., Härdtle, W., Lütkepohl, M. & Assmann, T. (2007) Dynamics in a butterfly–plant–ant system: influence of habitat characteristics on turnover rates of the endangered lycaenid Maculinea alcon. — Ecological Entomology 32: 536-543.

Schmitt, T. & Rákosy, L. (2007) Changes of traditional agrarian landscapes and their conservation implications: a case study of butterflies in Romania. — Diversity and Distributions 13: 855-862.

Schmitt, T., Hewitt, G.M. & Müller, P. (2006) Disjunct distributions during glacial and interglacial periods in mountain butterflies: Erebia epiphron as an example. — Journal of Evolutionary Biology 19: 108-113.

Wenzel, M., Schmitt, T., Weitzel, M. & Seitz, A. (2006) The severe decline of butterflies on western German calcareous grasslands during the last 30 years: A conservation problem. — Biological Conservation 128: 542-552.

Schmitt, T., Habel, J.C., Zimmermann, M. & Müller, P. (2006) Genetic differentiation of the marbled white butterfly, Melanargia galathea, accounts for glacial distribution patterns and postglacial range expansion in southeastern Europe. — Molecular Ecology 15: 1889-1901.

Schmitt, T., Habel, J.C., Besold, J., Becker, T., Johnen, L., Knolle, M., Rzepecki, A., Schultze, J. & Zapp, A. (2006) The Chalk-hill Blue Polyommatus coridon (Lycaenidae, Lepidoptera) in a highly fragmented landscape: How sedentary is a sedentary butterfly? — Journal of Insect Conservation 10: 311-316.

Schmitt, T., Cizek, O. & Konvicka, M. (2005) Genetics of a butterfly relocation: large, small and introduced populations of the mountain endemic Erebia epiphron silesiana. — Biological Conservation 123: 11-18.

Schmitt, T., Röber, S. & Seitz, A. (2005) Is the last glaciation the only relevant event for the present genetic population structure of the meadow brown butterfly Maniola jurtina (Lepidoptera: Nymphalidae)? — Biological Journal of the Linnean Society 85: 419-431.

Habel, J.C., Schmitt, T. & Müller, P. (2005) The fourth paradigm pattern of post-glacial range expansion of European terrestrial species: the phylogeography of the Marbled White butterfly (Satyrinae, Lepidoptera). — Journal of Biogeography 32: 1489-1497.

Schmitt, T., Varga, Z. & Seitz, A. (2005) Are Polyommatus hispana and Polyommatus slovacus bivoltine Polyommatus coridon (Lepidoptera: Lycaenidae)? The discriminatory value of genetics in taxonomy. — Organisms, Diversity & Evolution 5: 297-307.

Schmitt, T. & Hewitt, G.M. (2004) The genetic pattern of population threat and loss: a case study of butterflies. — Molecular Ecology 13: 21-31.

Schmitt, T. & Seitz, A. (2004) Low diversity but high differentiation: the population genetics of Aglaope infausta (Zygaenidae: Lepidoptera). — Journal of Biogeography 31: 137-144.

Krauss, J., Schmitt, T., Seitz, A., Steffan-Dewenter, I. & Tscharntke, T. (2004) Effects of habitat fragmentation on the genetic structure of the monophagous butterfly Polyommatus coridon along its northern range margin. — Molecular Ecology 13: 311-320.

Schmitt, T. & Hewitt, G.M. (2004) Molecular biogeography of the arctic-alpine disjunct burnet moth species Zygaena exulans (Zygaenidae, Lepidoptera) in the Pyrenees and Alps. — Journal of Biogeography 31: 885-893.

Schmitt, T. & Krauss, J. (2004) Reconstruction of the colonization route from glacial refugium to the northern distribution range of the European butterfly Polyommatus coridon (Lepidoptera: Lycaenidae). — Diversity and Distributions 10: 271-274.

Schmitt, T., Gießl, A. & Seitz, A. (2003) Did Polyommatus icarus (Lepidoptera: Lycaenidae) have distinct glacial refugia in southern Europe? Evidence from population genetics. — Biological Journal of the Linnean Society 80: 529-538.

Schmitt, T. (2003) Influence of forest and grassland management on the diversity and conservation of butterflies and burnet moths (Lepidoptera: Papilionoidea, Hesperiidae, Zygaenidae). — Animal Biodiversity and Conservation 26.2: 51-67.

Schmitt, T., Gießl, A. & Seitz, A. (2002) Postglacial colonisation of western Central Europe by Polyommatus coridon (Poda 1761) (Lepidoptera: Lycaenidae): evidence from population genetics. — Heredity 88: 26-34.

