Senckenberg Research Institute and Natural History Museum Frankfurt/M.

Terrestrial Zoology

At the Department of Terrestrial Zoology we study the diversity and evolution of land animals. Our extensive collections are the basis of this research, but also a vital platform for international collaboration and exchange. In six sections we examine the present-day, historical and ancient diversity of birds, amphibians, reptiles, spiders and insects and their evolution. The results of this research mainly contribute to Senckenberg’s Research Fields Biodiversity, Systematics and Evolution and Biodiversity and Environment.


Prof. Dr. Steffen U. Pauls
Head of Department Terrestrial Zoology / Head of Section Entomology III
Forschungsthemen / Research Interests
  • Molecular Systematics, Evolution and Phylogeography of Caddisflies (Trichoptera)
  • Integrative Taxonomy and Biodiversity Research
  • Population Genetics and Evolutionary Ecology of Aquatic Insects
  • Biodiversity Genomics with Museum Specimens

(* corresponding author; +equal contribution; undergraduate student)


Hjalmarsson, A.E.*, Jähnig, S.C., Cai, Q., Sharma, S., Tong, X., Li, F., Shah, D.N., Tachamo Shah, R.D. & Pauls, S.U.* (2019) Multi-locus molecular phylogeny establishes subgeneric groups and informs the biogeographic history of Himalopsyche (Trichoptera, Rhyacophilidae). Systematic Entomology 44: 973-984.

Ngera, M.F., Pauls, S.U., Holzenthal, R.W., Bagalwa, M., Bisimwa, M.A., Mushayuma, E.M. & Cammaerts, D.R. (2019) Contribution to the knowledge of the aquatic macroinvertebrate fauna of the streams of Kahuzi-Biega National Park, Democratic Republic of Congo. African Journal of Aquatic Sciences 44: 127-142 .


Bálint, M.*, Nowak, C., Márton, O., Pauls, S.U., Wittwer, C., Luis Aramayo, J., Schulze, A., Chambert, T., Cocchiararo, B. & Jansen, M. (2018) Accuracy, limitations and cost efficiency of eDNA‐based community survey in tropical frogs. Molecular Ecology Resources 18: 1415-1426.

Datry, T., Foulquier, A., Corti, R., von Schiller, D., Tockner, K., Mendoza-Lera, C., Clément, J. C., Gessner, M. O., Moleón, M., Stubbington, R., Gücker, B., Albariño, R., Allen, D. C., Altermatt, F., Arce, M. I., Arnon, S., Banas, D., Banegas-Medina, A., Beller, E., Blanchette, M. L., Blanco-Libreros, J. F., Blessing, J. J., Boëchat, I. G., Boersma, K. S., Bogan, M. T., Bonada, N., Bond, N. R., Brintrup Barría, K. C., Bruder, A., Burrows, R. M., Cancellario, T., Canhoto, C., Carlson, S. M., Cauvy-Fraunié, S., Cid, N., Danger, M., de Freitas Terra, B., De Girolamo, A. M., de La Barra, E., del Campo, R., Diaz-Villanueva, V. D., Dyer, F., Elosegi, A., Faye, E., Febria, C., Four, B., Gafny, S., Ghate, S. D., Gómez, R., Gómez-Gener, L., Graça, M. A. S., Guareschi, S., Hoppeler, F., Hwan, J. L., Jones, J. I., Kubheka, S., Laini, A., Langhans, S. D., Leigh, C., Little, C. J., Lorenz, S., Marshall, J. C., Martín, E., McIntosh, A. R., Meyer, E. I., Miliša, M., Mlambo, M. C., Morais, M., Moya, N., Negus, P. M., Niyogi, D. K., Papatheodoulou, A., Pardo, I., Pařil, P., Pauls, S. U., Pešić, V., Polášek, M., Robinson, C. T., Rodríguez-Lozano, P., Rolls, R. J., Sánchez-Montoya, M. M., Savić, A., Shumilova, O., Sridhar, K. R., Steward, A. L., Storey, R., Taleb, A., Uzan, A., Vander Vorste, R., Waltham, N. J., Woelfle-Erskine, C., Zak, D., Zarfl, C., & Zoppini, A. 2018. A global analysis of terrestrial plant litter dynamics in non-perennial waterways. Nature Geoscience 11: 497-503.