Schmitt, T. & Seitz, A. (2002) Influence of habitat fragmentation on the genetic structure of Polyommatus coridon (Lepidoptera: Lycaenidae): implications for conservation. — Biological Conservation 107: 291-297.

Schmitt, T. & Seitz, A. (2002) Postglacial distribution area expansion of Polyommatus coridon (Lepidoptera: Lycaenidae) from its Ponto-Mediterranean glacial refugium. — Heredity 89: 20-26.

Schmitt, T. & Seitz, A. (2001) Intraspecific allozymatic differentiation reveals the glacial refugia and the postglacial expansions of European Erebia medusa (Lepidoptera: Nymphalidae). — Biological Journal of the Linnean Society 74: 429-458.

Schmitt, T. & Seitz, A. (2001) Allozyme variation in Polyommatus coridon (Lepidoptera: Lycaenidae): identification of ice-age refugia and reconstruction of post-glacial expansion. — Journal of Biogeography 28: 1129-1136.

Schmitt, T., Varga, Z. & Seitz, A. (2000) Forests as dispersal barriers for Erebia medusa (Nymphalidae, Lepidoptera). — Basic and Applied Ecology 1: 53-59.


other peer-reviewed Articles

Schmitt, T. & Ehl, S. (2020) Some like it cold: Wie Permutterfalter der Gattung Boloria in ihren extremen Gebirgslebensräumen überleben. — Mitteilungen der Deutschen Gesellschaft für allgemeine und angewandte Entomologie 22.

Schmitt, T., Bartonova, A., Konvicka, M., Korb, S., Kramp, K. & Fric, Z.F. (2020) Was ein Schmetterling uns über die Dynamik der Steppen des westlichen Eurasiens erzählt: Die Phylogeographie von Proterebia afra (Satyrinae) — Mitteilungen der Deutschen Gesellschaft für allgemeine und angewandte Entomologie 22.

Schmitt, T. (2019) Insektenvielfalt und ihre Funktion in Ökosystemen. — Natur und Landschaft 94: 222-229.

Krahner, A., Dathe, H.H. & Schmitt, T. (2018) Wildbienen (Hymenoptera, Aculeata: Apiformes) des Mittleren Moseltals: Die Weinbausteillagen im Klotten-Treiser Moseltal. — Contributions to Entomology 68: 107-131.

Habel, J.C., Teucher, M., Schmitt, T. & Ulrich, W. (2018) Documenting the chronology of declining ecosystem health along East African rivers. — Remote Sensing in Ecology and Conservation 4: 34–43.

Schmitt, T. (2015) Biogeographie – einmal um die Welt. — Mitteilungen der Deutschen Gesellschaft für allgemeine und angewandte Entomologie 20: 87-100.

Schmitt, T. (2015) Biology and biogeography of the chalk-hill blue Polyommatus coridon – insect of the year 2015 for Germany, Austria and Switzerland. — Nota lepidopterologica 38: 107-126.

Habel, J.C. & Schmitt, T. (2014) Flügelgeschichten – Was Schmetterlingsflügel über die Evolution und den Zustand von Populationen erzählen. — Entomologie heute 26: 123-137.

Heiser, M. & Schmitt, T. (2014) Bekannte Zentren und neue Grenzen: Die Biogeographie der Libellen Eurasiens. — Entomologie heute 26: 87-109.

Schmitt, T. (2014) Zweck von Naturschutzgebieten. — Entomologie heute 26: 169-170.

Beierkuhnlein, C., Jentsch, A., Hahn, I. & Schmitt, T. (2011) Natürliche Laboratorien der Biogeographie – Inseln: Ursprung der Vielfalt. — Biologie in unserer Zeit 41: 384-394.

Habel, J.C., Meyer, M. & Schmitt, T. (2011) Der biologische Wert peripherer Populationen am Beispiel der Tagfalterarten Lycaena helle und Parnassius apollo. — Entomologie heute 23: 93-102.

Schmitt, T. (2011) Schmetterlinge – wer kommt, wer geht? Einwanderungsrouten nach Mitteleuropa. — Biologie in unserer Zeit 41: 324-332.

Schmitt, T. (2009) Mediterran, kontinental und arkto-alpin: Die drei biogeographischen Grundmuster Europas und des Mittelmeerraumes am Beispiel von Schmetterlingen. — Entomologie heute 21: 3-19.

Schmitt, T. (2009) Gene als Berichterstatter der Evolution: Die molekulare Biogeografie der europäischen Schmetterlinge. — Praxis der Naturwissenschaften – Biologie in der Schule 58/3: 30-34.

Veith, M. & Schmitt, T. (2008) Möglichkeiten und Grenzen der Naturschutzgenetik. — Zeitschrift für Feldherpetologie 15: 119-138.

Schmitt, T. (2007) Gene berichten über die Vergangenheit: Die molekulare Biogeographie europäischer Tagfalterarten. — Entomologica Romanica 12: 31-39.

Schmitt, T. (2006) Die Bedeutung der molekularen Biogeographie im Naturschutz. — Entomologie heute 18: 133-141.

Schmitt, T. (2005) Molekulare Biogeographie des Gemeinen Bläulings Polyommatus icarus (Lepidoptera, Lycaenidae). — Entomologie heute 17: 195-204.

Schmitt, T., Lambracht, E., Scheelke, K., Tietze, D.T. & Seitz, A. (2002) Wie viele Tagfalterarten gibt es in einem Habitat? Eine ökologische Fallstudie in Norditalien (Lepidoptera: Rhopalocera). — Nachrichten des Entomologischen Vereins Apollo 23: 1-6.

Schmitt, T. & Seitz, A. (2001) Intraspecific structuring of Polyommatus coridon (Lycaenidae). — Nota lepidopterologica 24: 53-63.

Schmitt, T. (2000) Anmerkungen zur Biologie von Saturnia pavonia (Linnaeus, 1761) im südwestlichen Hunsrück (Lepidoptera: Saturniidae). — Nachrichten des entomolo­gischen Vereins Apollo, N.F. 21: 173-176.



Schmitt, T. (2020) Molekulare Biogeographie: Gene in Raum und Zeit. UTB. — Haupt Verlag, Bern, 504 pp.

Habel, J.C., Meyer, M. & Schmitt, T. (eds) (2014) Jewels in the mist – A synopsis on the highly endangered butterfly species Violet Copper, Lycaena helle. Pensoft, Sofia, 235 pp.

Schmitt, T. (1999) Phylogeographie europäischer Tagfalter basierend auf populations­genetischen Analysen unter besonderer Berücksichtigung von Polyommatus coridon und Polyommatus icarus (Lepidoptera: Lycaenidae) sowie Erebia medusa und Maniola jurtina (Lepidoptera: Satyrinae). — Dissertation, Universität Mainz, 200 pp.


Congress volumes

Schmitt, T., Blank, S.M., Köhler, A., Kramp, K. & Weyer, J. (eds.) (2015) Entomology congress 02.–05.03.2015 Frankfurt/Main. Program and Abstracts. Frankfurt am Main, 120 pp.


Book Chapters (mostly peer-reviewed*)

Türkay, M., Fritz, U., Schmitt, T., Xylander, W., Ernst, R., Kallweit, U., Klass, K.-D., Nuss, M., Päckert, M., Schmidt, C., Reimann, A., Schniebs, K., Stefen, C., Weck-Heimann, A. & Zarske, A. (2018) Frankfurt, Dresden, Görlitz, Müncheberg: Senckenberg: Its zoological collections and their history. — In Beck, L.A. (ed.) Zoological collections of Germany, Natural history collections. Springer, Heidelberg: 317–371.

Schmitt, T. & Habel, J.C. (2017) Ausgeflattert: Warum haben wir ein Schmetterlingssterben? — In: Vössing, A. (ed.) Nationalpark-Jahrbuch Unteres Odertal 2017. Nationalparkstiftung Unteres Odertal, Schloss Criewen: 34–45.

*Schmitt, T. (2017) Molecular biogeography of the high mountain systems of Europe: an overview. — In: Catalan, J., Ninot, J.M. & Aniz, M.M. (eds) High Mountain Conservation in a Changing World. Springer, Heidelberg: 63–74.

Schmitt, T. (2017) Die Insektenfauna. — In: Blahy, B. & Flade, M. (eds.) Grumsin – Weltnaturerbe im Biosphärenreservat Schorfheide-Chorin. Natur+Text, Rangsdorf: 109–110.

*Schmitt, T. (2016) Molecular biogeography of the high mountain systems of Europe: an overview. — In: Catalan, J., Ninot, J. & Aniz, M. (eds) High Mountain Conservation in a Changing World. Springer, Heidelberg: in press.

Habel, J.C. & Schmitt, T. (2014) Why protecting relict populations? — In: Habel, J.C., Meyer, M. & Schmitt, T. (eds) Jewels in the mist – A synopsis on the highly endangered butterfly species Violet Copper, Lycaena helle. Pensoft, Sofia: 231-235.