Dossi, F.*, Leitner, P., Pauls, S. & Graf, W. (2018) In the mood for wood – habitat specific colonization patterns of benthic invertebrate communities along the longitudinal gradient of an Austrian river. Hydrobiologia 805: 245-258.

Graf, W.*, Pauls, S.U. & Vitecek, S. (2018) Isoperla vjosae sp. n., a new species of the Isoperla tripartita-group from Albania (Plecoptera: Perlodidae). Zootaxa 4370: 171–179

Hjalmarsson, A.E.*, Graf, W., Jähnig, S.C., Vitecek, S. & Pauls, S.U. (2018) Molecular association and morphological characterization of Himalopsyche larval types (Trichoptera, Rhyacophilidae). Zookeys, ZooKeys 773: 79-108.

Hoppeler, F.*, Winkelmann, C., Becker, J. & Pauls, S.U. (2018) Larval growth and metabolic energy storage of Micropterna lateralis (Trichoptera: Limnephilidae) in an intermittent stream: glycogen dominates in final instars. Hydrobiologia 806: 175-185.

Tonkin, J.D.*, Altermatt, F. Finn, D., Heino, J., Olden, J.D., Pauls, S.U. & Lytle, D.A. (2018) The role of dispersal in river network metacommunities: patterns, processes, and pathways. Freshwater Biology 63:141-163.

Waringer, J.*, Gonzalez, M.A., Martin, L., Martinez, J., Erzinger, F. & Pauls, S.U. (2018) DNA-based association and description of the larval stage of Apatania theischingerorum Malicky 1981 (Trichoptera, Apataniidae), with notes on its ecology. Zootaxa 4418: 161–170.


Szivák, I., Mikes. T., Szalontai, B., Kučinić, M., Vučković, I., Vadkerti, E., Kisfali, P., Pauls, S.U. & Bálint, M.* (2017) Ecological divergence of Chaetopteryx rugulosa species complex (Insecta, Trichoptera) linked to climatic niche diversification. Hydrobiologia 794:31-47. Full Text

Tonkin, J.D.*, Tachamo Shah, R.D., Shah, D.N., Hoppeler, F., +Jähnig, S.C. & +Pauls, S.U. (2017) Metacommunity structuring in Himalayan streams over large elevational gradients: the role of dispersal routes and niche characteristics. Journal of Biogeography 44: 62-74. Full Text

Vitecek, S.*, Vincon, G., Graf, W. & Pauls, S.U. (2017) High cryptic diversity in aquatic insects: an integrative approach to study the enigmatic Leuctra inermis species group (Plecoptera). Arthropod Systematics & Phylogeny 76: 497-521. Full Text

Vitecek, S.*, Kučinić, M., Previšić, A., Živić, I., Stojanović, K., Keresztes, L., Bálint, M., Hoppeler, F., Waringer, J., Graf, W. & Pauls, S.U.* (2017) Integrative taxonomy by molecular species delimitation: multi-locus data corroborate a new species of Balkan Drusinae micro-endemics. BMC Evolutionary Biology 17:129. Full Text


Hoppeler, F., Tachamo-Shah, R.D., Shah, D., Tonkin, J.D., Jähnig, S.C., Sharma, S., & Pauls, S.U.* (2016) Ecological characterisation of an unknown fauna using DNA taxonomy – An example with Himalayan aquatic insects. Freshwater Biology 61: 1905-1920.

Hoppeler, F.*, Rotter, B., Krezdorn, N. & Pauls, S.U. (2016) A larval Transcriptome of the limnephilid Caddisfly Micropterna lateralis. Aquatic Insects 37: 253-257.

Ibrahimi, H.*, Vitecek, S., Previšić, A., Kučinić, M., Waringer, J., Graf, W., Balint, M., Keresztes, L. & Pauls, S.U. (2016) Drusus sharrensis sp. n. (Trichoptera, Limnephilidae), a new species from Sharr National Park in Kosovo, with molecular and ecological notes. ZooKeys 559: 107–124. Full Text

Li, F., Shah, D.N., Pauls, S.U., Qu, X., Cai, Q. & Tachamo Shah, R.D.* (2016) Elevational shifts of freshwater communities cannot catch up climate warming in the Himalaya. Water 8(8):327 (doi:10.3390/w8080327) Full Text

Waringer, J.*, Graf, W., Bálint, M., Kučinić, M., Pauls, S.U., Previšić, A., Keresztes, L., & Vitecek, S. (2016) Project overview: the Drusinae (Insecta, Trichoptera) in a World of global change—bridging basic and applied research in a highly sensitive aquatic insect group. Zoosymposia 10: 432–438.

Waringer, J.*+, Previšić, A.+, Kučinić, M., Graf, W., Vitecek, S., Keresztes, L., Bálint, M. & Pauls, S.U. (2016) Larval morphology of the Western Balkans endemic caddisflies Drusus krusniki Malicky 1981, D. vernonensis Malicky 1989, and D. vespertinus Marinković-Gospodnetić 1976 (Trichoptera, Limnephilidae, Drusinae). Zootaxa 4083: 483-500. PDF

Zhou, X., Frandsen, P., Holzenthal, R., Beet, C.R., Bennett, K.R., Blahnik, R., Bonada, N., Cartwright, D., Chuluunbat, S., Cocks, G.V., Collins, G.E., de Waard, J., Dean, J., Flint, O., Razo Gonzalez, M., Hausmann, A., Hendrich, L., Hess, M., Hogg, I.D., Kondratieff, B.C., Malicky, H., Morinière, J., Morse, J.C., Mwangi, F.N., Pauls, S.U., Rinne, A., Robinson, J., Salokannel, J., Shackleton, M., Smith, B., Stamatakis, A., StClair, R., Thomas, J., Zamora-Muñoz, C., Ziesmann, T., & Kjer, K.M. (2016) The Trichoptera barcode initiative: strategies for integrative taxonomy and generating a species-level tree of life. Philosophical Transactions of the Royal Society. 371 : 20160025. Full Text


Favre, A.*, Päckert, M., Pauls, S.U., Jähnig, S. C., Uhl, D., Michalak, I., & Muellner-Riehl, A. (2015) The role of the uplift of the Qinghai-Tibetan Plateau for the evolution of Tibetan biotas. Biological Reviews 90: 236-253. PDF

Geismar, J., Haase, P., Nowak, C., Sauer, J. & Pauls, S.U.* (2015) Local population genetic structure of the montane caddisfly Drusus discolor is driven by overland dispersal and spatial scaling. Freshwater Biology 60: 209-221. Full Text

Ibrahimi, H.*, Mladen K., Vitecek, S., Waringer, J., Graf, W., Previšić, A., Bálint, M., Keresztes, L., & Pauls, S.U. (2015) New records for the Kosovo caddisfly fauna with the description of a new species, Drusus dardanicus sp. nov. (Trichoptera: Limnephilidae). Zootaxa 4032: 551–568. Abstract

Macher, J.-N., Rozenberg, A., Pauls, S.U., Tollrian, R., Wagner, R. & Leese, F.* (2015) Assessing the phylogeographic history of the montane caddisfly Thremma gallicum: a comparison of sequence data from mitochondria and restriction-site associated DNA markers (RAD). Ecology and Evolution 5: 648-662. Full Text

Tachamo-Shah, R.D.*, Sharma, S., Haase, P., Jähnig, S.+ & Pauls, S.U.+* (2015) The climate sensitive zone along an altitudinal gradient in central Himalayan rivers: a useful concept to monitor climate change impacts in mountain regions. Climatic Change 132: 265-278. Abstract

Vitecek, S.*, Graf, W., Kučinić, M., Oláh , J., Bálint , M., Previšić , A., Keresztes, L., Pauls, S.U.* & Waringer, J. (2015) A hairy case: The evolution of filtering carnivorous Drusinae (Limnephilidae, Trichoptera). Molecular Phylogenetics & Evolution 93: 249-260. Abstract

Vitecek, S.*, Kučinić, M., Oláh, J., Previšić, A., Bálint, M., Keresztes, L., Waringer, J., Pauls, S.U. & Graf, W. (2015) Description of two new filtering carnivore Drusus species (Limnephilidae, Drusinae) from the Western Balkans. Zookeys 513: 79-104. Full Text

Vitecek, S.*, Previšić, A., Kučinić, M., Bálint, M., Keresztes, L., Waringer, J., Pauls, S.U., Malicky, H. & Graf, W.* (2015): Description of a new species of Wormaldia from Sardinia and a new Drusus species from the Western Balkans (Trichoptera, Philopotamidae, Limnephilidae). ZooKeys 496: 85-103. Full Text