*Habel, J.C., Mattern, S., Meyer, M., Schmitt, T. (2014) Morphologic variability and signals of environmental stress: Wing-shape analyses in the Violet Copper Lycaena helle. — In: Habel, J.C., Meyer, M. & Schmitt, T. (eds) A synopsis on the highly endangered butterfly species Violet Copper, Lycaena helle. Pensoft, Sofia: 141-159.

*Habel, J.C., Finger, A., Meyer, M. & Schmitt, T. (2014) Isolated and unique: The conservation value of extant populations of Lycaena helle at the species’ western distribution edge. — In: Habel, J.C., Meyer, M. & Schmitt, T. (eds) Jewels in the mist – A synopsis on the highly endangered butterfly species Violet Copper, Lycaena helle. Pensoft, Sofia: 127-136.

*Habel, J.C., Meyer, M., Schmitt, T., Husemann, M., Varga, Z. (2014) The molecular biogeography of the Violet Copper Lycaena helle. — In: Habel, J.C., Meyer, M. & Schmitt, T. (eds) Jewels in the mist – A synopsis on the highly endangered butterfly species Violet Copper, Lycaena helle. Pensoft, Sofia: 111-124.

*Schmitt, T. & Habel, J.C. (2014) Biogeography: From the analysis of distributions to the distribution of genetic variants. — In: Habel, J.C., Meyer, M. & Schmitt, T. (eds) A synopsis on the highly endangered butterfly species Violet Copper, Lycaena helle. Pensoft, Sofia: 107-109.

Habel, J.C., Meyer, M. & Schmitt, T. (2014) Editorial. — In: Habel, J.C., Meyer, M. & Schmitt, T. (eds) A synopsis on the highly endangered butterfly species Violet Copper, Lycaena helle. Pensoft, Sofia: 1-2.

Schmitt, T. (2011) Molecular biogeography of terrestrial insects in Europe: case studies in butterflies. — In: Ujvárosi, L. & Marko, B. (eds.) The Carpathians as speciation centres and barriers. From case studies to general patterns. Cluj University Press, Cluj-Napoca: 51-82.

*Habel, J.C., Schmitt, T. & Assmann, T. (2010) Relict species: Some concluding remarks. — In: Habel, J.C. & Assmann, T. (eds.) Relict Species: Phylogeography and Conservation Biology. Springer, Heidelberg: 441-442.

*Habel, J.C., Assmann, T., Schmitt, T. & Avise, J.C. (2010) Relict species research: Some concluding remarks. — In: Habel, J.C. & Assmann, T. (eds.) Relict Species: Phylogeography and Conservation Biology. Springer, Heidelberg: 1-5.

*Habel, J.C., Drees, C., Schmitt, T. & Assmann, T. (2010) Refugial areas and postglacial colonizations in the western Palearctic. — In: Habel, J.C. & Assmann, T. (eds.) Relict Species: Phylogeography and Conservation Biology. Springer, Heidelberg: 189-197.

*Konvicka, M., Benes, J. & Schmitt, T. (2010) Ecological limits vis-à-vis changing climate: relic Erebia butterflies in insular Sudeten mountains. — In: Habel, J.C. & Assmann, T. (eds.) Relict Species: Phylogeography and Conservation Biology. Springer, Heidelberg: 341-355.

*Schmitt, T., Muster, C. & Schönswetter, P. (2010) Are disjunct alpine and arctic-alpine animal and plant species in the western Palearctic really “relics of a cold past”? — In: Habel, J.C. & Assmann, T. (eds.) Relict Species: Phylogeography and Conservation Biology. Springer, Heidelberg: 239-252.

Veith, M. & Schmitt, T. (2009) Naturschutzgenetik bei Amphibien: von der Theorie zur Praxis. — In: Hachtel, M., Schüpmann, M., Thiesmeier, K. & Weddeling, K. (eds.) Methoden der Feldherpetologie. — Zeitschrift für Feldherpetologie, Supplement 15: 153-184.

*Dennis, R.L.H. & Schmitt, T. (2009) Faunal structures, phylogeography and historical inference. — In: Settele, J., Shreeve, T.G., Konvicka, M. & van Dyck, H. (eds.) Ecology of Butterflies in Europe. Cambridge University Press, Cambridge: 250-280.