Waringer, J., Graf, W., Bálint, M., Kučinić, M., Pauls, S.U., Previšić, A., Keresztes, L., Ibrahimi, H., Živić, I., Bjelanovic, K., Krpač, V. & Vitecek, S. (2015). Larval morphology and phylogenetic position of Drusus balcanicusKumanski, Drusus botosaneanui Kumanski, Drusus serbicus Marinković-Gospodnetić and Drusus tenellus (Klapálek) (Trichoptera: Limnephilidae: Drusinae). European Journal of Entomology 112: 344-361. PDF

Waringer, J.*, Lubini, V., Hoppeler, F. & Pauls, S.U. (2015) DNA-based association and description of the larval stage of Apatania helvetica Schmid 1954 (Trichoptera, Apataniidae) with notes on ecology and zoogeography. Zootaxa 4020: 244–256. Abstract


Dijkstra, K.-D.B*, Monaghan, M.T. & Pauls, S.U. (2014) Freshwater Biodiversity and Insect Diversification. Annual Review of Entomology 59: 143–63. Abstract Full Text

Pauls, S.U.*, Alp, M., Bálint, M., Bernabò, P., Čiampor Jr., F., Čiamporová-Zaťovičová, Z., Finn, D.S., Kohout, J., Leese, F., Lencioni, V., Paz-Vinas, I. & Monaghan, M.T. (2014). Integrating molecular tools into freshwater ecology: developments and opportunities. Freshwater Biology 59: 1559-1576. Abstract

Previšić, A.*+, Schnitzler, J., Kučinić, M., Graf, W., Ibrahimi, H., Kerovec, M., & Pauls, S.U.*+ (2014) Micro-Scale Vicariance and Diversification of Western Balkan Caddisflies Linked to Karstification. Freshwater Science 33: 250-262. Full Text

Previšić A., Graf W., Vitecek S., Kučinić M., Bálint M., Keresztes L., Pauls, S.U. & J. Waringer. (2014) Cryptic diversity of caddisflies in the Balkans: the curious case of Ecclisopteryx species (Trichoptera: Limnephilidae). Arthropod Systematics & Phylogeny 72: 309-329. PDF

Shah, D.N., Domisch, S., Pauls, S.U., Haase, P. & Jähnig, S.* (2014) Current and future latitudinal gradients in stream macroinvertebrates richness across North America. Freshwater Science 33: 1136-1147. Abstract


Domisch, S.*, Araújo, M.B., Bonada, N., Pauls, S.U., Jähnig, S.C.+ & Haase, P.+ (2013) Modelling distribution in European stream macroinvertebrates under future climates. Global Change Biology 19: 752–762. Abstract

Pauls, S.U.*, Nowak, C., Bálint, M. & Pfenninger, M.* (2013) The impact of global climate change on genetic diversity within populations and species. Molecular Ecology 22: 925-946. Abstract

Kučinić, M.*+, Szivák+, Bálint, M., Delić, A., Pauls, S.U., & Vučković, I. (2013) Chaetopteryx bucari sp. n. a new species of from the Chaetopteryx rugulosa group from Croatia (Insecta, Trichoptera) with some molecular, taxonomical and ecological notes on the group. ZooKeys 320: 1-28. HTML

Theissinger, K.*, Bálint, M., Feldheim, K.A., Haase, P., Johannesen, J., Laube, I. & Pauls, S.U. (2013) Glacial survival and post-glacial recolonization of an arctic–alpine freshwater insect (Arcynopteryx dichroa, Plecoptera, Perlodidae) in Europe. Journal of Biogeography 40: 236-248. Abstract

Waringer, J.*, Graf, W., Bálint, M., Kučinić, M., Pauls, S.U., Previšić, A., Keresztes, L., & Vitecek, S. (2013) The larvae of Drusus franzressli Malicky 1974 and Drusus spelaeus (Ulmer 1920 (Trichoptera: Limnephilidae: Drusinae) with notes on ecology and zoogeography. Zootaxa 3637: 1–16. Abstract

Waringer, J.*, Graf, W., Bálint, M., Kučinić, M., Pauls, S.U., Previšić, A., Keresztes, L., & Vitecek, S. (2013) The larva of Drusus vinconi Sipahiler, 1992 (Trichoptera, Limnephilidae, Drusinae). ZooKeys 317: 69-80. HTML


Bagalwa, M., Zirirane, N., Pauls, S., Karume, K., Ngera, M., Bisimwa, M. & N. Mushagalusa (2012) Aspects of the Physico-Chemical Characteristics of Rivers in Kahuzi-Biega National Park, Democratic Republic of Congo. Journal of Environmental Protection 3: 1590-1595. PDF

Haase, P., Hering, D., Hoffmann, A., Müller, R., Nowak, C., Pauls, S.U., Stoll, S.U. & D. Straile (2012) Auswirkungen auf limnische Lebensräume. In Mosbrugger, V., Brasseur, G., Schaller, M. & Stribrny, B. (eds) Klimawandel und Biodiversität – Folgen für Deutschland. Wissenschaftliche Buchgesellschaft, Darmstadt, pp 91-105.

Kautz, S.*, Ballhorn, D.J., Kroiß, J., Pauls, S.U., Moreau, C.S., Eilmus, S., Strohm, E. & Heil, M. (2012) Host plant use by competing acacia‑ants: Mutualists monopolize while parasites share hosts. PLoS One 7: e37691 HTML

Malicky, H. & Pauls, S.U.* (2012) Crossbreeding of Chaetopteryx morettii and related species, with molecular and eidonomical results (Trichoptera, Limnephilidae). Annales de Limnologie – International Journal of Limnology 48: 13-19. Abstract

Pfenninger, M.*, Bálint, M., & Pauls. S.U. (2012) Methodological framework for projecting the potential loss of intraspecific genetic diversity due to global climate change. BMC Evolutionary Biology 12: 224. HTML

Sipahiler, F. & Pauls, S. (2012) Two new species of the genus Schizopelex McLachlan, from northern Turkey (Trichoptera, Sericostomatidae). Munis Entomology & Zoology 7: 184-190.


The following paper was featured in the Faculty of 1000 and ranked “Must Read”:

Bálint, M., Domisch, S., Engelhardt, C.H.M., Haase, P., Lehrian, S., Sauer, J., Theissinger, K., +Pauls, S.U.*, +Nowak, C.* (2011) Cryptic biodiversity loss linked to global climate change. Nature Climate Change 1: 313–318.Abstract

Bálint, M., Ujvárosi, L., Theissinger, K., Mészáros, N., Lehrian, S. & Pauls, S.U.* (2011) A Neglected Diversity Hotspot: Genetic Variation in the Carpathians. pp 189-205 in Habel, J. (ed.) Biodiversity Hotspots in Europe. Springer.

+Engelhardt, C.H.M., Haase, P. & +Pauls, S.U.* (2011) From the Western Alps across Central Europe: Postglacial recolonisation of the tufa stream caddisfly Rhyacophila pubescens (Insecta, Trichoptera). Frontiers in Zoology 8:10. HTML

Graf, W.*, Pauls, S.U., & Waringer, J. (2011) The larva of Ecclisopteryx malickyi Moretti, 1991 (Trichoptera: Limnephilidae: Drusinae) with comments on the genus. Zoosymposia 5: 136-142. PDF

Pauls, S.U.* & Bálint, M. (2011) Aquatic Insect Diversification and Speciation in the Carpathians – Case Studies on Caddisflies. In Ujvárosi, L. & Marko, B. (eds.) Romanian Carpathians, genetic centres or barriers in the process of postglacial isolation and speciation. Case studies in selected organisms. University Press, Cluj-Napoca.

Taubmann, J., Theissinger, K.*, Feldheim, K.A., Laube, I., Graf, W., Haase, P., Johannesen, J. & Pauls, S.U.* (2011) Modelling range shifts and assessing genetic diversity distribution of the montane aquatic mayfly Ameletus inopinatus in Europe under climate change scenarios. Conservation Genetics 12: 503-515. Abstract

Theissinger, K., Laube, I., Bálint, M., Johannesen, J., Haase, P. & Pauls, S.U.* (2011) Species distribution models and molecular data reveal the Pleistocene history of the cold-adapted mayfly Ameletus inopinatus(Ephemeroptera: Siphlonuridae) in Europe. Freshwater Biology 56: 2554–2566. Abstract

Waringer, J.*, Graf, W., Pitsch, T., Pauls, S.U., Previšić, A., & Kučinić, M. (2011) Description of the larval stage of Drusus mixtus (Pictet, 1834) (Trichoptera: Limnephilidae: Drusinae) with notes on ecology and zoogeography. Limnologica 41: 249-255. Abstract