Schmitt, T. & Varga, Z. (2009) Biogeography of the butterflies and larger moths of the Carpathian Basin and the Balkan Peninsula. — In: Stloukal, E., Hensel, K., Holec, P., Illyová, M., Jandzík, D., Jedlička, L., Joniak, P., Juráni, B., Kocian, Ľ., Košel, V., Krno, I., Kúdela, M., Miklós, P., Mikulíček, P., Obuch, J., Schmitt, T., Stankoviansky, M., Stloukalová, V., Varga, Z.S., Žiak, D. (eds.) Vývoj prírody Slovenska. Faunima, Bratislava: 143-166.

Schmitt, T. (2003) Análises Genéticas: um método importante para a compreensão e protecção dos Lepidópteros. — In: Maravalhas, E. (ed.) As Borboletas de Portugal. — Apollo Books, Stenstrup: 111-120.

Maravalhas, E., Garcia-Pereira, P. & Schmitt, T. (2003) Guia Monográfico Continente. — In: Maravalhas, E. (ed.) As Borboletas de Portugal. — Apollo Books, Stenstrup: 167-324.

Proceedings (partly peer-reviewed*)

Schmitt, T. (2006) Landschaftsstrukturen und molekulare Biogeographie. — Pp 465-472 in: Kulke, E., Monheim, H., Wittmann, P. (eds.) Grenzwerte. — Tagungsbericht und wissenschaftliche Abhandlungen 55. Deutscher Geographentag Trier 2005. Berlin, Leipzig, Trier.

Schmitt, T. (2003) Wie ökologische Nischen regionale Verbreitungsmuster beeinflussen: ein Fallbeispiel an Rhopaloceren und Zygaeniden (Lepidoptera) aus Südportugal. — Verhandlungen des Westdeutschen Entomologentag 2001: 167-176.

Schmitt, T. (2003) Biogeography and ecology of southern Portuguese butterflies and burnets (Lepidoptera). — In: Reemer, M., van Helsdingen, P.J. & Kleukers, R.M.J.C. (eds.) Changes in ranges: invertebrates on the move. — Proceedings of the EIS-Symposium, Leiden, 2-5 September 2001: 69-78.

Schmitt, T. (2003) Eiablageverhalten und Raupenfutterpflanzen von Tagfaltern und Widderchen in Rheinland-Pfalz und im Saarland (Lepidoptera). — Verhandlungen des Westdeutschen Entomologentag 2002: 93-104.

Schmitt, T. (2002) Stirbt der Rundaugen-Mohrenfalter Erebia medusa (Lepidoptera: Satyridae) regional aus? Ökologische und populationsgenetische Hinweise. — In: Gerken, B. & M. Görner (eds.) Planung contra Evolution? Über Evolution und Landschaftsentwicklung in Mitteleuropa. — Natur- und Kulturlandschaft 5: 177-184.

Schmitt, T. (2001) Arealgeschichte von Erebia medusa (Lepidoptera, Nymphalidae) im Würm und Postglazial. — Verhandlungen des Westdeutschen Entomologentag 2000: 253-265.

Schmitt, T. & Seitz, A. (2001) Influence of the ice-age on the genetics and intraspecific differentiation of butterflies. — Proceedings of the International Colloquium of the European Invertebrate Survey (EIS), Macevol Priory, Arboussols (66-France), 30 August 1999 – 4 September 1999: 16-26.

Schmitt, T. (2000) Glaziale Refugien und postglaziale Arealausweitung von Polyommatus coridon (Lepidoptera: Lycaenidae). — Verhandlungen des Westdeutschen Entomo­logentag 1999: 65-79.

*Schmitt, T. & Seitz, A. (1999) Analyse der genetischen Struktur und des Arealsystems mitteleuropäischer Populationen von Erebia medusa (Lepidoptera: Nymphalidae). — Ver­handlungen der Gesellschaft für Ökologie 29: 381-387. 

not peer-reviewed articles

Gelbrecht, J., Rämisch, F., Reinhardt, R., Richert, A. & Schmitt, T. (2015) Polyommatus coridon (Poda, 1761) – Insekt des Jahres 2015 (Deutschland, Österreich, Schweiz): Einwanderungsgeschichte, aktuelle Verbreitung und Gefährdung in Brandenburg und Berlin (Lepidoptera, Lycaenidae). — Märkische Entomologische Nachrichten: in press.

Schmitt, T. (2015) Wer kommt, wer geht: Einwanderungsrouten in die Großregion Saar-Lor-Lux und Rheinland-Pfalz am Beispiel von Schmetterlingen. — Abhandlungen der Delattinia 40: 49-56.

Schmitt, T. (2015) Der Silbergrüne Bläuling Polyommatus coridon – Anmerkungen zur Biologie und Biogeographie des Insekts des Jahres 2015 (Lepidoptera). — Entomologische Nachrichten und Berichte 59: 1-8.