Craft, K.J., Pauls, S.U., Darrow, K., Miller, S.E., Hebert, P.D.N., Novotny, V. & Weiblen, G.* (2010) Population genetics of ecological communities with DNA barcodes: An example from New Guinea Lepidoptera. Proceedings of the National Academy of Sciences 107: 5041-5046. Abstract

Haase, P.*, Pauls, S.U., Schindehütte, K. & Sundermann, A. (2010) First audit of macroinvertebrate samples from an EU-Water Framework Directive monitoring programme: human error greatly lowers precision of assessment results. Journal of the North American Benthological Society 29: 1279-1291. HTML

+Holzenthal, R.W.*, +Robertson, D.R., +Pauls, S.U., & +Mendez, P.K. (2010) Taxonomy and Systematics: Contributions to Benthology and Future Directions. Journal of the North American Benthological Society, 29: 147-169. HTML

Lehrian, S., Bálint, M., Haase, P. & Pauls, S.U.* (2010) Genetic population structure of an autumn emerging caddisfly with inherently low dispersal capacity and insights into its phylogeography. Journal of the North American Benthological Society 29: 1100-1118. HTML

Monaghan, M.T.*, Pauls, S.U., Hughes, J.M., & Robinson, C.T. (2010) Evolutionary ecology, speciation, and molecular systematic in freshwater organisms – Preface. Journal of the North American Benthological Society 29: 1024-1026. HTML

Pauls, S.U.*, Blahnik, R.J., Zhou, X., Wardwell, T.C. & Holzentahl, R.W. (2010) DNA barcode data confirm new species and reveal cryptic diversity in Chilean Smicridea (Smicridea) (Trichoptera: Hydropsychidae). Journal of the North American Benthological Society 29: 1058-1074. HTML

Pauls, S.U.*, Holzenthal, R.W., & Mwangi. F.N. (2010) Two new species of Triaenodes McLachlan, 1865 from streams in the Lake Kivu basin, South Kivu, Democratic Republic of the Congo (Trichoptera, Leptoceridae). Denisia 29: 277-285. Abstract

Waringer, J.*, Graf, W., Pauls, S.U., Previšić, A. & Kučinić, M. (2010) A larval key to the Drusinae species of Austria, Germany, Switzerland and the dinaric western Balkan. Denisia 29: 383-406. Abstract


Graf, W.*, Pauls, S.U. & Waringer, J. (2009) The Larva of Rhyacophila ferox Graf, 2006 (Trichoptera: Rhyacophilidae) from the Eastern Alps (Carinthia, Austria). Aquatic Insects 31: 111-117. Abstract

Graf, W.*, Waringer, J. & Pauls, S.U. (2009) A new morpho-ethological feeding group within larval Drusinae (Trichoptera : Limnephilidae): the alpinus-group sensu Schmid, 1956, including larval descriptions of Drusus franzi, 1956 and Drusus alpinus Meyer-Dür, 1875, with notes on their systematic position, ecology and zoogeography. Zootaxa 2031: 53-62. Abstract

Kautz, S.,* Pauls, S.U., Ballhorn, D.J., Lumbsch, H.T. & Heil, M. (2009) Polygynous supercolonies of the acacia-ant Pseudomyrmex peperi, an inferior colony founder. Molecular Ecology 18: 5180-5194. Abstract

Lehrian, S.*, Pauls, S.U. & Haase, P. (2009) Contrasting patterns of population structure in the montane caddisflies Hydropsyche tenuis and Drusus discolor in central European highlands. Freshwater Biology 54: 283-295. Abstract

Highlighted as Fast-Breaking Paper in Plant & Animal Science December 2010 by Thomson Reuters Essential Science Indicators:

Pauls, S.U.*, Theissinger, K., Ujvarosi, L., Bálint, M. & Haase, P (2009) Population structure in two closely related, partially sympatric caddisflies in Eastern Europe: historic introgression, limited dispersal and cryptic diversity. Journal of the North American Benthological Society 28: 517-536. PDF HTML

Theissinger, K., Feldheim, K.A., Seitz, A., Pauls, S.U.* (2009) Isolation and characterization of 11 polymorphic trinucleotide microsatellite markers in the stonefly Arcynopteryx compacta (Plecoptera: Perlodidae). Molecular Ecology Resources 9: 357-359. Abstract