Herrig, A., Schmitt, T., Weitzel, M. & Filz, K.J. (2014) Zusammensetzung und Bestandsentwicklung von Tagfalterzönosen (Lepidoptera: Rhopalocera) auf Brachflächen im Raum Trier über 40 Jahre. — Mainzer naturwissenschaftliches Archiv 51: 357-392.

Filz, K.J., Jänicke, D., Schmitt, T., Frede, M. & Weitzel, M. (2013) Untersuchungen zur Bestandsentwicklung der Tagfalterzönosen auf Kalkmagerrasen im Saar-Mosel-Gebiet. — Mainzer Naturwissenschaftliches Archiv 50: 383-397.

Filz, K.J. & Schmitt, T. (2013) Untersuchung der Arealdynamik des Kurzschwänzigen Bläulings (Cupido argiades, Pallas 1771) unter Klimawandelbedingungen mit Artverbreitungsmodellen in Europa (Lepidoptera: Lycaenidae). — Abhandlungen der Delattinia 38: 215-228.

Rákosy, L., Schmitt, T. & Varga, Z. (2011) Erebia oeme (Hübner, 1804) (Lepidoptera, Nymphalidae) in the Fagaras Mountains (Southern Carpathians). Entomologica romanica 16: 19-22.

Schmitt, T. (2005) Zygaena trifolii : observation of an autumn generation in southwestern Portugal (Lepidoptera : Zygaenidae). — Linneana Belgica 20: 48-50.

Schmitt, T. (2003) Cynthia virginiensis (Drury, 1773) : new records in central and southern Portugal (Lepidoptera. Nymphalidae). — Linneana Belgica 19: 95-98.

Schmitt, T. (2002) The Biology of Erebia medusa ([Denis & Schiffermüller],1775) in Central Europe (Lepidoptera). — Acta Biologica Debrecina 24: 113-129.

Schmitt, T. (2002) Biogeographie und regionale populationsgenetische Strukturierung von Polyommatus coridon (Lycaenidae: Lepoidoptera) im Saarland und in Rheinland-Pfalz. — Abhandlungen der Delattinia 28: 117-130.

Schmitt, T. & Stockhusen, N. (2001) Die Tagfalter und Widderchen des Bleichkopfs bei Jugenheim in Rheinhessen (Lepidoptera: Rhopalocera, Zygaenidae). — Pflanzen und Tiere in Rheinland-Pfalz (Berichtsjahr 2000) 11: 186-190.

Schmitt, T. (2001) Schmetterlinge und nachhaltige Forstwirtschaft – erste Ergebnisse aus dem Oettinger Forst. — GCD-Nachrichten 1/2001: 20-25.

Schmitt, T. (2001) Beobachtungen zum Eiablageverhalten und zu Raupenfutterpflanzen von Tagfaltern und Widderchen im südwestlichen Hunsrück im Jahr 2001. — Abhandlungen der Delattinia 27: 203-212.

Schmitt, T. (2000) Eine Erebia aethiopella (Hoffmannsegg, 1806) mit drei Fühlern (Lepidoptera: Nymphalidae, Satyrinae). — Nachrichten des entomologischen Vereins Apollo, N.F. 21: 191-192.

Schmitt, T. (2000) Vanessa atalanta (Linnaeus, 1758) am 13. März 1999 in Südwesthessen (Lepidoptera : Nymphalidae). — Nachrichten des entomologischen Vereins Apollo, N.F. 20: 320.

Schmitt, T. (2000) Beobachtungen zum Eiablageverhalten und zu Raupenfutterpflanzen von Tagfaltern im südwestlichen Hunsrück im Jahr 2000. — Abhandlungen der Delattinia 26: 143-148.

Schmitt, T. (1999) Die ökologischen Valenzen der Tagfalter des mittleren Löstertals. — Faunis­tisch-floristische Notizen aus dem Saarland 29: 585-614.

Schmitt, T. (1999) Eiablageverhalten und Raupenfutterpflanzen von Tagfaltern im südwestlichen Hunsrück. — Faunistisch-floristische Notizen aus dem Saarland 30: 659-670.

Schmitt, T. (1999) Nachtrag zu den ökologischen Valenzen der Tagfalter des mittleren Löstertals. — Faunistisch-floristische Notizen aus dem Saarland 30: 671-680.

Schmitt, T. (1998) Beobachtungen zur Eiablage von Lemonia dumi L. (Lepidoptera, Lemoniidae) im NSG Badstube bei Mimbach. — Faunistisch-floristische Notizen aus dem Saarland 28: 580-582.