Engelhardt, C.*, Pauls, S.U. & Haase, P. (2008) Population genetic structure of the caddisfly Rhyacophila pubescens, Pictet 1834, north of the Alps. Fundamental and Applied Limnology – Archiv für Hydrobiologie 173: 165-176. Abstract

Haase, P.*, Pauls, S.U., Engelhardt, C. & Sundermann, A. (2008) Effects of sampling microhabitats with low coverage within the STAR/AQEM macroinvertebrate sampling protocol on stream assessment. Limnologica 38: 14-22. Abstract

Pauls, S.U.*, Graf, W., Haase, P., Lumbsch, H.T. & Waringer, J. (2008) Grazers, shredders and filtering carnivores – The evolution of feeding ecology in the Drusinae (Trichoptera: Limnephilidae): insights from a molecular phylogeny. Molecular Phylogenetics and Evolution 46: 776-791. Abstract

Sundermann, A.*, Pauls, S.U., Clarke, R.T. & Haase, P. (2008) Within-stream variability of benthic invertebrate samples and EU Water Framework Directive assessment results. Fundamental and Applied Limnology – Archiv für Hydrobiologie 173: 21-34. Abstract

Theissinger, K., Feldheim, K.A., Taubmann, J., Seitz, A. & Pauls, S.U.* (2008) Isolation and characterization of 10 highly polymorphic di- and trinucleotide microsatellite markers in the mayfly Ameletus inopinatus(Ephemeroptera: Siphlonuridae). Molecular Ecology Resources 8: 1285-1287. Abstract

Waringer, J.*, Graf, W., Pauls, S.U. & Cianficconi, F. (2008) The Larvae of Drusus improvisus McLachlan, 1884, Drusus camerinus Moretti, 1981 and Drusus aprutiensis Moretti, 1981 (Trichoptera: Limnephilidae: Drusinae). Aquatic Insects 30: 69-79. Abstract

Waringer, J.*, Graf, W., Pauls, S.U., Vincentini, H. & Lubini, V. (2008) DNA based association and description of the larval stage of Drusus melanchaetes McLachlan, 1876 (Trichoptera: Limnephilidae: Drusinae) with notes on ecology and zoogeography. Limnologica 38: 34-42. Abstract


Pauls, S.U.*, Feldheim, K.A. & Haase, P. (2007) Isolation and characterization of microsatellite markers in the caddisfly Drusus discolor (Trichoptera: Limnephilidae). Molecular Ecology Notes 7: 150-152. Abstract

Waringer, J.*, Graf, W., Pauls, S. & Lubini, V. (2007) The Larva of Drusus nigrescens Meyer-Dür, 1875 (Trichoptera: Limnephilidae: Drusinae) with notes on its ecology, genetic differentiation and systematic position. Annales de Limnologie – International Journal of Limnology 43: 161-166. PDF


Haase, P.*, Murray-Bligh, J., Gunn, R., Lohse, S., Pauls, S. & Sundermann, A. (2006) An assessment of the impact of sorting and identification errors of macroinvertebrate samples. Hydrobiologia 566: 505-521. Abstract

Pauls, S.U.*, Lumbsch, H.T. & Haase, P. (2006) Phylogeography of the montane caddisfly Drusus discolor: evidence for multiple refugia and periglacial survival. Molecular Ecology 15: 2153-2169. Abstract

2002 to 2005

Graf, W.*, Lubini, V. & Pauls, S. (2005) Larval description of Drusus muelleri McLachlan, 1868 (Trichoptera: Limnephilidae) with some notes on its ecology and systematic position within the genus Drusus. Annales de Limnologie – International Journal of Limnology 42: 93-98. PDF

Haase, P.*, Lohse, S., Pauls, S., Schindehütte, K., Sundermann, A., Rolauffs, P. & Hering, D. (2004) Assessing streams in Germany with benthic invertebrates: development of a practical standardised protocol for macroinvertebrate sampling and sorting. Limnologica 34: 349-365. Abstract

Haase, P.*, Pauls, S., Sundermann, A. & Zenker, A. (2004) Testing different sorting techniques in macroinvertebrate samples from running waters. Limnologica 34: 366-378. Abstract

Pauls, S., Feld, C., *Sommerhäuser, M.* & Hering, D. (2002) New concepts in assessing lowland streams in Germany in accordance with the EU Water Framework Directive. Wasser & Boden 54: 70-77.