Schmitt, T. (1998) Blütenpräferenzen von Tagfaltern im südwestlichen Hunsrück (Lepidoptera). — Nachrichten des entomologischen Vereins Apollo, N.F. 19: 161-204.

Schmitt, T. & Heidenreich, A. (1998) Untersuchungen zur Verbreitung des Kleinen Perlmutterfalters Issoria lathonia (Linnaeus, 1758) in Rheinland-Pfalz und im Saarland 1996 (Lepidoptera, Nymphalidae). — Atalanta 28: 271-276.

Schmitt, T. (1998) Tagfalterbeobachtungen in rheinland-pfälzischen Naturschutzgebieten. — Mainzer naturwissenschaftliches Archiv 36: 217-251.

Schmitt, T. (1993) Biotopansprüche von Erebia medusa brigobanna FRUHSTORFER, 1917 (Rundaugen-Mohrenfalter) im Nordsaarland. — Atalanta 24: 33-56.

Schmitt, T. (1991) Groß-Schmetterlinge als Bioindikatoren unter besonderer Berück-sichtigung der Lokalfauna des nördlichen Saarlandes. — Faunistisch-floristische Notizen aus dem Saarland. 22: 93-99.

Schmitt, T., Summkeller, R., Werno, A. & Zahm, N. (1991) Neue und wiederentdeckte Arten für die saarländische Lepidopterenfauna. — Faunistisch-floristische Notizen aus dem Saarland 22: 100-115.

Schmitt, T. (1991) Energetischer und ökonomischer Vergleich verschiedener Landwirt-schaftsmethoden. — Geographie und Schule 13, Heft 71: 39-44.


Schmitt, T. (2019) Besprechung / Review: Brockhaus, T. 2018: Die Eiszeitlibellen der Alten Welt – Pleistozäne Biogeographie paläarktischer Libellen. – Zoologica – Original Contributions to Zoology 163, Schweizerbart Science Publishers, Stuttgart: 145 S. – ISBN: 978-3-510-55050-0. — Contributions to Entomology 69: 78.

Schmitt, T. (2019) Besprechung / Review: Sobczyk, T.; Stöckel, D.; Graf, F.; Jornitz, H.; Karisch, T. & Wauer, S. 2018: Die Schmetterlingsfauna (Lepidoptera) der Oberlausitz. Teil 5: Kleinschmetterlinge (Microleidoptera) 1. Teil. – Entomologische Nachrichten und Berichte (Dresden), Beiheft 22, Dresden: 439 S. – ISSN: 0232-5535. — Contributions to Entomology 69: 86.

Schmitt, T. (2019) Besprechung / Review:Neu, P.; Malicky, H.; Graf, W. & Schmidt-Kloiber, A. 2018: Distribution Atlas of the European Trichoptera. – Die Tierwelt Deutschlands, 84. Teil. – ConchBooks, Harxheim: 891 pp. – ISBN: 978-3-939767-93-0. — Contributions to Entomology 69: 187-189.

Schmitt, T. (2019) Geleitwort. — In: Brockhaus, T. Die Eiszeitlibellen der Alten Welt – Pleistozäne Biogeographie paläarktischer Libellen. — Zoologica 163: 3.

Schmitt, T. (2019) Das große Insektensterben: Auch häufige Arten verschwinden. — Senckenberg 2018: 14-17.

Schmitt, T. (2018) Schmetterlingsparadiese und warum sie bei uns so selten geworden sind. — Natur • Forschung • Museum 147: 172–177.

Schmitt, T. (2018) Mit Schmetterlingen zur Professur. — In: Verband für Biologie, Biowissenschaften und Biomedizin in Deutschland e.V. – VBIO (ed.) Perspektiven. Berufsbilder von und für Biologen und Biowissenschaftler, 10. Aufl.: 25–27.

Schmitt, T. (2017) Besprechung / Review: Gaedike, R.; Nuss, M.; Steiner, A. & Trusch, R. (Hrsg.) 2017: „Verzeichnis der Schmetterlinge Deutschlands (Lepidoptera)“. – Entomologische Nachrichten und Berichte, Beiheft 21, Dresden: 362 S. – ISSN 0232-5535. — Contributions to Entomology 67: 362.

Schmitt, T. (2017) Besprechung / Review: Hofmann, A. F. & Tremewan, W. G. 2017: The natural history of burnet moths (Zygaena Fabricius, 1775) (Lepidoptera: Zygaenidae), part 1. – Proceedings of the Museum Witt, Munich, Vilnius: 631 pp. – ISBN 978-3-940732-32-3. — Contributions to Entomology 67: 324.

Schmitt, T. (2017) Besprechung / Review: Willner, W. 2017: „Taschenlexikon der Schmetterlinge Europas – Alle Tagfalter im Porträt“. – Quelle & Meyer, Wiebelsheim: 450 S. – ISBN 978-3-494-01633-7. — Contributions to Entomology 67: 254.

Schmitt, T. (2017) Besprechung / Review: Kunz, W. 2016: “Species Conservation in managed Habitats. The Myth of a Pristine Nature“. – Wiley-VCH, Weinheim: 273 pp. – ISBN: 978-3-527-33845-0. — Contributions to Entomology 67: 116.

Schmitt, T. (2017) Insektensammlungen: Wichtige Archive für den Naturschutz. — Natur • Forschung • Museum 147: 142–145.

Habel, J.C., Segerer, A. & Schmitt, T. (2016) „Früher gab es mehr Schmetterlinge“ Langzeitstudie untersucht Entwicklung von Tagfaltern und Widderchen. — Naturmagazin 3/2016: 12-16.

Schmitt, T. (2016) Besprechung / Review: Klausnitzer, B.; Klausnitzer, U.; Wachmann, E. & Hromátko, Z. 2016: Die Bockkäfer Mitteleuropas, in zwei Bänden; 3., stark überarbeitete und erweiterte Auflage. Neue Brehm Bücherei, Band 499 – Verlags KG Wolf, Magdeburg: 692 S. — Contributions to Entomology 66: 156-157.

Schmitt, T. (2016) Besprechung / Review Keil, T. 2015: Die Widderchen Mittelasiens. – 19. Beiheft der Entomologischen Nachrichten und Berichte: 84 S. — Contributions to Entomology 66: 112.

Schmitt, T. (2015) Editorial: Neues Design außen mit altgewohnter Qualität innen. — Abhandlungen der Delattinia 40: 5-6.

Schmitt, T. (2015) Besprechung / Review Kudrna, O.; Pennersdorfer, J. & Lux, K. 2015: Distribution Atlas of European Butterflies and Skippers. – Wissenschaftlicher Verlag PEKS, Schwanfeld, Deutschland. – ISBN 978-3-938249-71-0. — Contributions to Entomology 65: 326.

Schmitt, T. & Groll, E. (2015) Editorial. — Contributions to Entomology 65: 1-2.

Köhler, A. & Schmitt T. (2015) Die Entomologentagung 2015 in Frankfurt am Main: eine Nachlese. DGaaE-Nachrichten 29: 11-14.

Schmitt, T. (2015) Bericht über die Entomologentagung 2015 vom 2. bis 5. März in Frankfurt (Main). — Entomologische Nachrichten und Berichte 59: 74-76.

Nagel, P. & Schmitt, T. (2011) Nachruf auf Paul Müller 11.10.1940 – 30.05.2010. — Mitteilungen der Deutschen Zoologischen Gesellschaft 2011: 46-50.

Schmitt, T. & Schreiber, H. (2010) Nachruf Prof. Dr. Dr. h. c. mult. Paul Müller (1940-2010). Abhandlungen der Delattinia 35/36: 585-591.

Konvička, M., Kuras, T., Beneš, J., Čižek, O. & Schmitt, T. (2008) Endemičtí okáči vysokých Sudet: Vliv změn klimatu a hospodaření na populace vysokohorských motýlů. — Vesmír 87: 516-521.

Schmitt, T. (2006) Rettet die bunten Gaukler der bunten Sommerwiesen. — NABUlletin 2006.2: 15-17.

Schmitt, T. (2004) Mit Schmetterlingen zum Professor für Molekulare Biogeographie. — Pp. 20-21 in: Verband Deutscher Biologen (ed.) Perspektiven. Berufsbilder von und für Biologen, 4. Aufl.

Schmitt, T. & Stockhusen, N. (2001) Die Tagfalter und Widderchen auf den NABU-Flächen am Bleichkopf. — NABUlletin 2001.1: 26-27.

Schmitt, T. (1996) Magerwiesen – Kleinodien im Nordsaarland. — Naturschutz im Saarland 96.2: 30-32.

Schmitt, T. (1994) Kein Sommer ohne Schmetterlinge. — In: Vogel- und Pflanzenschutz­verein Otzenhausen (Hrsg.) Begleiter durch die heimische Natur: 69-73.

Prof. Dr.  Zsolt Végvári
Wissenschaftler, Redaktionsleitung Zeitschriften
Assistant Editor
Cornelia Grunow
Editorial Assistant